INTRODUCTION The West Indies lie in an arc extending some 1,800 miles long from the Serpent's Mouth (between South America and Trinidad) to the Yucatan Channel (between Cuba and Mexico) on the west, and the Straits of Florida (between Cuba and Florida) on the north. They are comprised primarily of three island groups. These are the Bahamas on the northeast, the Greater Antilles southward, and the Lesser Antilles on the southeast (fig. 1). The Bahamas are flat whereas the Antilles are relatively mountainous. Coral reefs dot the eastern rim of this island chain, especially among the Bahamas, where new land is forming. There is a great differential in off-land depth of the seas. Thus in the Gulf of Batabano, between Cuba and the Isle of Pines, the water is so shallow that the bottom may be seen; off the northwest coast of Puerto Rico there is an abyss, the Milwaukee Deep, of some 9,220 m. A more extensive account can be found in Roberts (1940). Several important physical and ecological factors probably have exerted an important influence on the dispersals and/or population sizes of species of pselaphid beetles in the past and continuing into the present. Such factors include (1) the possibility of land bridges, short and long. If such existed in the historical past they could be utilized by population fragments walking to their present locations. Such movements would be slow, from one available habitat niche to another; random as to direction; presupposing long intervals of time. (2) The Antillean marine currents may have been important. Such currents include (a) the Caribbean Current westward from the Atlantic Ocean, through numerous passages between the islands of the Lesser Antilles; (b) Florida Current, a warm marine river, moving northwest and north at about 4 miles an hour through the Straits of Florida; (c) the Antilles Current which flows northwestward on the Atlantic side of the Bahamas. Such currents, and other minor ones, may have contributed pselaphids from one island to another, or from the Spanish Main to an island, by rafting. 1 2 FIELDIANA: ZOOLOGY. VOLUME 68 Such dispersal by natural rafts could have happened through geological time, at least over the last 40 million years (Oligocene) into the present. Since pselaphids are predators, and of relatively small size, quite a few could survive on a small floating mass of debris if the latter contained such organisms as small fly larvae, mites, or Collembola. General ecology, including foods, has been summarized by Park (1964, 1965). In recent times such possible dispersals by rafting probably would be west and northwest from South America on the Caribbean Current and the Antilles Current, and northeast from Yucatan toward Florida and Cuba. Both the Florida Current and Antilles Current would tend to converge near the Bahamas and extreme southern peninsular Florida, forming the Gulf Stream. (3) Hurricanes. Even this name is West Indian, named for the Carib god, Huracan. On the average about 14 hurricanes occur between late June and early November, several of which are severe each year. These storms originate in the Atlantic just north of the equator, wax in wind velocity from 75 to 150 miles per hour and are accompanied by heavy rains. They work an erratic course through the Caribbean region and may blow small arthropods into new habitats. Pselaphidae are not usually regarded as strong fliers and any individuals exposed, or air-borne during such a period would be available for automatic transfer, possibly for many miles. Additionally, the general wind pattern might disperse some of these beetles. Such winds are the northeast trades which are preponderantly northeast in January and west in July. Such recurrent winds in recent times could be subtractive rather than additive for pselaphids of the West Indies, except with the proviso that there would be a chance of wind- transport from the Leeward Islands westward, and from Puerto Rico to Hispaniola, Jamaica, and Cuba, and from the Bahamas toward extreme southern Florida and Cuba. (4) Volcanic action and earthquakes. The Antilles south of the Tropic of Cancer have been and probably will continue to be subject to earthquakes. Furthermore, Mont Pelee of Martinique and the closely associated peaks of nearby islands should be noted here for volcanic disturbances. These latter are sporadic but may be locally important in disturbance of pselaphid populations. (5) Activities of man. As the human population increases in size direct and indirect effects on the local pselaphid populations are PARK ET AL.: PSELAPHID BEETLES 3 probable. Such effects would be generally adverse since pselaphids are primarily inhabitants of top-soil and floor molds. Urban and suburban sprawl, agriculture, and industry would tend to have a subtractive force, and lead to reduced populations or local extinction of these beetles. This would be hastened where soil erosion was involved. An exception might be the multiplication by man of a suitable pselaphid habitat through unconscious co- operation, e.g., in the West Indies where banana plantations are operated, fallen banana leaves tend to accumulate a variety of pselaphid genera. The West Indian pselaphid fauna is poorly known. The present report adds to our information but also serves to bring up to date the extent of our ignorance. A preliminary analysis was included in a general Neotropical study (Park, 1942). In later reports the author has received a loan of pselaphids from South Bimini Island in the Bahamas through the courtesy of Dr. Mont A. Cazier, American Museum of Natural History (Park, 1954a); a gift of pselaphids from Jamaica by Dr. Peter F. Bellinger (Park, 1955); additional pselaphids from Jamaica were given by Dr. Bellinger and Dr. C. Clayton Hoff (Park, 1960). Other material has been received and has added to our information. Much of this is being reported upon here. It includes the gift of a small but choice collection by Dr. Glenn E. Haas, United States Public Health Service, from Puerto Rico; a gift of a larger collection from numerous areas of the Antilles by the Rev. E. J. Pearce, of the Barbados; the loan of material from several West Indian islands through the courtesy of Dr. Milton W. Sanderson, Illinois Natural History Survey; and, finally, the loan of a varied collection from several Antillean localities through the courtesy of Dr. P. J. Darlington, Museum of Comparative Zoology, Harvard University. In this latter instance an unique specimen, representing a new genus and species from Hispaniola, has been described previously (Park, 1954a). The author thanks Dr. Leland R. Pederson of Northwestern University for a painstaking review of the introduction of this paper and he thanks Barbara Verrier for assistance in the preparation of material and illustrations. (The editors thank Mrs. Grace Finger and Mrs. Robyn Gardner for bibliographic and indexing help). SYSTEMATIC TREATMENT The following keys are designed for the discrimination of genera and species and all higher taxa known to inhabit the West Indies. They will not apply necessarily to other Neotropical regions. Full references to previously described species, and West Indian type species, will be found in the checklist and references. All measurements for new species descriptions are in miUimeters. Key TOTHK Subfamilies OF West Indian Pselaphioae 1 Three-segmented tarsi, of which the first two segments are relatively very short, and the third (distal) segment is relatively long 2 Three-segmented tarsi, of which the first segment is very short, and the last two segments are relatively very long PSELAPHINAE 2 (1) Abdomen with five free visible tergites FARONINAE Abdomen with dorsal surface composed largely of a single disc which is formed by the fusion of the first three visible tergites. CLAVIGERINAE Subfamily FARONINAE Faronitae Jeannel, 1949, Paris Mus. Nat. d'Hist. Nat. N. S., 29 (1), p. 15. Key to West Indian Tribes of FARONINAE Antennae 11-segmented FARONINI Antennae nine-segmented PYXIDICERINI Tribe FARONINI Faronini Raffray, 1890a. Rev. d'Entomol., 9, p. 82. Genus Caccoplectus Sharp' Caccoplectus Sharp. 1887, Biol. Centr.-Amer., 2 (1), p. 22. Type-species: Caccoplectus celatus Sharp, loc. cit., p. 22, Guatemala. 'Donald S. Chandler has revised Caccoplectus, removed the genus from Faronini. and placed it back into Holozodini of Raffray. Chandler, D. S., A Revision of the Genus Caccoplectus (Coleoptera: P.selaphidae). Coleopterists Bull., 29 (4), Jan. 1976, pp. 301-316. PARK ET AL.: PSELAPHID BEETLES 5 Caccoplectus bellingeri Park, 1955, Bull. Chicago Acad. Sci., 10 (7), p. 101. Type (male?) beaten from bushes on Morce's Gap Trail, St. Andrew Parish, Jamaica, between 1,500 and 5,(X)0 ft. elevation. Tribe PYXIDICERINI Pyxidicerini Raffray, 1903, Ann. Soc. Entomol. France, 74, p. 6. Genus Bythinoplectus Reitter Bythinoplectus Reitter, 1883, Deut. Entomol. Z., 27, p. 37. Type-species: Bythinoplectus foveatus Reitter, loc. cit., p. 37, St. Thomas, Virgin Islands. Key to .species of Bythinoplectus 1 Lateral pronotal margins rounded, without distinct apical or ba.sal angles. bahamicus Park Lateral pronotal margins with distinct apical angles 2 2 (1) Pronotal margins suddenly produced as an obtu.se, lobular angle at apical three-fourths of length acutangulus Raffray Pronotal apical angles distinct and the lateral margins thence almost obliquely straight to ba.se 3 3 (2) Pronotum much longer than wide foveatus Reitter Pronotum with length and width subequal laminatus Park, n. sp. Bythinoplectus laminatus Park, new species Description. — Type male: Head 0.20 long x 0.34 wide; pronotum 0.24 x 0.24; elytra 0.33 X 0.41; abdomen 0.46 X 0.38; total length 1.24. Cuticula light yellowish brown (apparently teneral); head and pronotum granulate-punctate, elytra finely punctulate, abdomen subimpunctate; pubescence sparse on head and pronotum, short and appressed; abdomen with longer setae. Head with the usual median rastrum of the genus, transversely divided into a shorter frontal, and a longer vertexal portion; a pair of minute vertexal foveae, inconspicuously placed on either side of the rostral base; eyes large; tempora short and much wider than long; face deeply excavated to form the palpal fos.sae. These two fos-sae separated by a thin clypeo-frontal lamina. This lamina bears an oval, translucent window at center of base, and is erected ventrally into a bidentate process, each tooth of which bears an arcuate seta. Maxillary palpi as for genus. Convex gula with basal fovea. Antennae 9-segmented as for genus; 1 large and granulate-punctate with dorsal face flattened and ventral face concave; 2 smaller, truncate-obconical; .3-7 .subequally wide, transverse and very small; 3 briefly obconical in shape; 4-7 transver.se- moniliform; 8 slightly larger, tran.sversely obtrapezoidal; 9 large, as wide as 1, with truncate base and subovate outline, this segment bearing con.spicuous special setae. 6 FIELDIANA: ZOOLOGY. VOLUME 68 Pronotum with distinct but rounded anterior angles, and with the lateral margins thence obliquely straight to base. A nude lateral fovea near margin at basal two-fifths; a broadly subcircular nude fovea at discal center; base with a pair of narrowly separated, transversely oval nude foveae. Elytra with rounded humeri; each elytron bifoveate near base, the sutural fovea at origin of an entire sutural stria, discal fovea at origin of a short, broad impression extending to basal fourth; flank lacking subhumeral fovea. Abdomen with five visible tergites, the first four with strong lateral margins; first four tergites progressively slightly longer posteriorly; last tergite semicircular, about as long as second. Basal abdominal carinae present on first three tergites, separated by about 46 per cent of total segmental width and about 18 per cent of segmental length. Six visible stemites; first short; second to fourth progressively slightly longer posteriorly; fifth shorter than fourth, with an inconspicuous transverse elevation at center near base; sixth about as long as first, with a small, inconspicuous depression and posterior margin medianly concave. Legs slender, simple; three-segmented tarsi with first tarsomere short and minute; second and third tarsomeres subequally long, the second slightly inflated and third bearing a single tarsal claw. This species is described from one specimen, the male type, in the author's collection. It was not dissected for the aedeagus as it appears teneral, and if so the copulatory apparatus might be unsuitable for study. It was collected by Rev. E. J. Pearce on the forest floor near Pembroke, Tobago on April 22, 1965. It is most closely comparable to the type species, Bythinoplectus foveatus Reitter, 1883, also based on one specimen from St. Thomas, Virgin Islands, but the latter differs markedly in body proportions, the pronotum being described as much longer than wide and the elytra only shghtly wider than pronotum. Subfamily PSELAPHINAE (Reorganized by Raffray. 