448 MR. L. DE NICtviLLE ON NEW OR [May 3, 5. Descriptions of some new or little-known Butterflies from India, with some Notes on the Seasonal Dimorphism obtaining in the Genus Melanitis. By Lionel de NiCEVILLE, F.E.S. [Eeceived May 3, 1887.] (Plates XXXIX. & XL.) Lethe nicetella, n. sp. (Plate XXXIX. fig. 5, c^ .) Hab. Sikkim, Expanse. S ^ r8 to 2 inches. Male. Upperside : both wings brown, with a brilHant golden- bronzy sheen when fresh, the outer margins with a regular band of the ground-colour devoid of this gloss. Fore wine/ with a short obscure ochreous subapical fascia at the bifurcation of the fourth and fifth subcostal nervules. Hind wing with a submarginal series of five round black spots with pale outer rings, of which the second from the apex is the largest, the uppermost and the fourth subequal, and the next largest, the third the smallest, though but little smaller than the fifth or anal. Underside : both wings paler than above. Fore wing with an obscure dark bar across the middle of the cell, an oblique discal band beyond the cell, prominent at the costa, be- coming obsolete towards the anal angle, beyond this band the ground- colour is abruptly paler ; the subapical fascia as on the upperside, but more extended violet-white ; two marginal fine dark lines en- closing a line of the ground-colour, the inner one defined with a somewhat coarser pale line, all these lines extending from the second median nervule to the apex. Hind wing with two basal and two discal highly irregular violet lines, the one furthest from, the base with an irregular dark brown fascia placed outwardly against it ; the discocellular nervules defined on both sides with a fine violet line ; a submarginal series of six ocelli, the first, second (usually), fifth, and geminated sixth composed of a black centre with a pure white pupil, an ochreous, a dark brown, and an outer violet ring ; the second (occasionally), third, and fourth ocelli composed of a small dark brown centre with a violet pupil, a violet, a dark brown, and an outer violet ring ; marginal lines as in the fore wing, but the innermost line violet, very prominent towards the anal angle. Cilia cinereous, dark brown at the ends of the veins. — Female. Upper- side : both wings paler. Fore wing with the discal band of the underside showing through. Underside ; both wings also paler. L. nicetella may be known from L. nicetas, Hewitson, from Kulu and Sikkim, on the underside of the fore wing in both sexes by not possessing three small subapical ocelli divided by the discoidal ner- vules, and in the discal band being much less prominent (in L. nicetas it is broadly outwardly defined with ochreous) ; in the iiind wing in having at least two and often three of the ocelli with small dark brown instead of deep black centres (in L. nicetas all the ocelli are P.Z.S. 1887,?!. XXXIX. I 'W.Purkiss litK , TMEW INDIAN BUTTERFLIES Han^iarb imp. OCR text unavailable for this page.P Z.S.1887 Pl.XL. "A.rurK-.ss liii'. NEW raDIAlT 3"jt:i?.?l:zs Han.'idrb ir mp I 1887.] LITTLE-KlNOWN BUTTERFLIES FROM INDIA. 449 similarly formed), and in the outer discal violet line having a dark brown fascia placed against it (in L. nicetas this fascia is much more deeply scalloped and of a rich ochreous colour). I have described L. niceiella from a laro;e series of males and a single female collected in Sikkim by Mr. Otto MoUer. In Sikklm, judging from the number of specimens before me of each species, L. nice(ella\s much the commoner of the two. Lethe tamuna, n. sp. (Plate XXXIX. fig, 6,2 ■) Hub. Little Nicobar. Expanse. $ 2v inches. Female. Upperside : both wings dull dark fulvous. Fore wing with the apical half fuscous, bearing two pure white spots below the costa, placed midway between the apex and the median ochreous band, the upper of the two spots much the smaller, and divided into two portions by the fourth subcostal nervule ; below tliese spots in the upper discoidal interspace is an obscure oval black spot ; near the margin are four bright ochreous lunulas placed between the veins from the lower discoidal nervule to the inner angle, beyond these lunnles in each interspace is a fine ochreous line ; across the disk of the wing, from the middle of the costa to near the inner angle, is a broad bright ochreous band, its inner edge nearly sti'aight and even, its outer edge produced into points between the veins, the lower portion of the