BioStor
(specimen not known in GBIF)
 | A review of the Centrolenid frogs of Ecuador, with descriptions of new species John D Lynch and William E Duellman (1973) Occasional Papers University of Kansas Museum of Natural History 16: 1-66 |
 | Frogs of Columbia Doris M Cochran and Coleman J Goin (1970) Bull U S National Museum 288: 1-655 |