1903. 1904; Park, 1942, 1951; Jeannel, 1949, 1950) In the pselaphine tribe Brachyglutini the genus Bythinogaster has such narrow margins on the sides of the first three visible tergites that these latter may be overlooked, and the abdomen appear marginless. This genus can be separated from West Indian Tanypleurini by the penultimate segment of the maxillary palpi. This segment is longitudinally lenticular and bears a conspicuous pad of short pubescence on its external face. Key to West Indian Trihesof PSELAPHINAE 1 All trochanters short, with the femora inserted very obliquely on the trochanters, so that each femur is quite near its associated coxa 2 PARK ET AL.: PSELAPHID BEETLES 7 Mesotrochanters and metatrochanters long and more or less inflated distally (clubbed), with femora inserted on the distal faces of trochanters, so that each femur is distant from its associated coxa 5 2 (1) Mentum wide and largely covering the mouth and mouth-parts; cardo of each maxilla extended obliquely as an acute external projection. JUBININI Mentum small, not covering mouth and mouth-parts; maxillary cardo not so formed 3 3 (2) First three visible tergites with lateral margins 4 Abdomen without margins, or with the first visible tergite bearing a longitudinal carina on each side TANYPLEURINI 4 (3) Venter with six or seven sternites fully visible EUPLECTINI Venter with only five sternites fully visible from side to side. BRACHYGLUTINI 5 (1) Last segment of maxillary palpi wider than long, and with a slender palpal cone at apex and a distinctive appendage on its external face; general body pubescence generally in the form of flattened scales. CTENISTINI Last segment of maxillary palpi otherwise formed 6 6 (5) Last segment of maxillary palpi elongate ovoidal and apically acute, with short stiff setae and a minute apical palpal cone TYRINI Last segment of maxillary palpi with the internal face occupied in whole or in part by a broad, entire sulcus or sulcoid impression. HAMOTINI Tribe JUBININI Jubinini Raffray, 1903, Ann. Soc. Entomol. France, 74, p. 25; Genera Insectorum, 64, p. 25. Jubus and its allied genera (cf. Park, 1952a, p. 14) have the V- shaped or Y-shaped carinal pattern on the ventral surface of the head strongly formed. Balega has the ventral surface of the head bearing a median, longitudinal sulcoid impression and a pair of shorter, obliquely converging sulcoid impressions. Such impressions may be weakly formed. Between these three impressions the cuticula may be raised in weakly developed folds so that specimens of Balega may be incorrectly placed in Jubus unless this matter is realized. Stratus has the ventral surface of the head adorned more simply, with either a single median longitudinal sulcoid impression, or a median fovea. 8 FIELDIANA: ZOOLOGY, VOLUME 68 Key TO Genera of Jubinini 1 Ventral surface of head bearing a pair of sharply defined obliquely converging carinae which unite posteriorly to form a V-shaped or Y- shaped structure Jubus Schaufuss Ventral surface of head lacking this carinal 2 2(1) Pronotal disc bisected by a strongly developed median, longitudinal sulcus Balega Reitter Pronotal disc lacking a median longitudinal sulcus Stratus Schaufuss Genus Balega Reitter Balega Reitter, 1883, Deut. Entomol. Z., 27, p. 43. Type-species: Balega elegans Reitter, loc. cit, p. 43, St. Thomas, Virgin Islands. Key to Specie.s of Balega 1 Head obviously elongate, about V/z times longer than wide, in a length- width ratio of 5/3.4 (length from labral suture to occiput, and width, the widest point behind the eyes) longiceps Park, n. sp. Head much shorter relatively, from as long as wide to not more than a length-width ratio of 3.6/3 2 2 (1) Vertexal sulci very narrow, about the diameter of the vertexal foveae. elegans Reitter Vertexal sulci posteriorly much broader than vertexal foveae, and enclosing them 3 3 (2) Eyes large, subreniform, of more than 30 coarse ocular facets; antennal segment 10 distinctly longer than 11 male intermedia Park Eyes rudimentary, of two ocular facets; antennal segments 8, 9, and 10 slightly transverse and subequal female intermedia Park Balega longiceps Park, new species Description. — Type male: Head 0.43 long x 0.31 wide; pronotum 0.43 X 0.41; elytra 0.67 x 0.69; abdomen 0.55 x 0.67; total length 2.1. Cuticula bright yellowish brown; head subimpunctate; pronotum micro- granulate; elytra micropunctulate; abdomen subimpunctate with a few micro- granules; pubescence conspicuous, long and hyaline. Head elongate-trapezoidal; subreniform eyes of more than 30 coarse ocular facets, the eyes flattened and occupying the median fifth of head length; tempora elongate, gently rounded into cervicum, about one-third longer than eyes; two nude vertexal foveae on a line passing through posterior margins of eyes, these foveae small, about the size of an ocular facet. A long glabrous vertexal sulcus extends anteriorly from a point behind and surrounding each vertexal fovea; these two sulci unite distally to form a short broader median sulcus which bisects the frontal rostrum PARK ET AL.: PSELAPHID BEETLES 9 (the three sulci form a furculoid impression). Occiput bearing two blackened cusps. Front occupied largely by the acetabulae of the first antennomeres, these latter being separated medianly by a subvertical frontal strap which in turn unites with the simple clypeus. Labrum transverse with a deeply concave distal margin; maxillary palpi as for genus. Ventral surface of head bisected by a longitudinal sulcoid impression which deepens anteriorly, and bears a small elongate-oval gular fovea near center; on either side of the anterior part of this impression is a short, broad sulcoid impression; these two short oblique impressions converge upon, but do not join the median impression. Antennae long, about .72 mm.; 11-segmented with an inconspicuous club of the last four segments; 8-10 subequal in width and length; 11 nearly as long as the preceding three united, base truncate and apex rounded. Pronotum with a lateral fovea at each side at basal fifth; extending forward from each fovea is a broad arcuate impression to about center of lateral margin; lateral foveae connected by a transverse sulcus at basal fifth; pronotal disc bisected by a longitudinal sulcus from apical fifth to transverse sulcus; a longitudinal carina extends from center of transverse sulcus to bisect basal margin; an acute spine on lateral margin opposite each lateral fovea. Elytra with rounded humeri; each elytron bifoveate; sutural fovea at basal origin of entire sutural stria, and discal fovea at origin of poorly defined impression through basal third; these foveae more or less united by a biarcuate ridge which is more defined over each fovea; flank with a subhumeral fovea at origin of an entire longitudinal sulcoid impression, this latter bounded laterally by a carina which arches over the foveal rim. Abdomen with five visible tergites, the four of which have broad, well-defined margins; first three subequally long and fourth distinctly longer than third tergite; fifth short and subvertical. Six sternites of which the first is short and inconspicuous, second and third subequally long, fourth shorter than third, fifth medianly shorter than fourth, sixth about as long as fourth. Legs long and slender, relatively simple; each tarsus with a pair of claws, the lateral claw relatively large, the mesial claw slender, shorter, and set at an angle. This new species is based on the unique male type, deposited at the Museum of Comparative Zoology. Collected by Dr. P. J. Darlington, Jr. at about 6,000 ft. elevation, near Loma Vieja, S. Constanza, Dominican Republic in August, 1936. It is set apart from other known West Indian species of the genus as a consequence of its very long head. By virtue of its relatively broad vertexal sulci it is most closely allied so far to Balega intermedia Park, 1960. Balega intermedia Park, 1960 Balega intermedia Park, 1960, Trans. Amer. Microscop. Soc, 79 (1), p. 5. This species was erected on an unique female type, collected at Dolphin Head, Hanover Parish, Jamaica. A male of this species was 10 FIELDIANA: ZOOLOGY. VOLUME 68 in material loaned to the author, and will be deposited in the Museum of Comparative Zoology. It was collected by Dr. P. J. Darlington, Jr. at Whitfield Hall, Blue Mountains, Jamaica near 4,500 ft. elevation, August 13-20, 1934. The male differs from the female in the following particulars: (1) eyes are large, subreniform, about as long as the tempora in lateral view, and composed of about 32 coarse ocular facets; (2) antennal segment 10 distinctly longer than 9; (3) legs more robust than in the female, the femora especially more inflated. Genus Stratus Schaufuss Stratus Schaufuss, 1872, Nunquam Otiosus, 2, pp. 246, 452. Type-species: Stratus ursinus Schaufuss, loc. cit, p. 452. Key to Species of Stratus Pronotum with a conspicuous antebasal spinoid tooth each side; size relatively large (2.9 mm.) dominicanus Park, n. sp. These pronotal teeth absent; size relatively small (1.5 mm.) hirsutus Park, n. sp. Stratus dominicanus Park, new species Description. — Type male: Head 0.5 long x 0.38 wide; pronotum 0.52 x 0.55 (just apical of lateral spines); elytra 0.91 x 1.00; abdomen 0.95 x 0.9; total length 2.9. Cuticula shining reddish brown; head sparsely punctulate, pronotum sparsely punctate, elytra and abdomen subimpunctate; pubescence flavous, suberect and arcuate on tempora and base of pronotum but tending to become prostrate on elytra and abdomen, setae forming an obvious fringe on posterior elytral margins and tergites. Head with a pair of large, coarsely faceted eyes of about 36 facets, these eyes hirsute with short, bristling setae between the facets; tempora rounded and almost twice as long as eyes; a pair of nude vertexal foveae, each with the diameter of more than an ocular facet, mutually closer than either to its adjacent eye; an arcuate vertexal sulcus extending from each fovea and uniting anteriorly in a common rostral sulcus (this furculoid design is deep and the sulci slightly wider than a vertexal fovea); occiput with a pair of median cusps, these cusps separated by a broad median impression which narrows distally to a point on a line passing through posterior eye margins; ventral surface of head with a weakly developed median sulcal impression which is widened at each end and bears a circular submental fovea and a sharply defined, elongate gular fovea; maxillary palpi as for genus. Antennae 11-segmented; segments 1-7 elongate; poorly defined club of last four segments; 8 and 9 with length and width subequal; 10 slightly longer; 11 elongate, about one-third longer than 9 and 10 united (these comparisons do not include the conspicuous antennal articulations). PARK ET AL.: PSELAPHID BEETLES 11 Pronotum with unmarked disc; a large foveoid impression on either side at basal two-fifths, these connected by a transverse antebasal sulcus; basal two-fifths medianly interrupted by a carina from floor of sulcus to basal margin; a strong acute tooth at each side foveoid impression. Elytron bifoveate, the basal rim of each fovea carinoid; entire sutural stria from mesial fovea, a vague discal impression for basal fifth of elytral length; flank with subhumeral fovea which is unusually posteriad in position at basal two-fifths of elytral length and a broad and deep longitudinal sulcus from fovea. Metathoracic wings present. Abdomen with five tergites in a median length ratio of 2.0/2.6/2.0/2.4/1.4 and first four with broad lateral margins; six visible sternites of which the first is very short and the second to sixth in a median length ratio of 2.0/2.4/2.0/1.2/1.6 and the venter simple. Legs as described for Balega longiceps Park, page 9. Aedeagus 0.36 long and 0.5 deep (including the long curved distal tube). This organ dissected from a paratype (fig. 2). Female: as described for male with the following differences: (1) small eyes of five large facets, these set in a sharply defined oval impression; (2) antennal segment 8 longer than wide and 11 about as long as 9 and 10 united; (3) tergite ratio 3.0/3.0/2.6/2.0/1.4 and stemite ratio of second to sixth 3.0/2.4/2.0/1.2/0.6 and simple. This new species is based on a series of five specimens (three males and two females). Type male and half of the paratypes are deposited in the Museum of Comparative Zoology and the other paratypes are retained in the author's collection. The series collected by Dr. P. J. Darlington, Jr. in cloud forest near Valle Nuevo, Dominican Republic in August, 1938 at about 6,000 ft. elevation. The most closely related ally is Stratus hirsutus but the