band composed of two spots (the lower one very small) in the submedian interspace. Hindwing with the ocelli of the underside more or less showing through by transparency ; a series of bright ochreous lunules with inner dark borders placed near the outer margin between the veins, with a darker ochreous line beyond. Underside : both wi?igs dull brown. Fore wing with a whitish subbasal line crossing the middle of the discoidal cell from the subcostal to the submedian nervure ; the broad discal band as above, but with its edges more even and wider at its lower end ; beyond it are four ill-shaped ocelli with black pupils dotted with white, a pale violet ring, then a browner ring and an outer pale violet ring ; the margin marked much as above. Hind wing with a subbasal line in continuation of that on the fore wing, not reaching the abdominal margin ; a discal series of ocelli placed on a pale violet band, which more or less follows their outline ; the upper ocellus very large, its centre deep black dotted with white, then a broad rich ochreous ring, outwardly defined with a fuscous ring ; the next largest ocellus is in the first median interspace, with two small, equal-sized ocelli in the interspaces above and below it, and two very small and indistinct ocelli diAuded by the discoidal nervule; the bright ochreous lunules on the margin of the upperside white, almost silvery, on the underside. Lethe tamuna is a local form of the widely-distributed L. europa, Fabricius, which occurs in the Andamans (but is replaced apj)a- rently in the Nicobars by the specits under notice), almost through- out India (though not in Ceylon), the Malay |)eninsula and islands, and has been recoidcd from China. L. tamuna 2 "'f^y b^ ^^ once 450 MR. L. DE NICEVILLE ON NEW OR [May 3, distinguished from the same sex of L. europa by its more tawny coloration on the upperside, the discal band bright ochreous instead of white, with its outer edge more irregular, and by the underside of the hind wing having four well-formed perfect ocelli ; L. europa the black middle portion of all the ocelli except the upper one is entirely disintegrated and broken up into black dots. Mr. E. H. Man obtained a single specimen of this interesting species on Little Nicobar. Lethe gulnihal, n. sp. (Plate XXXIX. fig. 7, 6 •) Hah. Bhutan. Expanse. S 2-45 inches. Male. Upperside: both wings dark brown, with a distinct rich vinous gloss. Fore wing unmarked, except that the narrow discal band of the underside shows through paler on the upperside, the wing being somewhat paler beyond. Hind wing with the four middle ocelli of the underside showing through indistinctly. Underside : both wings dull brown without any vinous gloss. Fore wing with a short narrow ferruginous line across the middle of the cell, another similar one towards its end, the lower discocellular nervule marked with ferruginous ; a slightly outwardly-curved discal line from the subcostal nervure to just below the first median nervule, beyond which are four somewhat obscure small perfect ocelli placed between the nervules from the upper discoidal to the first median nervule ; a fine dark brown marginal line bordered on either side with paler and an anteciliary similar dark line. Hind wing with a pair of narrow ferruginous lines across the disk from the costal to the sub- median nervure, the outer one very irregular ; the lower disco- cellular nervule marked with ferruginous ; a series of six small distinct perfect ocelh composed of a black centre with a white pupil, a yellow, a black, and lastly a pale violet ring ; the upper ocellus out of line, placed furthest from the margin and the largest, the three lollowing subequal, the fifth a little larger, but not so large as the first, the sixth the smallest of all and geminate ; two fine dark marginal lines enclosing a fine ochreous line, the inner one. inwardly defined with pale lunules. The secondary sexual characters (" male marks ") of L. gulnihal are very peculiar and interesting. On the upperside of the hind wing it possesses the tuft of long black hair which is found in L. scanda, L. bhairava, L. latiaris, L. minerva, L. siha/a, and L. dyn- sate ; in addition it has a large oval patch of deep black lustrous scales, which is bounded above by the first subcostal nervule, partially inwardly and beneath by that portion of the subcostal nervure between the bases of the first subcostal and discoidal