Systematics of the Bufo coccifer complex (Anura: Bufonidae) of Mesoamerica

] ds>^-Scientific Papers Natural History Museum The University of Kansas 31 Mcirch 2003 Number 38:1-27 Systematics of the Bufo coccifer Complex (Anura: Bufonidae) of Mesoamerica ubrary Joseph R. Mendelson III', Bhck^ L. Williams-, Christoi'hek A. Sheil\ ANL^ Daniel G. Mulcahv* -^m ' Dcimiiiiieiit of Ucrpctolo^ii, Zoo Atlniitu, S(H) Cherokee Ave SE, Atlanta, GA j03]5 H '-'•^y^-rrY -Department ofhite^rative Biology, University of California, Berkeley, CA 94720-3140 V. ''c-^^^ ^Department of Biolo<;y, John Carroll Universiti/, University Hei;^hti, OH 4411S ^Department of Biology, Utah State Universiti/, Logan, UT S4322-5305 CONTENTS ABSTRACT 2 RESUMEN 2 INTRODUCTION 2 Al K\'OVVLEDGMENTS 5 MATERIALS AND METHODS 5 Morphological Da la 5 Molfxular Data 6 ACCOUNTS OF SPECIES 6 Biifo coccifer 6 Bufo cycladen 11 Bufo ibarrai 13 Bufo pisiinius 15 Bufo porteri 17 Bufo signifcr 20 MORPHOMETRIC ANALYSES 22 MOLECULAR ANALYSES 23 LITERATURE CITED 24 APPENDIX 26 © Natiiivil History Museum, The University of Kjnsds EmS^ MtiVf LF'TarV '^^^ ^'^' ' "^'■*""''*^2 Museum of Compara*ye Zoo'o^ Harvard Univ6rs[t>'