latter is quickly separated by the characters noted in the preceding key. Stratus hirsutus Park, new species Description. - Type male: Head 0.25 long X 0.24 wide; pronotum 0.31 X 0.32; elytra 0.49 x 0.51; abdomen 0.49 x 0.47; total length 1.5. Cuticula shining reddish brown; subimpunctate except outer walls of vertexal sulci and submental area which are microgranulate; pubescence of head and pronotum relatively long, bristling and shaggy; on elytra and abdomen the setae are shorter and more appressed. Head with a pair of flattened, coarsely faceted eyes of about 30 facets; vertexal foveae and sulci as noted for Stratus dominicanus; base of head with a pair of occipital cusps and a relatively deep median sulcus extending between these cusps from occipital margin nearly to a line through posterior eye margins; ventral surface of head with a deep median sulcus which broadens both apically and basally and bears a basal fovea. Antennae with poorly differentiated club of last four segments; 8-10 subequal; 11 twice as long as wide. 12 FIELDIANA: ZOOLOGY, VOLUME 68 Pronotum as for dominicanus except that the strong juxtafoveal teeth of the latter are obsolete in the present species. Elytra as for dominicanus. Abdomen with five visible tergites in median length ratio of 2.0/1.3/1.1/1.0/0.05 and first four with broad lateral margins; six visible sternites in median length ratio of 0.3/1.3/1.3/1.0/0.05/0.05 and the venter simple. Legs as for dominicanus except that the accessory tarsal claw is more arcuate. Aedeagus 0.09 long and 0.19 deep (including the long, biarcuate tube). Usually in the case of a species based on an unique male I have not dissected for the aedeagus. In this case the type was mounted on a point in two pieces, barely held together by the articulating membranes, so the above description was taken from the point- mount and then dissected and preserved on a slide (fig. 3). Based on the male type, deposited at the Museum of Comparative Zoology and collected by Dr. P. J. Darlington, Jr. in August, 1938 at Costanza, Dominican Republic at between 3,000 and 4,000 ft. elevation. Its nearest ally is dominicanus. The aedeagi of dominicanus (fig. 2) and hirsutus (fig. 3) are similar generically, attesting to their obvious relationship, but these organs are very distinctive at the species level. Genus Jubus Schaufuss Jubus Schaufuss, 1872, Nunquam Otiosus, 2, p. 455. Type-species: Jubus spinicollis Schaufuss, loc. cit. p. 456, Santa Fe de Bogota, Colombia. This genus was revised by Raffray, 1893, and Park, 1952. Key to Species of Jubus Vertexal sulci very narrow and the vertexal foveae not wider than the sulci, so that the sulci appear to arise de novo from the vertex insularis Raffray Vertexal sulci broad; vertexal foveae large and pubescent and lie in a common excavation between the eyes clavatus Raffray Tribe TANYPLEURINI Tanypleurini Jeannel, 1949, Mem. Mus. Nat. d'Hist. Nat., N. S., 29, p. 79. Key to Subtribes of Tanypleurini Elytron with two, three, or four antebasal foveae DALMODIINA Elytron with antebasal foveae absent TANYPLEURINA Subtribe DALMODIINA Dalmodiina Park, 1951, GeoL Surv. Alabama, Mus. Paper 31, p. 61. PARK ET AL.: PSELAPHID BEETLES 13 Figs. 2-4. 2. Stratus dominicanus n. sp., aedeagus, left lateral face of paratype male. 3. Stratus hirsutus n. sp., aedeagus, left lateral face of type male. 4. Insulomodes tobagoensis n. g. and sp., aedeagus, ventral face of paratype male. Key to Genera of Dalmodiina 1 Elytron with four antebasal fovea Insulomodes Park, n. g. Elytron with two antebasal fovea 2 2 (1) Pronotal disc with a strong median longitudinal carina. Euharmophola Park Pronotal disc simply convex Dalmonexus Park Insulomodes Park, new genus Tanypleurini differing from other genera of this tribe in having in common the following features: (1) well-developed eyes in both sexes and a pair of vertexal foveae; (2) antennae 11-segmented, distantly articulated, and not geniculate; (3) 4-segmented maxillary palpi; segment 1 minute; 2 elongate, arcuate, pedunculate; 3 short, rounded-triangular; 4 largest, elongate-oval with narrowed apex bearing a palpal cone; (4) pronotum with a simple convex disc; three antebasal foveae, a lateral each side and a median, these three foveae united by an arcuate antebasal sulcus; (5) elytron quadrifoveate and flank with a subhumeral fovea; (6) abdomen with five 14 FIELDIANA: ZOOLOGY. VOLUME 68 tergites; no abdominal margins per se, but with firet tergite bearing each side two short carinae; six stemites in fenuile; male with seven stemites, the seventh obliquely divided into a right and left aedegeal plate; first stemite in both sexes fully visible posteriad of thorax; (7) pro-, meso-, and metastema not medianly carinated; (8) mesocoxal cavities closed by narrow processes from meso-, and metastemum; (9) metacoxae widely separated; (10) tarsi 3-segmented; segment 1 short, 2-3 long and the last bearing a single claw. Type-species: Insulomodes tobagoensis Park, new species. This new genus resembles the neotropical Dalmodes (Reitter, 1882) but the latter has six sternites in both sexes, and the male sixth sternite is not divided into two aedeageal plates; it also is allied to Buris (Fletcher, 1928, p. 227) and Dalmoburis (Park, 1942, p. 266) in having the male seventh sternite similarly divided, but differs from these two genera in that they have no antebasal foveae on the elytra. Key to Species of Insulomodes 1 Seven visible stemites, the seventh obliquely, longitudinally divided into a right and left aedeagal plate (males) 2 Six visible sternites, the sixth not so divided (females) „ 3 2(1) Mesotrochanters each bearing a blunted median tooth at center of ventral face tobagoensis Park, n. sp. Mesotrochanters not toothed excavatus Paris, n. sp. 3(1) Last stemite not much longer than first, transversely semicircular in outline and weakly flattened medianly tobagoensis Park, n. sp. Last stemite twice as long as first, rounded ogival in outline with a narrow elongate-triangular impression at base. excavatus Park, n. sp. Insulomodes tobagoensis Park, new species Description. - Type male: Head 0.22 long x 0.26 wide; pronotum 0.30 x 0.36; elytra 0.46 x 0.60; abdomen 0.53 x 0.50; total length 1.5. Cuticula shining reddish brown; head and pronotum subimpunctate; elytra with sparse punctures; abdomen sparsely punctulate; pubescence moderately abundant and appressed. Head rounded trapezoidal; a pair of flattened eyes of about 24 coarse facets; tempore rounded, as long as the eyes; vertex with a pair of nude foveae which are mutually separated by twice the distance of either fovea from its adjacent eye, these foveae connected by broadly arcuate sulcus; front short and simple; labrum large with the distal margin suddenly wider; ventral surface of head transversely convex between the eyes, and with 2 gular foveae at base. Antennae ll-segmented; 1 elongate, large and parallel-sided; 2 smaller, elongate with slightly rounded sides; 3-8 smaller than second, subequally wide. 3 elongate PARK ET AL.: PSELAPHID BEETLES 15 obconical, 4-8 subequal, subquadrate; club of last three, 9 wider than eighth, transverse trapezoidal; 10 larger than ninth, transverse trapezoidal; 11 largest segment, four times longer than 10 and a fifth wider. Pronotum with simple convex disc; a nude lateral antebasal fovea each side and a nude median antebasal fovea, these three connected by an arcuate transverse sulcus. Elytra with rounded humeri, each elytron with four large, nude antebasal foveae, entire sutural stria from mesial fovea; flank with a nude subhumeral fovea and an entire carina arising from the foveal rim. Metathoracic wings present. Aedeagus 0.30 long X 0.15 wide (fig. 4), dissected from a paratype. Abdomen with five visible tergites in median length ratio of 2.0/1.8/1.6/1.6/1.2. No lateral abdominal margins per se; first tergite bearing each side two marginal carinae which extend from basal margin through basal half of tergite length, inner stria straight and outer stria sinuous; first tergite bearing a pair of basal abdominal carinae which are stout, straight and divergent, separated distally by 24 per cent of segmental width and they are 40 per cent of segmental length. Seven stemites in median length ratio of 1.4/1.0/0.8/0.6/0.4/1.2/0.8; first sternite fully visible side to side and extending well posteriad of metacoxae; third sternite bearing a small, elongate, distinctive lamina which tends to bear vertical striations and arises from center of posterior margin and is abruptly reflected anteriorly; sixth sternite bearing a median subcircular impression and posterior segmental margin medianly briefly elevated; seventh sternite transversely rounded-triangular in outline and divided obliquely and slightly asymmetrically into a right and left aedegeal plate (these two plates are articulated laterally and probably are moved laterally at copulation to allow extrusion of the aedeagus, but when these plates are appressed the line of separation may be difficult to observe). Legs stout with femora swollen medianly and tibiae swollen distally; mesotrochanters each bearing a small, blunted, subcyclindrical tooth at center of ventral face; metatibiae rather suddenly arcuate for distal third; tarsi with a single large claw. Female: with minor proportional differences, as described for the male except for the following exceptions: (1) only six visible sternites, in median length ratio of 1.6/1.2/1.0/0.8/0.4/1.8 with the last sternite transverse and almost semicircular in outline and weakly flattened medianly; (2) no lamina on third sternite; (3) mesotrochanters not toothed.' Within the known population fragment of this species there is the expected amount of minor variation in size and structure. As 'Where females are preserved with the terminal tergite and sternite sufficiently separated there is revealed a pair of weakly sclerotized rounded-triangular processes. This pair is articulated laterally and may be critically involved in clasping a part of the aedeagus during copulation. They may represent a part of the next sternite. This would be the seventh if the first six visible sternites are considered, but morphologically the first two sternites are membranous and not visible in prepared specimens, and this would indicate that the two processes of the female belong to the true ninth sternite. The pselaphid abdomen is thought of as having 10 segments by Park (1942). This view agrees with Blackwelder (1936) for Staphylinidae, and with Tanner (1927) for Coleoptera. In this case the ninth and tenth segments would be involved in the genitalia. 16 FIELDIANA: ZOOLOGY. VOLUME 68 between males the length and degree of bluntness of the mesotrochantal spine and the size of the lamina on the third sternite varies as does also the amount of mesial flattening of the sixth sternite in females. This species is based on four males and two females (type and five paratypes) collected by the Rev. E. J. Pearce beneath bark and in rotten log mold in the vicinity of Hillsborough Dam and Mt. Scarborough, Tobago on April 3 to 5, 1961. Not included in the above type series are six males and six females from beneath bark on the Woodford Estate near St. John's Grenada collected by the Rev. Pearce May 5, 1962 and one male taken by the Rev. Pearce under bark on December 9, 1960 near Diego Martin, Trinidad. These 19 specimens are in the author's collection. Since this new species is known from Grenada, Tobago, and Trinidad it is probably a member of the large mainland fauna of South America and should occur in Venezuela. Insulomodes excavatus Park, new species Description. - Type Male: Head 0.30 long x 0.40 wide; pronotum 0.36 x 0.43; elytra 0.55 X 0.76; abdomen 0.81 x .68; total length 1.7. Head with small, short but deep flattened eyes of about 26 facets; tempora rounded and as long as eyes; vertex, front, labrum and ventral surface of head as for /. tobagoensis; maxillary palpi as for genus. Antennae 11-segmented, distantly articulated, segments 1-4 as for tobagoensis; 5- 10 progressively more transverse and progressively produced mesially; 11 about as long as 7-10 united (antennal description checked against a paratype slide-mount at 430 diameters). Pronotum and elytra as for tobagoensis. Metathoracic wings present. Abdomen with five visible tergites in median length ratio of 2.2/2.2/2.2/2.4/2.8 with no lateral margins per se, the lateral carinae as for tobagoensis but heavier; first tergite with basal abdominal carinae strong and very divergent, 33 per cent the length and separated by 33 per cent the width of segment; last tergite vertical, rounded trapezoidal. Seven stemites in median length ratio of 1.2/1.2/0.8/0.4/0.2/2.4/1.2 with sixth sternite complex: median half of sternite deeply subcircularly excavated, with a small median tumulus at