nervules, the patch not nearly reaching the margin (this feature occurs in L. bhairava in a somewhat modified form); lastly, the inner margin of the fore wing is deeply outwardly bowed, which is a unique feature, the bowed portion beneath the submedian nervure clothed with difi'erently formed and modified scales to those on the rest of the wing, this portion of the wing being clearly defined on 1887.] LITTLE-KNOWN BUTTERFLIES FROM INDIA, 451 the underside by a large deep black oval patcli. In L. bhairava there is a large patch of black scales on the inner margin of the fore wing ou the upperside, but it extends into the submedian inter- space. L. gulnihal is nearest to L. bhairava, its markings being very similar ; but it is smaller, and the bowed inner margin of the fore wing will at once distinguish the males of the two species. Two male specimens have been obtained in Bhutan by the native collectors of Messrs. Otto Mciller and A. V. Knyvett, in whose collections the specimens above described are deposited. Lethe brisanda. (Plate XXXIX. fig. 8, rS •) L. brisanda, de Niceville, Journ. A. S. B. vol. Iv. pt. ii. p. 249. n. 1, pi. xi. fig. U, female (1886). Hab. Bhutan. Expanse, rj 2'4 inches. Male. Differs from fresh native Sikkim specimens of L. dinarbas on the UNDERSIDE of the fure wing in the lilnc baud across the middle of the discoidal cell being much narrower, with uneven instead of straight edges ; the discal oblique band lilac throughout, straight, and of equal width throughout (in L. dinarbas the upper portion of the band only is lilac, this portion too in that species being wider) ; in having an additional ocellus in the subcostal inter- space (in a long series of L. dinarbas before me there are always three ocelli only). On the hind wing the discal bands are deep brown instead of ferruginous, and the lilac washings througiiout much more brilliant. Should the above-given characters prove constant in a large series of specimens, the males of L. brisanda and L. dinarbas will be easily distinguishable, the latter species being apparently very uniformly marked. The females are abundantly distinct. A single mide has been obtained near Buxa, Bhutan, by Mr. A. V. Knyvett's native collectors. Melanitis bethami, n. sp. Hab. Pachmarhi, Central Provinces, 3500 feet. Rainy-season form. Expanse. S 2'8, 5 3 inches, Male. Upperside: bothwings sooty hl&ck. Fore wing With the outer margin paler, the apex very slightly truncate, there being a small notch only below the lower discoidal nervule, a well-defined subcostal nearly round ochreous patch beyond the cell divided into three portions by the discoidal nervules. Hind wing unmarked. Underside : both wings paler than above, densely and evenly striated with darker, the outer margin ferruginous. Fore wing with four small obscure discal ocelli, of which the two in the second median and upper discoidal interspaces are the largest. Hind wing with a series of six submarginal ocelli, the second from the apex minute, the anal one geminated, the other four subequal and much 452 MR. L. DE MCivILLE ON NEW OR [May 3, larger. — Female. Upperside : both wings much paler than in tlie male. Fore icing slightly more truncate than in the male, with the v\hole apical thiid of the wing ochrcous, on which is placed a round black spot with a white centre in the second median interspace, with one suiall obsolete ocellus in the interspace below and three above it. Hind wing with the outer margin paler, a round black spot with a white centre in the first median interspace. Underside : both ivings with the ground-colour pale purplish, thickly striated with brown. Fore wing with an obscure ochreous fascia before the middle of the wing, a more prominent discal one from just beyond the middle of the costa directed towards the anal angle, which it does not reach ; the ocelli as in the male, but rather larger, the outer margin broadly ochreous. Hind wing with an outwardly curved discal ochreous fascia, its outer margin sharply defined ; the ocelli as in the male ; the outer margin broadly ochreous. Cilia throughout blackish in both sexes. Dry-season form. L\rpanse. S 3'3, $ 3'2 to 3"3 inches. Male. Upperside : both wings deeper black than in the rains form, the outer margins ashy. Fore iving highly falcate ; with a large rich ochreous, inwardly almost ferruginous, patch, wide at the costa, narrowed to a point at the first median