basal margin and a small rounded cusp on distal margin set obliquely each side of excavation and projecting obliquely above this cavity; seventh sternite divided obliquely into a right and a left aedeageal plate, articulated laterally for extrusion of aedeagus (in a slide mount of a paratype male the distal rim of the excavation, between the cusps, is bounded by a row of smaU, distinctive tubercles seen clearly at 430 diameters). Legs strong, femora medianly swollen; tarsi as for tobagoensis; all tibiae thickened distally; mesotibiae suddenly angulated at center and the point of PARK ET AL.: PSELAPHID BEETLES 17 angulation concave ventrally; mesotrochantere not toothed; metacoxae distantly articulated; metatibiae slowly arcuate mesially for distal third. Female generally similar to male with the following exceptions: (1) only six visible stemites in median length ratio of 1.2/1.0/0.8/0.6/0.4/2.4 with the sixth twice the length of first and rounded-ogival in perimeter, from a direct dorsal view this sixth stemite has a narrowly elongate-triangular foveoid impression for basal third; (2) mesotibiae normally arcuate. Described from five males and four females collected by Rev. E. J. Pearce between April 3 to 5, 1961 in Tobago. The type male and three paratype females were taken under bark at Hillsborough Dam. Four males, including one teneral, and one female, all paratypes were taken in a rotten log at Mr. Scarborough. Type and paratypes in the O. Park collection. Genus Euharmophola Park Euharmophola Park, 1960, Trans. Amer. Microscop. Soc, 79 (1), p. 23. Type-species: Euharmophola carinata Park, loc. cit., p. 23. Euharmophola carinata Park, 1960, Jamaica (rotting log mold, 12 miles south of Falmouth, Trelawny Parish, Cornwall County). Genus Dalmonexus Park Dalmonexus Park, 1942, Northwestern Univ. Stud. Biol. Sci. Med., 1, p. 272. Type-species: Dalmonexus seeversi Park, loc. cit., Barro Colorado Island, Panama. Dalmonexus tobagoensis Park, new species Description. - Type male: Head 0.26 long x 0.34 wide; pronotum 0.27 x 0.38; elytra 0.46 x 0.58; abdomen 0.39 X 0.52; total length 1.4. Cuticula shining reddish- brown, subimpunctate with sparse bristling, suberect pubescence. Head with a pair of rather small, subcircular eyes of about 28 small facets; eyes inconspicuous from above being set far down on genae and on a line with mandibular articulations, this lateral view shows the tumid vertex seen as a high, rounded tumulus. Head from dorsal view relatively complex: a median vertexal carina crosses occiput to basal portion of vertex; tumid vertex bears medianly in apical portion a shallow, glabrous depression; apical of this depression the vertex is abruptly transversely excavated to the interantennal line of the front; this frontal wall bears medianly a mat of very short, golden setae arranged in two confluent triangular masses and this wall is undercut each side; distal floor of excavation slightly higher medianly and the floor drops abruptly beyond antennal articulations to form a short, simple fronto-clypeal strip; labrum with distal margin suddenly expanded laterally; ventral surface of head simply convex with a broad basal foveoid depression; maxillary palpi as for genus. 18 FIELDIANA: ZOOLOGY. VOLUME 68 Antennae ll-segmented, distantly articulated; segment 1 relatively large and elongate; 2 smaller, elongate; 3-7 subequal, submoniliform; 8 slightly wider, slightly transverse; club of last three segments; 9 larger than eighth, transverse trapezoidal; 10 larger than ninth, transversely trapezoidal; 11 largest segment, as long as 7-10 united. Pronotum with each lateral margin incised near basal third, a lateral fovea in this incisure and these two antebasal foveae united by an arcuate sulcus; disc simply convex. Elytra bifoveate, entire sutural stria from sutural fovea and a vague, short discal impression from discal fovea; flank not fovea te but bearing a carina for apical half of length. Metathoracic wings present. Abdomen with five visible tergites in median length ratio of 2.4/1.0/1.0/1.0/1.2/. No lateral margins per se but first tergite with two strong, straight, subentire lateral carinae each side and second and third tergites with one entire carina each side. Six visible sternites in median length ratio of 0.8/1.0/0.4/0.2/0.2/1.2, sixth stemite punctulate and with a small foveoid depression at apical margin; first stemite fully visible beyond thorax. Pro-, and mesosterna not carinate medianly; metastemum medianly, longitud- inally sulcoid. Legs relatively heavy and femora swollen medianly; mesotrochanters each with a short, stout tooth at lateroventral angle; metatibiae arcuate in apical third; tarsi with a single claw. Female: generally as for male except that (I) the vertex normally convex, so that the median, longitudinal carina from the occiput curves briefly on the base of the vertex; vertex-frontal area simple, so that the pair of vertexal foveae are clearly visible, one at base of each antennal incisure, and these foveae connected by a broadly arcuate sulcus; (2) antennal segment 10 transversely ovidal; 11 not as long, in a ratio of 1.6/2.0 as for the male, but still as long as preceding four segments united; (3) six sternites in median length ratio of 0.8/1.2/0.4/0.6/0.6/1.2 with sixth stemite weakly depressed at center of base; (4) mesotrochanters not toothed. Description based on one pair (type and paratype) in author's collection. Collected by Rev. E. J. Pearce under bark on April 5, 1961 at Hillsborough Dam, Tobago. The only ally is D. seeversi Park (1942, p. 272) from Barro Colorado Island, Panama Canal Zone, but the male seeversi has a much more complex vertexal- frontal area with only the median part of the vertex uncarinated, and the female seeversi is unknown. Subtribe TANYPLEURINA sensu strictiore Tanypleurina Jeannel, 1949. Mem. Mus. Nat. d'Hist. Nat., N. S.. 29. p. 79. Key to Genera of Tanypleurina Elytral flank with a subhumeral fovea Buris Fletcher Elytral flank with subhumeral fovea absent Bythinophysis Raffray PARK ET AL.: PSELAPHID BEETLES 19 Genus Buris Fletcher Buris Fletcher, 1928, Ann. Entomol. Soc. Amer., 21, p. 227. Type-species: Buris brunneus Fletcher, loc. cit.. Vera Cruz, Vera Cruz, Mexico. Buris ensipes (Raffray), 1891, Ann. Soc. Entomol. France, ser. 6, 10, 316. Described from San Esteban, Venezuela. Our records are from Trinidad, and Antigua, Leeward Islands. This species was described in the genus Dalmodes Raffray (1908, p. 39). [Park, 1942, placed ensipes in Bythinophysis, but now places it in Buris. J.W. - M.W.S.] Genus Bythinophysis Raffray Bythinophysis Raffray, 1908, Genera Insectorum, 64, p. 266. Ty{>e-species: Bythinophysis punctipennis Raffray, loc. cit., p. 266, Cayenne, French Guiana. Bythinophysis humilis (Raffray), 1908, Ann. Soc. Entomol. France, 77, p. 39. Guadeloupe, Leeward Islands. This species was described in the genus Dalmodes. Tribe EUPLECTINI sensu latiore Euplectini Park, 1942, Northwestern Univ. Stud. Biol. Sci. Med., 1, p. 63. Key to Subtribes of EUPLECTINI 1 Front of head prolonged into a conspicuous elongate rostrum, with antennae articulated at distal end of rostrum and subcontiguous. RHINOSCEPSINA Front not as described 2 2 (1) Tarsi with two claws, a large primary claw and a short accessory claw. TROGASTRINA Tarsi with either a single claw, or a claw and an accessory seta 3 3 (2) Mesocoxae well separated; the coxal cavities separated by distinct processes from the meso-, and metasternum PANAPHANTINA Mesocoxae contiguous, the coxal cavities confluent 4 4 (3) Presternum longitudinally bisected by a median carina. BIBLOPORINA Prostemum not bisected by such a carina or a carinoid fold 5 5 (4) Antennae with club formed by the last three or four segments. EUPLECTINA sensu strictiore' 'Two Antillean genera, Latomelba and Hanfordia, have an intermediate antennal club and might be placed in either Euplectina or Trimiina on this character. Because they have gular capitate setae and other features they are assigned to the Trimiina, q. v. 20 FIELDIANA: ZOOLOGY. VOLUME 68 Antennal club formed by the last segment only, segment very large and as long as the preceding three to six segments united TRIMIINA (see footnote p. 19) Subtribe RHINOSCEPSINA Rhinoscepsina Bowman, 1934, Pselaphidae of N. Amer., p. 144. Genus Rhinoscepsis LeConte Rhinoscepsis, 1878, Proc. Amer. Phil. Soc, 17, p. 382. Type-species: Rhinocepsis bistriatus LeConte, loc. cit., Florida, U.S.A. Rhinoscepsis insularis Park, 1955, Bull. Chicago Acad. Sci., 10 (7), p. 105. Described from Morce's Gap, St. Andrew Parish, Surrey County, Jamaica, 5,500 ft. elevation. Subtribe TROGASTRINA Trogastrina Jeannel, 1949, Mem. Mus. Nat. d'Hist. Nat., N. S., 29, p. 75. Key to Genera of Trogastrina Pronotal disc bisected by a deep longitudinal sulcus; each lateral margin bearing six teeth Eurhexius Sharp Pronotal disc with a hardly discernible median longitudinal impression; lateral pronotal margins crenulated only Rhexinia Raffray Genus Eurhexius Sharp Eurhexius Sharp, 1887, Biol. Centr.-Amer., 2 (1), p. 41. Type-species: Eurhexius vestitus Sharp, loc. cit., Panama. Eurhexius incertus Park, new species Description. — Type measurements: Head 0.24 long x 0.38 wide; pronotum 0.38 X 0.41; elytra 0.48 x 0.61; abdomen 0.55 x 0.58; total length 1.6. Cuticula shining reddish brown throughout, sparingly punctulate and with appressed pubescence. Head very transverse with prominent eyes which are subcircular in lateral view and about twice as long as the tempora; temporal angles rounded; a median longitudinal sulcus from occiput to a line through eye-centers on vertex; usually pair of mutually distant small tubercles on occipital margin; pair of mutually distant vertexal foveae connected by a broadly arcuate sulcus; fronto-clypeal area simple and punctures larger; mandibles stout; maxillary palpi as for genus; ventral surface of head bearing longer, bristling genal setae; deeply concave in center of gular area and from this region a median longitudinal carina extends to basal margin of the mouth. PARK ET AL.: PSELAPHID BEETLES 21 Antennae 11 -segmented, distantly articulated; segment 1 elongate, subcylindri- cal; 2 subcircular, as wide as first but one-fifth as long; 3-6 subequal in width, very transverse-moniliform, about twice as wide as long; 7 and 8 slightly wider, transverse- moniliform; well-developed club of 9-11; 9 twice as wide as eighth, transverse- trapezoidal; 10 larger than ninth, twice as wide as long; 11 largest segment, slightly more than twice as long as wide and about as long as 8-10 united. Pronotum orbicular, deeply bisected by an entire longitudinal sulcus; an arcuate sulcus uniting a median and a large lateral antebasal fovea each side; lateral margins each with six teeth, these teeth becoming smaller distally. Elytron antebasally quadrifoveate with entire sutural stria but no subhumeral fovea. Metathoracic wings present. Abdomen with five visible tergites in median length ratio of 2.4/1.4/1.4/1.6/1.2 with the fifth stemite vertical and the first four bearing strong lateral margins. Six visible stemites, the first bearing distally an especially strong row of prostrate setae; these six stemites in median length ratio of 1.0/2.0/1.2/0.8/0.6/0.6 and medianly at base of third to fifth stemites is a very small glabrous area. Legs unmodified except for the usual lamina found in the genus in both sexes at distal three-fourths of ventral face of profemora; tarsi as for genus, with a strong primary claw and a shorter secondary claw. Described on a unique specimen collected by Rev. E. J. Pearce under bark on April 5, 1961 at Hillsborough Dam, Tobago. The type in author's collection. The sex of the type is uncertain. The rather large eyes, distinctly longer than the tempora suggest a male, but not invariably so in the genus. On the other hand, the length ratio of the fifth and sixth sternites suggest a female. The species is most closely allied to hambletoni Park (1952) from Vicosa, Minas Gerais, Brazil. Both sexes are known for this latter species, and the female hambletoni is quickly differentiated from incertus since, among other features, it has obliquely suboval eyes and the fifth stemite is from one-half to two-thirds as long as the sixth stemite. The male hambletoni has eyes only as long as tempora, and the fifth sternite is only about one-eighth the length of the sixth sternite. Genus Rhexinia Raffray Rhexinia Raffray, 1890a, Rev. d'Entomol., 9, pp. 104, 106. Type-species: Rhexinia angulata Raffray, loc. cit., p. 196, New Grenada, (Colombia?) Rhexinia versicolor Raffray, 1908, Ann. Soc. Entomol. France, 77, p. 37, Guadeloupe, Leeward Islands Subtribe PANAPHANTINA Panaphantina, Jeannel, 1950, Lechevalier, Paris, p. 76 22 FIELDIANA: ZOOLOGY. VOLUME 68 Key to Genera op Panaphantina Mesocoxae distant, separated by subtruncate, contiguous processes of mesostema and metastema; elytron trifoveate; male with seven visible stemites Thesium Casey Mesocoxae narrowly separated by distally rounded and overlapping processes of mesostema and metastema; elytron quadrifoveate; male with only six visible stemites Thesiectus Park Genus Thesiectus Park Thesiectus Park. 1960. Trans. Amer. Microscop. Soc. 79 (1), p. 8. Type-species: Thesiectus prohus Park. loc. cit.. p. 9. Thesiectus probus Park. Collected from a rotten log, 12 miles south of Falmouth, Trelawny Parish, Jamaica. Genus Thesium Casey Thesium Casey. 1884, Philadelphia. Penn.. pp. 94. 117. Type-species: Thesium cavifrons (LeConte). designated by Bowman. 1934, Pselaphidae of N. Amer.. p. 144. Penn.? Eastern North America. In this genus the males have seven stemites, the seventh being an ovoidal to circular aedeagal plate whereas the females have only six stemites. Key to Species of Thesium 1 Pronotal disc bearing a subcircular small fovea about its own diameter from the apical pronotal margin affabilis Park Pronotal disc bearing an elongate-oval fovea which is much larger, occupying the median half or more 2 2(1) First antennal segment triangular in cross-section, the mesial and lateral surfaces strongly obliquely flattened to form a fine dorsal edge, this edge elevated into a laminoid wing antennalis Park First antennal segment not as described 3 3 (2) Antennae exceptionally stocky with tenth segment three times wider than long .pearcei Park, n. sp. Tenth antennal segment not this transverse 4 4 (3) Male with frontal declivity opaque, subgranulate and bearing at its apical tip a membranous hom or a hom-like tuft of recurved setae. cornutum Park Male with frontal area quite different: bearing a relatively large circular, glabrous depression in the center of which is a small penicillate process which projects anteriorly frontalis Park PARK ET AL.: PSELAPHID BEETLES 23 Thesium pearcei Park, new species Description. — Type male. Head 0.16 x 0.22 wide; pronotum 0.19 x 0.23; elytra 0.29 X 0.34; abdomen 0.34 X 0.30; total length 0.98. Cuticula uniform yellowish brown; subimpunctate; sparse pubescence, being longer and less appressed on abdomen. Head with prominent eyes one-third longer than rounded tempora; vertex with a pair of pubescent foveae united by an angulated sulcus which terminates at base of the elevated frontal margin; face declivous between antennal acetabulae and the declivity bisected medianly by a carina, the tip of which is seen as a minute tubercle on frontal margin; ventral surface of head with submental area microgranulate; maxillary palpi as for genus. Antennae 11-segmented, articulated on either side of frontal rostrum; segment I subquadrate, simple; 2 shorter than first and subquadrate; 3 to 10 increasingly transverse, the club of 10 and 11 only; 10 very transverse, three times wider than long; 11 largest segment, five times longer than tenth and terminating in a very short conoidal apex. In general the antennae are exceptionally stocky and segments 3 to 10 unusually wider than long. Pronotum slightly narrowed at base and apex and only slightly wider than head; lateral margins crenulated; a pubescent lateral fovea each side, set in a larger depression; a median nude foveoid depression; these three areas united by a transverse antebasal sulcus, but this sulcus interrupted each side, between lateral fovea and median depression by tubercle; disc with a large, elongate oval foveoid impression from near apical margin almost to the transverse sulcus. Elytra with well-developed, rounded humeri; elytron trivoeate, the foveae pubescent; sutural stria entire; sutural and discal foveae approximate and separated from humeral fovea by a longitudinal plicoid fold; impression from humeral fovea extending through basal half of elytral length; flank with a pubescent subhumeral fovea and a carinoid line at apical four-fifths. Metatathoracic wings present. Abdomen with five visible tergites, the first four bearing strong lateral margitis; median length ratio of 1.0/1.0/1.0/1.0/0.8 and last tergite vertical. Seven sternites visible, in median length of ratio of 0.6/1.0/0.8/0.8/0.2/0.4/1.0 with six medianly concave and seventh an asymmetrically articulated, ovoidal aedeagal plate. Presternum medianly bisected by a carina and mesocoxal cavities well -separated by sternal process, as usual in genus; metasternum bearing a narrow Y-shaped impression, each arm in the Y extending to mesocoxal cavity. Legs simple, with tarsi bearing a single large claw. This species is based on a single male, the type in author's collection. It was collected by Rev. E. J. Pearce, in whose honor it is named, under a board on April 5, 1961 at Hillsborough Dam, Tobago. In a key based on study of type specimens (Park, 1952b, pp. 115-116) of species known from the Neotropical Region, pearcei keys out with noctumalis Park (1952b, p. 117) from near Mexico, D. F. but these two species are very different from one another. This nocturnalis has the lateral pronotal margins coarsely and obviously crenulated and antennal segments 3 to 10 relatively slender and less 24 FIELDIANA: ZOOLOGY. VOLUME 68 transverse, whereas pearcei has the lateral pronotal margins just discernibly microcrenulate and antennal segments 3 to 10 trans- verse progressively to the tenth which is three times longer than wide. Furthermore, this 1952 key was published when the genus Thesium was unknown from the West Indies. Since then four species of the genus have been described from Jamaica (Park, 1955, 1960). T. pearcei is quickly discriminated from this Jamaican assemblage as a consequence of its unique antennae {vide supra); probably its nearest allies will be discovered in the large and still poorly known South American fauna. Subtribe BIBLOPORINA Bibloporina Park, 1951, Geol. Surv. Alabama, Mus. Paper, 31, p. 64 Genus Biblomimus Raffray Biblomimus Raffray, 1903, Ann. Soc. Entomol. France, 72, p. 62. Type-species: Biblomimus minutus Raffray, loc. cit., p. 62, St. Vincent, (Windward Islands) Key to Species of Biblomimus 1 Antennal segment 10 very transverse, twice as wide as ninth. minutus Raffray Antennal segment 10 not appreciably wider than ninth but twice as long. ...2 2(1) Seven visible stemites, seventh ovate longitudinally. male impressa Raffray Six visible stemites, sixth triangular female impressa Raffray Subtribe EUPLECTINA s. str. Euplectina Jeannel, 1949, Mem. Mus. Nat. d'Hist. Nat., N. S., 29, p. 45 Key to Genera of Euplectina Ventral surface of head with stout, subspiniform, strongly capitate setae. Thesiastes Casey Ventral surface of head with simple, aciculate, semiappressed setae. Euplectus Leach Genus Thesiastes Casey Thesiastes Casey, 1893, Ann. New York Acad. Sci.. 7, pp. 444. 457. Type-species: Thesiastes debilis (LeConte), designated by Bowman, 1934, Pselaphidae of N. Amer.. p. 144, Florida, U.S.A. PARK ET AL.: PSELAPHID BEETLES 25 Key to Species ok Thesiastes Relatively small (0.9 mm.); pronotal disc with a large oblong median fovea; first two visible tergites with a pair of basal abdominal carinae which bound more than one- third of discal width liliputanus Raffray Relatively large (1.2 mm.); pronotal disc with a very long, relatively narrow median fovea; basal abdominal carinae not evident hopensis Park, n. .sp. Thesiastes hopensis Park, new species Description. - Type female: Head 0.21 long X 0.27 wide; pronotum 0.27 x 0.31; elytra 0.29 x 0.41; abdomen 0.41 x 0.41; total length 1.2. Cuticula shining reddish brown; head, pronotum and elytra microgranualte- punctate; pubescence appressed, longer on abdomen with fourth tergite bearing arcuate guard setae. Head with prominent, relatively coarsely faceted eyes; tempora rounded, shorter than eyes; a pair of vertexal foveae united by an inverted V-shaped sulcus which is deepest and broader around each fovea; occiput deeply medianly sulcate; face simple and convex-declivous; maxillary palpi as for genus; ventral .surface of head with abundant short capitate setae. Antennae 11-segmented, widely separated; segment 1 subquadrate; 2 narrower than first, elongate-cylindrical; 3 to 8 much shorter, third subquadrate and obtrapezoidal, fourth to eighth as wide as third, subspherical; club well developed of last three segments with 9 transverse trapezoidal, distinctly larger than eighth; 10 larger than ninth, transverse trapezoidal, 11 largest, one-fourth longer than wide, suboviform and one-fourth longer than 9 and 10 united. Pronotum with three antebasal foveae united by an arcuate sulcus; a sulcoid impression extends anteriorly from each lateral fovea; disc with a long median fovea from which extends a sulcoid impression nearly to the anterior margin. Elytra with rounded humeri; elytron trifoveate with entire sutural stria and a short discal impression; flank with a subhumeral fovea and an arcuate carina which curves sharply above base of fovea and extends to posterior elytral margin. Metathoracic wings present. Abdomen with five visible tergites in median length ratio of 1.2/1.2/1.2/1.0/1.0 with first three bearing strong lateral margins; basal abdominal carinae not evident on first two tergites; fifth tergite with a dark brown area medianly (which could be confused with a seventh sternite on casual inspection) and its posterior margin slightly indented medianly. Six visible sternites with the sixth medianly .slightly extended to fit the incisure of the fifth tergite. Metasternum convex. Legs rather short with femora slightly swollen. A noteworthy feature is found on the ventral face of each profemur: a longitudinal row of about eight microtubercles, each of which bears a very short, stiff white seta. Described from four females. Type and two paratypes in the collections of the Illinois Natural History Survey and one paratype in author's collection. Collected by H. B. Mills on May 6, 1950, in the Hope Gardens at Kingston, Jamaica. The new species is allied distantly to T. liliputanus Raffray, 1903, p. 543 from Grenada. 26 FIELDIANA: ZOOLOGY. VOLUME 68 Genus Euplectus Leach Euplectus Leach. 1817, Zool. MLst-.. 3, p. H(). Type-spet-ies: Euplectus nanus Reichenbach. 1816. Lipsiae, p. 69. "Europe." Sex can be determined by the number of sternites. Females have six visible, the last simple and subtriangular. Males have seven visible, the seventh being longitudinally divided. Kky TO Species of Euplectus 1 Lateral pronotal margins neither sinuate near base nor having a marginal tooth or tubercle in sinuation near lateral antebasal foveae. exiguus Raff ray Lateral pronotal margins sinuate near base and having either a small tubercle or tooth in sinuation near lateral antebasal foveae 2 2(1) Vertexal foveae with interfoveal sulcus obsolete apically; male fourth sternite evenly convex and unmodified insularis Raffray Vertexal foveae united by a strong interfoveal sulcus 3 3 (2) Male fourth sternite bearing a median, transverse fossa. iUepidus Raffray Male with first four sternites unmodified 4 4 (3) Male with fifth sternite transversely impressed near basal margin for median half of width ^ouyai'ensis Park. n. sp. Male with fifth sternite bearing three deep, transverse impressions near basal margin, these three separated by two elevations. episcopalis Park, n. sp. Euplectus gouyavensis Park, new species Description. - Type male: Head 0.24 long X 0.33 wide; pronotum 0.31 X 0.33; elytra 0.43 x 0.41; abdomen 0.53 x 0.43; total length 1.5. Cuticula shining reddish brown with paler elytra; head and pronotum sparsely punctate, elytra more densely punctate; abdomen subimpunctate except seventh sternite which is obviously punctate; pubescence appressed. Head with eyes and tempora of same length; occiput deeply notched medianly; vertex with a pair of nude vertexal foveae united by an inverted U-shaped sulcus; area enclosed by this sulcus glabrous and depressed; face subvertical and transversely recessed; labrum with distal margin concave; mandibles strong; maxillary palpi as for genus. Antennae 1 1 -segmented, distantly articulated; segment 1 elongate; subcylindri- cal; 2 smaller, quadrate; 3 to 8 smaller, third briefly obconical. fourth to seventh moniliform. eighth slightly larger, trapezoidal; club of last three segments with 9 twice as wide as long. 10 larger than ninth and twice as wide as long. 11 largest segment, one-third longer than wide and as long as 8-10 united. Pronotum with three nude antebasal foveae united by a transverse sulcus; the median foveoid area extends to base, and continues apically to unite with the PARK ET AL.: PSELAPHID BEETLES 27 longitudinal discal sulcus which extends to apical three-fourths of pronotal length; lateral margins incised near lateral foveae and bearing a small, blunt tooth at incisure. Elytron trifoveate, entire sutural stria, a short discal impression