nervule, outwardly banded by the ashy marginal area, and just extending into the apex of the cell. Hind wing unmarked, the tail much longer tlian in the rains form. Underside rery dark, very irregularly striated ; all the ocelli obsolete. Fore wing with a large wedge-shaped subapical pale violet patch, behind which are three obscure straight fasciae compored of pale mottles. Hind wing with the basal half of the wing much darker than the outer half, and sharply defintd, a patch of ochreous mottles at the end of the cell. The general character of the markings of the underside is similar to that of M. duryodana, Felder, and as they show but little variation in a long series of specimens, 1 have described them somewhat minutely. — Female. Upperside : both wings much paler than in the n.ale, and somewhat purplish, with no distinct outer ashy margin. Fore loing even more falcate than in the male, euorn ously more so than in the corresponding sex of the rains fcrm. 'J^ie apical two thirds rich ochreous, extending well into the disccichd cell, and reaching the anal angle, enclosing spots as in the female of the rains form, but which, however, so far as the two lower ones are concerned, are vaiiable in size, in one specimen having their black portions much lengthened inwardly ; the outer margin just mottled with ashy. Hind wingv,\{\\ the outer margin narrowly mottled with feriuginous and ashy, a small submarginal white dot in the first median inter- space (sometimes present in the male), sometimes with another smaller one in the interspace above. Underside : both wings with all the ocelli obsolete and much paler than in the male, being fer- ruginous-ochreous, fairly evenly covered with blotches of dark brown irrorations ; the fasciae as in the femi'Ie of the rains form, but blackish 1887.] LITTLE-KNOWN BUTTERFLIES FROM INDIA. 4.53 instead of ochreous. The markings of the underside of this sex, too, seem to be fairly constant in a number of specimens. My recent experiments proving by breeding from tlie egg that M. leda and M. ismene are but seasonal forms of one species, and the acquisition of both forms of M. bethami, has thrown a flood of light into my mind regardina; the Butterflies of this difficult genus. It mav now, I think, be accepted as au axiom that in all tropical and subtropical countries in which the year is divided into two well- marked seasons, a dry and a wet, the 3Ielanites that occur there will also have two well-marked forms, — a rains form, with slightly falcated fore wing, short tail to hind wing, and prominent ocelli on both wings on tlie underside; and a dry-season form, which has the fore wing highly falcate, a long tail to hind wing, and obsolete ocelli below. In the ' Lepidoptera of Ceylon,' in addition to M. leda and M. ismene (one species), Mr. Moore records only M. tainbra ; but in the Indian Museum, Calcutta, is a specimen marked by Mr. Moore himself " M. suyudana," which certainly differs from the form Mr. Moore has figured and described as M. tambra. I possess in all six specimens of this group from Ceylon, and though they present but slight variation in the size of the ocelH (it should be remembered that Ceylon has a very equable chmate throughout the year), there is a well-marked difference in outline, what I should call M. tambra being the rains form, and M. suyudana the dry- season form. In Sikkim we have, besides M. leda and M, isinene, M. zitenius, of which Herbst has figured the dry-season form and Mr. Distant^ the rains form ; and M. aswa, Moore, the strongly occllated rains form, and M. bela, Moore, and M. duryodana (tiie two latter I now believe to be but varieties of one form), the dry- season form of a third species. An intimate knowledge of the species of Melanitis occurring in other parts of the Old World, to be obtained only by living amongst them and carefully noting their different forms and the seasons when they occur, and by breeding them from the egg, would, I feel sure, reveal the fact of the seasonal dimorphism which occurs in all the species of the genus, and I trust that notice being now drawn to the subject, collectors and entomologists will devote attention to it. To return to M. bethami, I have described the rains form from a pair taken on the 8th August, and the dry-season form from five pairs taken between the 15th and 27th October at Pachmarhi, by Mr. J. A. Betham, after whom I have much pleasure in naming the species. CyLLOGENKS JANETS, U. Sp. Hab. Bhutan. Expanse. S 3"5, $ 3'55 inches. Male. Upperside : both wings deep dull brown, almost black. Fore wing with a broad rich ochreous curved subapical band, attenuated towards the anal angle, which it hardly reaches, the rich ^ Rbopalocera Malayuua, p. 41i!. ii. 3, pi. xxx\iii. fig. 2,(5 (1886). 