for basal fifth near humeral fovea; flank with a subhumeral fovea. Abdomen with five visible tergites, the first three strongly margined, in a median length ratio of 1.2/1.4/1.6/2.2/1.4 and widest through the third tergite; fifth subvertical; first two bearing a pair of basal abdominal carinae, with those of second divergent, 45 per cent of tergite length and separated by 25 per cent of tergite width. Seven sternites, the venter being concave in profile and in median length ratio 1.0/0.8/0.8/0.6/0.2/0.6/1.8 with the first four unmodified; fifth sternite transversely impressed near basal margin for median half of width, this ver\' short impression extending anteriorly beneath the fourth sternite; seventh sternite divided longitud- inally into a right and left aedeageal plate of which the left is slightly larger; the line of union ("pygidial carina") is slightly inclined to the morphological right side. The inclination of the "pygidial carina" to the morphological right side places gouyaiensis in the genus Pycnoplectus (Casey, 1897) following the revision of the North American Euplectus-complex by Wagner (1975). Since not all of the Neotropical species have been studied with reference to the inclination of the "pygidial carina," and the fauna is poorly known, these species have been left in Euplectus for the present. Metasternum longitudinally medianly sulcoid; legs unmodified, with femora slightly tumid medianly and tarsi with a single large claw. Described from the unique male type, in author's collection; collected by the Rev. E. J. Pearce under bark on April 27, 1962 near Gouyave, Grenada. The nearest ally of this new species is illepidus Raffray also from Grenada, Windward Islands but this species has a fossa on the fourth sternite. Euplectus episcopalis Park, new species Description. — Type male: Head 0.22 long x 0.31 wide; pronotum 0.31 X 0.31; elytra 0.39 X 0.41; abdomen 0.45 X 0.38; total length 1.4. Cuticula as for gouyavensis except that elytra are not paler. Head with eyes one-fourth shorter than tempora: vertexal foveae and uniting sulcus as for gouyavensis except the enclosed area is medianly slightly elevated; face subvertical with median area not as recessed as lateral areas; labrum with concave apical margin slightly everted medianly; mandibles strong and maxillary palpi as for genus. Antennae 11-segmented, distantly articulated; segment 1 elongate and subcylin- drical; 2 smaller, slightly elongate; 3 to 6 smaller, submoniHform; 7 to 8 slightly larger; club of last three segments, with 9 almost three times wider than long, trapezoidal; 10 about the same size but shorter and slightly more than three times wider than long; 11 largest segment, as long as 8 to 10 united. Pronotum as for gouyavensis the discal fovea is much shorter, elongate-oval, and occupying the median half of disc. Elytra as for gouyavensis. 28 FIELDIANA: ZOOLOGY. VOLUME 68 Abdomen with five visible tergites, first three strongly margined, in a median length ratio of 1.0/1.0/1.0/2.0/1.2 and widest through third tergite; Hfth subvertical; basal abdominal carinae on first two tergites, those on second divergent, 25 per cent of tergite length and separated by 25 per cent of tergite width. Seven stemites in a median length ratio of 0.8/0.8/0.8/0.6/0.4/0.2/1.8 and concave in profile; first four stemites unmodified; fifth stemite complexly modified: the basal half bears three transverse excavations, a deep, glabrous excavation each side, and a deeper and wider median excavation; these three excavations are separated by a pair of elevations and all three extend anteriorly beneath the fourth stemite; the median excavation is setose basally. Seventh stemite as for gouyavensis. Metastemum and legs as for gouyavensis. Described from the unique male type, in author's collection; collected by the Rev. E, J. Pearce in a banana stalk on April 29, 1960 at Campbeltown, Tobago. Its nearest ally is gouyavensis described from Grenada in this paper, but gouyavensis has a single, simple, very short transverse impression near base of the fifth stemite in contrast to the much more complex modifications of this stemite in episcopalis. Other characters separating these two new species include the eye/tempora length ratio and the length of the basal abdominal carinae of the second tergite. Subtribe TRIMIINA Trimiina Bowman, 1934, Pselaphidae of N. Amer., p. 8. Key of Genera to Trimiina 1 Dorsal head outline of male unique in subtribe, obviously wider than any other part of body, and large eyes wholly invisible from above. Malleoceps Park Dorsal head outline never wider than elytra in either sex 2 2 (1) Both pronotum and elytra lacking foveae Hanfordia Park Pronotum and/or elytra bearing foveae 3 3 (2) Metacoxae relatively distant from each other, separated by the width of their mesial articular surfaces or more at bases 4 Metacoxae subcontiguous or slightly separated at bases 6 4 (3) Elytron with subhumeral fovea present 5 Elytron with subhumeral fovea absent Dalmomelha Parte 5 (4) Basal fifth of pronotum medianly bisected by a high, laminoid carina from basal bead to transverse antebasal sulcus Lemelba Park Pronotum lacking this feature Haasiella Park, n. g. PARK ET AL.: PSELAPHID BEETLES 29 6 (3) Dorsal outline of elytral humeri subdentate to dentate as a consequence of the position of subhumeral fovea and adjacent cuticula. Actium Casey Elytral humeri rounded, whether subhumeral foveae are present or absent.? 7 (6) Pronotum with lateral foveae present and mesiad of pronotal margins and (discounting pubescence) wholly visible or partially visible from a dorsal view 11 Pronotum with lateral foveae either absent, or vestigial, or well formed, but if the latter, wholly invisible from a dorsal view and located on the flanks of pronotum 8 8 (7) Flank of elytron bearing a subhumeral fovea 9 Flank of elytron with subhumeral fovea absent Melba Casey 9 (8) Head slightly to obviously wider than pronotum; males (seven visible sternites, females with six visible stemites) with fourth stemite each side bearing a pair of distinctive spatulate setae Latomelba Park Head as wide or narrower than pronotum; males (same stemite numbers as for Latomelba) with such spatulate setae absent 10 10(9) Head quadrate in dorsal view; males (same stemite numbers as for Latomelba) with head dilated distally Quadrelba Park Head narrowed distally in both sexes 16 11(7) Vertex with the usual pair of foveae between the eyes 13 In addition to the pair of interocular foveae, vertex bears a second pair, a fovea or foveoid pit slightly behind and mesad of each antennal prominence 12 12 (11) Eyes normal, of 30 facets or more (fig. 7) Ramelbida Park Eyes rudimentary, of 9 facets (fig. 6) female Ma//eoceps Park 13 (11) Head through eyes slightly wider than pronotum (figs. 8, 10) 14 Head through eyes not wider than pronotum Trimiovillus Park 14 (13) Antennae distantly articulated from each other 15 Antennae closely articulated on a broad, short rostrum. Sandersonella Park, n. g. 15 (14) Elytral flank with no trace of subhumeral fovea Trimiosella Raffray Elytral flank with a subhumeral fovea or foveoid impression. Hispanisella Park, n.g. 16 (10) Vertexal fovea united by an entire sulcus Allomelba Park Vertexal foveae free Zolium Casey Genus Malleoceps Park Malleoceps Park, 1954, Nat. Hist. Misc., 138, p. 1. Type-species: Malleoceps darUngtoni Park, loc. cit., p. 1, Sanchez, Dominican Republic. aO FIELDIANA: ZOOLOGY. VOLUME 68 Key to Species of Malleoceps 1 Six viable stemites female pear cei Park, n. sp. Seven stemites visible (males) 2 2(1) Vertex with a median tubercle between two deep fossae (fig. 6). pearcei Park, n. sp. Vertex convex, lacking median tubercle or fossae (fig. 5) darlingtoni Park Malleoceps pearcei Park, new species ■ Description. - Type male: Head 0.22 long x 0.43 wide; pronotum 0.33 x 0.24; elytra 0.38 x 0.38; abdomen 0.41 x 0.34; total length 1.34. Cuticula shining yellowish brown; occiput micropunctulate; pronotum, elytra subimpunctate; abdomen sparingly micropunctulate; pubescence sparse, short and appressed except for two special setae on venter noted later. Head (fig. 6a) obviously wider than the rest of the body; a pair of well -developed eyes of more than 30 facets, but wholly invisible from above; vertex in the form of a transverse rounded hexagon, bearing two deep median concavities; the lateral wing each side is more or less divided into an anterior oval, imbricated field and a subovate, sparingly imbricated lateral field; in the extreme lateral angle of the lateral field is a semicircular area (if one looks at the undersurface at this point, a rather large globular cavity is seen through the cuticula placed beneath this circular area. The limited material available for study did not allow preparation of slide-mounts, so it could not be ascertained whether the circular area was a membrane-covered orifice, e.g., a possible vibration perceptor, or whether the circular area was a pore, e.g., a possible chemoreceptor complex). The two vertexal concavities are glabrous and separated by a median, longitudinal lamina; arising from the posterior end of this lamina is a remarkable tubercle which seems to be surmounted by about five tortuous, possibly setoid processes. Occiput bears a minute cusp on either side of this tubercle, connected to the tubercle by a fine carina. Face below the two vertexal wings subvertical and simple; ventral surface of head simply convex with a pair of basal gular foveae and a few aciculate setae. Maxillary palpi as for genus. Antennae short, 11-segmented, distantly articulated at a (X)int exactly ventral to the junction of the anterior and lateral field of each vertexal wing; segments 1 and 2 subelongate, subequal; 3 to 8 small, subequal; 9 slightly larger and transverse; 10 larger, trapezoidal; 11 largest segment, twice as long as wide, subcorneal, as long as 5 to 10 united. Pronotum nearly glabrous with strongly convex disc; anterior two thirds orbicular; posterior third narrowing and bearing a biarcuate sulcus composed of a lateral impression down each flank and a large semicircular median impression. Elytra with prominent, rounded humeri; each elytron bifoveate; mesial fovea surrounded by a short impression which continues as an entire sutural stria; humeral fovea with similar short impression; flank with a shallow, nude subhumeral fovea which extends distally as a weak sulcoid impression bounded laterally by a carinoid ridge. Abdomen with five visible tergites, first three with strong lateral margins, in median length ratio of 1.6/1.0/0.8/0.8/0.6; first tergite with straight, divergent basal PARK ET AL.: PSELAPHID BEETLES 31 6a 6c Figs. 5-6. 6. Malleoceps darlingtoni Park, a, dorsal view of head of male, b, aedeagus. c, antenna (after Park, 1954). 6. Malleoceps pearcei n. sp., a, dorsal view of head of male paratype. b, top detail of vertexal tubercle, c, dorsal view of head of female paratype. abdominal carinae 22 per cent of segmental length and separated by 40 per cent of segmental width; fifth tergite rounded-triangular. Seven visible stemites in median length ratio of 0.6/1.0/0.8/0.4/0.4/0.4/0.2 with third, fourth, and sixth stemites each with a median glabrous impression; fourth bearing at the extreme latero-ventral angle each side a long (0.05), hyahne, stiff slightly spatulate process (the anatomy and position of these two processes are similar to the four processes of Latomelba quadrisicca Park, 1955, p. 113). Legs slender and unmodified except the ventral margins of the meso- and metatrochanters which are laminoid; tarsi with a single claw. Metastemum broadly concave; metacoxae slightly separated. Female essentially as for male with the following exceptions: 1) head (fig. 6c) relatively simple, 0.22 long by 0.24 wide; eyes rudimentary, of nine facets, visible in dorsal view; a pair of nude vertexal foveae between eyes but these foveae not united by an interfoveal sulcus; instead a sulcoid impression extends distally from each fovea ending in a weak foveoid impression mesial of each antennal prominence (this quadrifoveoid condition reminds one of the head of Ramelbida quadrifoveata (Raffray) shown in Figure 7 but the anterior pair of pits in this latter species are much more pronounced and foveoid); rounded tempora one-fourth longer than eyes; 2) six visible sternites in median length ratio of 0.6/1.4/1.0/1.0/0.2/1.4; 3) fourth 32 FIELDIANA: ZOOLOGY. VOLUME 68 stemite lacking special setoid processes; 4) meso- and metatrochanters simple; 5) metastemum much less concave. This remarkable species erected on two males and two females (male type and three paratypes) in collection of the author. Collected by Rev. E. J. Pearce, in whose honor it is named, in dead leaves at 2,700 ft. elevation near Rubias, Puerto Rico on January 24. 1961. The species is most closely allied to M. darlingtoni Park, but has many similarities in structure with Latomelba and Trimiosella. To include this new species in the genus, Malleoceps must be emended to contain species in which the metacoxae are either contiguous or slightly separated, and to restrict the remarkably wide head to the male sex. Hispanisella Park, new genus Trimiina having the following combination of structural features: (1) Head slightly wider than pronotum and bearing a pair of vertexal foveae between the eyes (fig. 8); (2) maxillary palpi 4 -segmented, segment 1 very small, 2 elongate- pedunculate, 3 short and rounded-triangular, 4 largest and longer than 2, elongate- conical with an oblique base; (3) antennae 11-segmented, distantly articulated from each other, tenth antennal segment bilaterally symmetrical and eleventh largest and forming the club; (4) ventral surface of head bearing aciculate setae; (5) pronotum with a lateral fovea each side, visible from a dorsal view, and connected to a median impression by a sulcoid impression, disc simply convex; (6) elytra with rounded humeri, bifoveate, and flank with a subhumeral foveoid impression; (7) males with seven stemites, the seventh a small aedeagal plate, and females with six stemites; (8) metacoxae slightly separated; (9) mesocoxae in confluent cavities; (10) tarsi bearing a single claw. Type-species: Hispanisella haitiana Park, new species. The nearest known alhed genus is Triangusella, known from one species, T. acuta Park (1952) (p. 132), from tropical Veracruz, Mexico, but acuta has vestigial eyes of six facets in the male, vertexal foveae united by an entire interfoveal sulcus, and a much deeper subhumeral elytral fovea. In the West Indian fauna Hispanisella appears to be most closely allied to Malleoceps as a consequence of the great similarity of the aedeagi (figs. 5, 9). Key to Species of Hispanisella Male head with eyes inconspicuous in dorsal view, and with an oval, imbricated tumulus above each antennal base (fig. 8) haitiana Park, n. sp. Male head with eyes conspicuous, and vertex greatly modified; excavated between eyes anterior to median tumulus (fig. 10) hirsuta Park, n. sp. PARK ET AL.: PSELAPHID BEETLES 33 Hispanisella haitiana Park, new species Description. - Type male: Head 0.26 long x 0.27 wide; pronotum 0.24 X 0.24; elytra 0.38 x 0.38; abdomen 0.33 x 0.38; total length 1.2. Cuticula shining reddish brown; subimpunctate except for special areas noted later; pubescence short, inconspicuous, appressed. Head (fig. 8) with eyes inconspicuous from above but in reality well formed, far down on side of head (almost twice their height from vertex), with about 30 small facets; vertex with a pair of minute foveae, diameter less than that of an ocular facet, and not united by an interfoveal sulcus; an obliquely placed oval, imbricated tumulus above each antennal articulation; frontal margin between tumuli gently arcuate and medianly bearing a mat of short, stiff setae; front slightly transversely excavated and then joining a large clypeal rostrum; posterior margin of rostrum bearing a short median cusp; ventral surface of head with a few aciculate setae; maxillary palpi 4- segmented, 1 minute, 2 elongate-pedunculate, 3 short and rounded-triangular, 4 longer than 2 and elongate-conical with an oblique base. Antennae 11-segmented, distantly articulated, segments 1 and 2 rather large and elongate subequal; 3 to 7 small and moniliform; 8 about the same as seventh but narrowed apically; 9 wider than eighth and transverse-trapezoidal; 10 of same shape as ninth but wider; 11 largest, one-fourth longer than wide, conical and as long as 6 to 10 united. Pronotum (fig. 8) with anterior two-thirds orbicular and basal third constricted; a small fovea each side visible from above, and a large median impression united by sulcoid depressions; disc simply convex. Elytra with rounded oblique humeri; elytron bifoveate with a depression extending posteriorly from each fovea (as in MaUeoceps); entire sutural stria from mesial foveal depression; flank with a weak subhumeral fovea. Abdomen with five visible tergites in median length ratio of 1.2/1.0/1.0/0.6/0.8 with first three bearing strong lateral margins; first tergite with minute basal abdominal carinae separated by almost 30 per cent of segmental width; last tergite rounded-triangular. Seven visible stemites in median length ratio of 0.6/1.2/0.6/0.4/ 0.2/0.8/0.2 with sixth medianly flattened. The seventh sternite is so small, and articulated so evenly with the sixth, that it may be overlooked. It is a small semicircular aedeagal plate externally, fitting into a semicircular notch in the sixth (actually this sternite is larger and partially membranous, and most of it lies beneath the sixth sternite). This plate is quite obvious when the aedeagus is exserted (fig. 9), when it is seen to swing to the morphological right. The aedeagus (fig. 9) is 0.22 long X 0.08 wide and is quite similar in general anatomy to that of Malleoceps (fig. 5). Legs slender, unmodified, tarsi with a single claw. Metastemum gently sulcoid medianly. Metacoxae slightly separated. Female essentially as for male with the following exceptions: (1) head just discemibly wider than pronotum, as a consequence of her small eyes of only about 14 facets; vertexal tumuli absent; vertexal foveae larger than in the male; face gently declivous, the front and clypeus continuous and simple; (2) only six visible stemites. Described from nine males and six females (type male and 12 paratypes in the collections of the Illinois Natural History Survey; 84 FIEUDIANA: ZOOLOGY, VOLUME 68 two paratypes in author's collection). Collected May 28, 1950 at Shada Headquarters near Refuge, Haiti, by H. B. Mills. This species appears to be allied to Malleoceps darlingtoni Park (1954) on the basis of the comparative anatomy of the aedeagus. Hispanisella hirsuta Park, new species Description. - Type male: Head 0.22 long x 0.26 wide; pronotum 0.21 x 0.25; elytra 0.36 X 0.45; abdomen 0.34 x 0.40; totol length 1.1. Cuticula shining reddish brown, sparingly micropunctulate; pubescence very short, appressed and inconspicuous except for special areas noted later. Head (fig. 10) not including eyes, rounded-triangular; eyes conspicuous, subcircular, at their own height from vertex. Vertex greatly modified: medianly, behind the eyes, it is elevated into a prominent tumulus. This tumulus is declivous posteriorly to the arcuate occiput; it is declivous anteriorly and here bears a patch of stiff setae, and this declivity terminates in an acute tooth. Medianly, between the eyes, the vertex is deeply transversely excavated beneath the acute tooth, with a vertexal fovea either side of excavation. Anterior to this excavation the vertex is less excavated and bears medianly a tubercle which is surmounted by about five contorted setose processes, unusually like the organ on the male vertex of MeUeoceps pearcei (fig. 6). Anterior to this organ the frontal margin narrows and bears between the antennae an abrupt, high, setose roll. Posterio- lateral portions of the head concave, produced, and bearing oblique rows of very short, stiff, truncate setae. Ventral margin of head nearly glabrous, with a few aciculate setae. Antennae 11-segmented, distantly articulated; segments 1 and 2 elongate; 3 to 10 much smaller and subequally wide, 3 elongate-obconical, 9 slightly wider than eighth and trapezoidal, 10 larger than ninth, trapezoidal; 11 largest segment, conical, two- fifths longer than wide and as long as 6 to 10 united. Pronotum with anterior three-fourths transversely orbicular; a lateral fovea each side, wholly visible from above, at constriction of basal fourth, and these three foveae united by an antebasal sulcus; disc simply convex. Elytra with rounded humeri; elytron deeply bifoveate with entire sutural stria; flank with a just discernible subhumeral foveoid impression. Abdomen with five visible tergites in median length ratio of 1.4/1.0/0.8/0.8/0.8 and first three with strong lateral margins. Seven visible stemites in median length ratio of 0.4/1.4/0.6/0.4/0.2/0.2/0.8, last stemite a transversely oval aedeagal plate with acute lateral angles. Mesocoxae in confiuent cavities; metacoxae slightly separated; meta trochanters with ventral faces laminoid. Legs slender, with tarsi bearing a single claw. Described from a unique male, the type (deposited in the collection of the lUinois Natural History Survey) and collected at Tombeau Cheval, Haiti on May 24, 1950 by H. B. Mills. This species also shows close affinity with Malleoceps; it appears to be most closely allied to H. haitiana. PARK ET AL.: PSELAPHID BEETLES 35 Figs. 7-9. 7. Ramelbida quadrifoveata (Raffray), dorsal view of head of female type (modified from Raffray, 1903). 8. Hispanisella haitiana n. g. and sp., dorsal view of head and pronotum of male. 9. Hispanisella haitiana n. g. and sp., a, aedeagus exserted, as seen on a point-mount, with aedeagal plate swung to morphological right, b, aedeagus as seen through a slide-mount which had been in cold 15 per cent KOH for two days. Genus Dalmomelba Park Dalmomelba Park, 1954, Amer. Mus. Nov., 1674, p. 6. Type-species: Dalmomelba cazieri Park, loc. cit., p. 6. Dalmomelba cazieri Park, 1954. South Bimini (Bahamas). Genus Trimiosella Raffray Trimiosella Raffray, 1898, Rev. d'Entomol., 17, p. 236. Type-species: Trimiosella anguina (Reitter), 1883, Deut. Entomol. Z., 27, p. 42. Trimiosella anguina (Reitter) loc. cit., St. Thomas (Virgin Islands). Haasiella Park, new genus Trimiina having the following combination of structural features: (1) Head narrower than pronotum; a pair of vertexal foveae between eyes; eyes well developed; ventral surface of head with aciculate setae; (2) antennae 11-segmented, moderately closely articulated on either side of the frontal margin; segment 10 bilaterally symmetrical; (3) maxillary palpi 4 -segmented; segment 1 minute; 2 36 FIELDIANA: ZOOLOGY, VOLUME 68 elongate pedunculate; 3 slightly longer than wide and ovoidal; 4 largest, conical with truncate base; (4) pronotum with disc simply convex and base not bisected by a median vertical carina; (5) elytral humeri rounded; elytron bifoveate; flank with a subhumeral fovea; (6) abdomen with five visible tergites and seven visible stemites in male (only sex known); (7) mesocoxae in confluent cavities; metacoxae distantly articulated; (8) tarsi with a single claw. Type-species: Haasiella medicina Park, new species. This new genus is named in honor of Dr. Glenn E. Haas, of the U.S. Public Health Service. Haasiella medicina Park, new species Description, - Type male: Head 0.17 long X 0.17 wide; pronotum 0.71 x 0.21; elytra 0.31 x 0.34; abdomen 0.31 x 0.31; total length 0.9. Cuticula shining reddish brown; micropunctulate; pubescence scant on head, rather abundant on pronotum, elytra, and abdomen but very short. Head (flg. 11) rounded-triangular in dorsal view, with flattened eyes of about 30 facets, the eyes ovoidal, longer than wide; tempora shorter than eyes; a pair of vertexal foveae between eyes, each fovea set in a large depression and these depressions narrowing rapidly to form a complete interfoveal sulcus; frontal margin between antennae a rounded-triangular lobe which is then declivous to clypeolabral suture; posterior margin of this frontal roll triangularly impressed medianly, and bounded posteriorly by interfoveal sulcus; occiput medianly indented and this indentation extended anteriorly to an acute point between vertexal foveae; under surface of head bearing a few aciculate setae. Antennae 11-segmented; moderately approximate on either side of the triangular frontal roll; segment 1 a truncated triangle, with base slightly wider than apex, base slightly wider than segment is long; 2 longer than first and ovoidal; 3 to 8 much smaller, moniliform; 9 very slightly wider than eighth, transverse-moniliform; 10 obviously wider than ninth, trapezoidal; 11 forming the club, largest segment, three- fifths as wide as long, ovoidal and about as long as segments 3 to 10 united. Pronotum with anterior two- thirds transversely orbicular with the narrower posterior third sharply set off by the antebasal sulcus (reminiscent of the pronotal outline of the North American Dalmosella Casey (1897, p. 570) and Lemelba Park); disc simply convex; a large, deep median antebasal impression which is joined briefly by a right and left lateral antebasal sulcus; these sulci extending down the flanks of the pronotum. Elytra with rounded humeri; elytron deeply bifoveate; entire sutural stria and a short discal depression; flank with a subhumeral fovea and longitudinal carina. OpposUe: Figs. 10-16. 10. Hispanisella hirsuta n. g. and sp., a, dorsal view of head of male, b, top detail of vertexal tubercle. 