454 MR. L. DE NIC^VILLE ON NEW OR [May 3, ochreous colour extending along the first median nervule for a short distance ; cilia black. Hind vnng with the outer margin somewhat broadly ochreous, sprinkled with dark-brown or blackish irrorations ; cilia black, tipped with white on the three upper indentations between the veins. Underside : both wings very variegated, the ground-colour apparently being ochreous, thickly irrorated with dark-brown striae ; a discal obscure purplish fascia, inwardly bounded by a dark line. Fore wing with three short subcostal bands reaching the middle of the cell, and a broader subapical one, ochreous (these fasciae are formed by those portions of the ground being free from irrorations), an obscure purplish patch at the apex, an irregular series of five round violet-white spots placed between the veins mid- way between the cell and the outer margin, the two in the median interspaces large, the other three very small. Hind wing with the inner edge of the black discal line marked with an ochreous bar at the costa, the outer margin showing more of the ochreous ground- colour than the rest of the wing ; five discal viulet-white spots between the veins, the three lower ones large, the two upper small, the lower ones surrounded by a black ring, the spot in the first median interspace the largest of all. — Female. Upperside : fore wing with the subapical band broader, richer-coloured, and extending along the costa ; the black apical patch crossed by yellow veins; the r.iedian nervules also marked with yellow near the band. Hind wing with the outer margin richer ochreous. Underside much paler ; the darker irrorations far less dense ; no trace of the diffused pur- plish fasciae. I have placed this species somewhat doubtfully in the genus Cyllogenes, the chief recorded structural character of which is the presence of a large deep black patch in the male, this " sexual mark " being entirely absent in my species, the sexes being prac- tically marked alike. C. janetce is, however, much more closely allied to C. suradeva, Moore (hitherto the type and only species in the genus), than to any other described species, the yellow band on the fore wing and the upperside being a striking feature, which is common to both species. On the underside the blind ocellated spots are precisely similar in both species ; and the purj)lish fasciae on the underside of the male of C. janeta is of the same tint as obtains on the upperside of both sexes of C. suradeva. The truncation of the apex of the fore wing in both sexes of C. janetce (more especially in the male) is a good structural character by which to separate the two species in both sexes. It agrees also structurally with Cyllogenes suradeva in the almost similar extraordinary character which obtains also in the genus Parantirrhcea of Wood-Mason, and which for Cyllogenes has not hitherto been noticed, viz. : in the male the three median nervules of the fore wing are considerably further apart at their apices than in the female, owing to the fact that the lower one has to supply the place normally taken by the submedian nervure, as it reaches the outer margin but a very short distance anterior to the anal angle ; while the submedian nervure is very short, slightly sinuous, and reaches the inner margin at considerably less than half 1887.] LITTLE-KNOWN BUTTERFLIES FROM INDIA. 435 the length of the margia from the base. In the female, however, the median riervules and submedian nervure are quite normal, the neuration being very similar to that of Melanitis. Described from two male examples in the collection of Mr. A. V. Knyvett (after whose wife I have named it), and a female in that of Mr'. Otto Moller. EuTHALIA DUDA. E. duda, Staudinger, Ex. Schmett. part i. p. 152, pi. liii., male (1886). Hah. Near Buxa, Bhutan ; Sikkim. Expanse. $ 4*5 inches. Female. Upperside : fore wing differs from the same sex of E. durga, Moore, which also occurs in the same locality, in the four white spots between the veins beyond the discoidal cell being much smaller, their inner ends excavated ; the lowest one in the lower discoidal interspace the smallest and most deeply excavated, and shilted outwards, thus breaking the line of the spots ; the four spots from the third median nervule to the inner margin also much smaller, the two lower ones with their outer edges highly diffused instead of ha\ing them sharply defined. Hind wing with the discal white band narrower, its outer edge not defined with a line of the ground- colour as in E. dvrga ; its inner edge much more even ; the diffused fascia immediately beyond it bluish purple instead of green. Un- derside : fore wing with the black increasing submarginal band springing posteriorlv from the two apical white spots twice as wide ; the discal macular band on both tvings as on the upperside. A single female of this quite distinct species was obtained by Mr. A. V. Knyvett's native collectors near Buxa. Since my description was written, I have received Dr. O. Staudiu- ger's work quoted above ; and from it I learn that he has described this species from two male specimens from Sikkim, collected many years ago by the late W. S. Atkinson. Catapcecilma delicatum, n. sp. C. bubases, de Niceville (nee Hewitson), Journ. A. S. B. vol. liv. pt. ii. p. 118, pi. ii. figs. 11, male, \, female (1885). Through the kindness of Mr. W. L. Distant in sending me an advance copy of the figure of the true C. bubases, Hewitson, which is shortly to appear in the final part of his ' Rhopalocera Malayana,' I am at once enabled to see that Hewitson's description of his species does not apply to the species which occurs in Sikkim, but to which it is somewhat closely allied. "When redescribing and figuring this species I was very uncertain whether or no to give it a name, and wrote: — " Hewitson's description of C. bubases is very meagre ; a comparison of his Malaccan female type with Sikkim specimens may disclose specific differences." This latter conjecture turns out to be a correct one, so I name the Sikkim species C. delicatum. 4:)6 MR. L. DE NicfiviLLE ON NEW OR [May 3, Cheritrella, nov. gen. Fore wing with the costal marc/in slightly and regularly arched ; the outer margin highly truncated from the apex to the termination of the third median nervule, this truncated portion, moreover, being concave ; below the third median nervule to the inner angle the margin is nearly straight and oblique ; inne?' margin straight. Gostal nervure very short, not nearly reaching to opposite the apex of the cell ; the first, second, and third subcostal nervules at regular dis- tances apart before the origin of the upper discocellular nervule, the fourth springing from the third about the middle of its length ; upper discocellular nervule directed outwards, middle and loiver upright and concave ; the bases of the second and third median nervules half the distance apart of the bases of the second and first ; submedian nervure straight. Hind wing with the costal margin strongly arched at the base, thence to apex nearly straight ; outer margin to base of long tail at termination of first median nervule straight but waved from thence to anal lobe at right angles ; abdo- minal margin strongly convex at base and highly excavated above the anal lobe ; a short tail at termination of submedian nervure. Costal nervure strongly arched at base, afterwards nearly straight ; Jirst subcostal nervule originating some little distance before apex of cell, nearly straight ; discocellular nervules of nearly equal length, concave, outwardly oblique, the origin of the third median nervule at the lower end of the cell, the second just before its end, that of the first being fully four times as distant from that of the second as is the latter from the third ; submedian nervure nearly straight ; internal nervure very short, ending on the abdominal margin above the deep excavation, and highly sinuous. Nearest to Tichei-ra, with which it agrees in having no secondary sexual characters in the male, in the neuration of the hind wing ; also in the length and position of the tails, the anal lobe, and the deep excavation above it, but differs from it in the truncation of the fore wing, the costal nervure terminating long before the apex of the cell, and the fourth subcostal nervule spiinging from the third about its middle instead of considerably nearer the apex. Cheritrella TRUNCirENNis, n.sp. (Plate XXXIX. figs. 4, c? ; 3,?.) Hah. Sikkim. Expanse.