11. Haasiella medicina n. g. and sp., dorsal view of head. 12. Sandersonella transversa n. g. and sp., dorsal view of head of male. 13. Lemelba davisi Park, dorsal view of male (after Park, 1953). 14. Lemelba hirsuta n. ■p.. dorsal view. 18. Actium opacum Park, dorsal view of male (after Park, 1960). 16. Actium haaai n. sp., dorsal view of male. 37 38 PIELDIANA: ZOOLOGY, VOLUME 68 Abdomen with Ave visible tergiteB in median length ratio of 1.0/0.8/0.8/1.0/0.4 with first three having narrow lateral margins and the last rounded-triangular. Apparently seven visible stemites in median length ratio of 0.4/0.6/0.4/0.4/0.2/1.0/0.2 with third stemite bearing each side a glabrous, egg-shaped tumulus; sixth having a subcircular impression at center of posterior margin; seventh minute and hardly discernible. Mesocoxae in confluent cavities; metastemum deeply and broadly concave; legs with metacoxae widely separated by a transverse distance equal to the median length of the third tergite; femora medianly swollen; mesotibiae wider medianly and at this point bearing a small spine on the mesial face; tarsi with a single short claw. Oescribed from a unique male in author's collection. Collected by Glen E. Haas from berlesate of forest floor debris on Big Tree Trail at 1,750 ft. elevation in the Luquillo Forest, Puerto Rico on December 5, 1959. This species is most closely allied to species of Lemelba but lacks the median, vertical, antebasal carina of this genus. It has no known close allies. There are two ambiguous items. The antebasal pronotal sulci cross the flanks and, obscured by pubescence, may be slightly deeper and wider to form a "lateral pronotal fovea." Second, the seventh stemite in the unique type is obscured by ejecta and may not be present. Only further examination of additional material can settle this question, but it is believed that this stemite is present. Sandersonella Park, new genus Trimiina having following combination of structural features: (1) Head (fig. 12) slightly wider than pronotum; vertex with a pair of foveae between the eyes; ventral surface of head with capitate setae: (2) antennae U-segmented, articulated on either side of a broad, short frontal rostrum with segment 10 bilaterally symmetrical and 11 forming the antennal club; (3) maxillary palpi 4-segmented, segment 1 short, 2 elongate-pedunculate; 3 rounded-triangular and short; 4 largest, subcorneal; (4) pronotum with a small antebeisal fovea each side which is slightly visible form above, and a larger median fovea, all three united by an antebasal sulcus; (5) elytral humeri obliquely rounded, elytron bifoveate at base and a weakly formed subhumeral fovea on flank; (6) male (only sex known) with five visible tergites and seven visible stemites; (7) mesocoxae in confluent cavities and metacoxae slightly separated; (8) tarsi with a single claw. Type-species: Sandersonella transversa Park, new sp)ecies This genus has no close neotropical allies and is diffit ilt to place in the tribe Euplectini. The single-segmented antennal club places it in the subtribe Trimiina. The capitate setae allies it with Latomelba, Quadrelba, and AUomelba in so far as the ventral surface of the head is concerned. The rather closely articulated PARK ET AL.: PSELAPHID BEETLES 39 antennae on the short, broad rostrum gives a head outUne which is unique in the fauna under consideration. This genus is named in honor of Dr. Milton W. Sanderson, of the Illinois Natural History Survey. Sandersonella transversa Park, new species Description. - Type male: Head 0.15 long x 0.19 wide; pronotum 0.17 x 0.17; elytra 0.26 X 0.27; abdomen 0.21 x 0.26; total length 0.8. Cuticula shining yellowish brown, subimpunctate except for special areas noted later; pubescence very short and inconspicuous. Head (flg. 12) with vertex organized in two areas, e.g., a very transverse basal portion, and a short, broad rostrum; eyes are hardly visible from above, yet they are well-developed, of about 20 small facets, and far down on the sides of the head; tempore very short, about one-half of eye length; the transverse basal part of the vertex has at each antero-lateral angle a small, circular tumulus (reminding one of the larger, oval tumuli of Hispanisella haitiana Park); the central part of this basal portion bears a Y-shaped sulcoid impression, and between the stem of the Y and each tumulus is a vertexal fovea. The rostrum is limited laterally by the antennal incisures, and its base is bounded by the arms of the Y-shaped impression; this rostrum is surmounted by a small tubercle and then becomes declivous anteriorly. Ventral surface of the head short and bears about 12 capitate setae. Maxillary palpi described in the generic diagnosis. Antennae 11-segmented, articulated on either side of the rostrum and so relatively approximate, being separated by the width of the last antennal segment; segment 1 elongate; 2 as wide but orbicular; 3 to 8 much smaller, moniliform; 9 slightly wider; 10 distinctly wider than ninth, trapezoidal; 11 largest segment, similar to Melba, as long as segments 4 to 10 united. Pronotum with anterior two-thirds suborbicular, posterior third suddenly narrower; a minute lateral fovea each side of constriction, just visible from above and its position obscured by pubescence; a relatively large median fovea which is connected with lateral foveae by a narrow antebasal sulcus; disc convex and unmodified. Elytra with rounded humeri, but with the subhumeral area slightly set off and approaching the condition found in Actium Casey; elytron deeply bifoveate; entire sutural stria; discal impression nearly to middle of elytral length; flank with a very weakly developed subhumeral foveoid impression. Abdomen with five visible tergites in a median length ratio of 1.0/0.8/0.6/0.6/0.6/ with three bearing lateral margins, first with minute basal abdominal carinae separated by about 25 per cent of segmental width; fifth tergite vertical. Seven visible stemites in a median length ratio of 0.2/0.8/0.2/0.2/0.2/0.009/0.6, the seventh in the form of a transversely ovate aedeagal plate. Mesocoxae in confluent cavities; metasternum convex; metacoxae slightly separated by half of the length of their mesial faces. Legs short, slender; tarsi with a single, short claw. Described from one male (the type in the collection of the 40 FIELDIANA: ZOOLOGY. VOLUME 68 Illinois Natural History Survey), collected on May 24, 1950 at Tombeau Cheval, Haiti by H. B. Mills. Genus Ramelbida Park Ramelbida Park. 1942. Northwestern Univ. Stud. BioL Sci. Med.. 1. p. 112. Type-species: Ramelbida quadrifoveata (Raffray). 1903. Ann. Soc. Entomol. France, 72. p. 537. Ramelbida quadrifoveata (Raffray), St. Thomas (Virgin Is- lands) Genus Hanfordia Park Hanfordia Park, 1960, Trans. Amer. Microscop. Soc, 79 (1), p. 10. Type-species: Hanfordia absoluta Park, loc. cit., p. 10. Hanfordia absoluta Park, beneath bark of dead branches on Cooper's Hill, Red Hills, St. Andrew Parish, Surrey County, Jamaica. Genus Lemelba Park Lamelba Park. 1953, Nat. Hist. Misc., 121. p. 1. Type-species: Lemelba davisi Park, loc. cit., p. 2. This genus was described from a single species within the neotropical tip of peninsular Florida, and in the key which follows the type-species is included although it has not been recorded from the West Indies. Key to Species of Lemelba 1 Special setae of the abdomen relatively thin-shafted, very arcuate, and often coiled at tip; a small cusp at center of pronotum which almost touches the median bisecting antebasal carina (fig. 14). hirsute Park, n. sp. Special abdominal setae slightly arcuate, thick-shafted, not apically coiled; pronotum without cusp 2 2(1) Eyes almost invisible from a direct dorsal view; metacoxae not contiguous but separated by a transverse distance that equals about one-fourth the metastemal length; type species known from the Florida Everglades (fig. 13) davisi Park Eyes visible from above; metacoxae separated by a transverse distance that equals about one-half the metastemal length millsi Park, n. sp. Lemelba millsi Park, new sp>ecies Description. - Type male: Head 0.12 long x 0.17 wide; pronotum 0.17 X 0.21; elytra 0.29 x 0.34; abdomen 0.19 X 0.29; total length 0.8. PARK ET AL.: PSELAPHID BEETLES 41 Cuticula shining yellowish brown; subimpunctate; general body pubescence short but abundant; abdomen with thick-shafted, slightly arcuate special guard setae, which probably are special tactile receptors. Of these latter there appear to be one or two on the lateral margin of first tergite, about six in a transverse row of the second, third, and fourth tergites, and none on the fifth tergite. Head rounded-triangular, with a pair of eyes that are slightly visible from above, subcircular in lateral view, of about 30 very small ocular facets; a pair of vertexal foveae between the eyes united by a weakly formed interfoveal sulcus; occiput medianly with a broadly indented area; ventral surface of head with about 10 capitate setae. Antennae U-segmented, distantly articulated, segment 1 rounded-triangular, slightly wider at base than long; 2 elongate, slightly larger than first; 3 to 8 much smaller, third subobconical, fourth to eighth submoniliform; 9 slightly larger than eighth, trapezoidal; 10 distinctly larger than ninth, trapezoidal; 11 largest segment, truncate-conical, as long as segments 5 to 10 united, apical truncature bearing a circlet of long setae. Pronotum with anterior three-fourths transversely orbicular, the basal fourth suddenly narrower and bearing a straight transverse antebasal sulcus which ends each side in a foveoid impression but these latter not obvious from above, probably as a consequence of the pubescence; basal fourth medianly bisected by a strong carina from basal bead to the anterior margin of the sulcus; disc simple. Elytra with rounded humeri; elytron bifoveate with entire sutural stria from sutural fovea, and discal fovea in an impression; flank with weakly formed subhumeral fovea. Abdomen with five visible tergites bearing the remarkable special setae as noted previously, and first three tergites with strong lateral margins. Seven sternites in median length ratio of 0.7/0.34/0.09/0.08/0.009/0.17/0.26 with seventh a transversely ovoidal aedeagal plate. Metastemum broadly and deeply concave. Metacoxae separated by about one-half of the metasternal length. Profemora and mesofemora inflated, especially the latter. Three-segmented tarsi bearing a single, relatively thick tarsal claw. Described from a unique male, the type, deposited in the Illinois Natural History Survey and collected by Dr. Harlow B. Mills, in whose honor this species is named. Collected June 4, 1950, near Cienfuegos, Soledad, Cuba. This new species is most closely allied to the type-species, Lemelba davisi Park (1953, pp. 1, 2), known only from the Florida Everglades where it is relatively abundant in floor debris. Lemelba hirsuta Park, new species Description. - Type female: Head 0.19 long x 0.21 wide; pronotum 0.21 X 0.21; elytra 0.31 x 0.38; abdomen 0.28 x 0.33; total length 1. Cuticula shining reddish brown; subimpunctate; pubescence sparse but long, giving a shaggy appearance to the insect. In addition there are very long, recurved, 42 FIELDIANA: ZOOLOGY. VOLUME 68 hyaline special setae on pronotum, elytra, and abdomen. Some of these setae on the abdomen are 0.07 nun. long, not including the often recoiled tips, and if the latter are included some about 0.1 mm. long, or half as long as the head (fig. 14). Head rounded-triangular with the eyes readily seen from above, the eyes with about 34 ocular facets and in lateral view about twice as long as the tempora; a pair of deep impressions in each of which is a small vertexal fovea, these foveae united by an interfoveal sulcus; occiput medianly indented, this narrowing to a subacute point on the vertex between the vertexal foveae; face simply declivous, convex; ventral view of head reveals 16 capitate setae. Antennae as described for millsi. Pronotum with shape as noted for millsi but a lateral fovea is partially visible from above at each end of the transverse antebasal sulcus; additionally, a short, acute cusp briefly overhangs the sulcus just above, but does not touch the median bisecting carina (fig. 14). Elytra as described for millsi except for the shaggy pubescence noted previously, and relatively much broader. Abdomen with five visible tergites, with strong lateral margins and proportions as shown in Figure 14. Six visible stemites in a median length ratio of 0.03/0.9/0.3/0.2 /0. 1/0.8 with the last rounded-triangular. Metacoxae well separated by a transverse distance which is almost one-half the metastemal length. Three-segmented tarsi, with an arcuate tarsal claw. Described from a single female, the type specimen, in the Museum of Comparative Zoology. Collected by Dr. P. J. Darlington in July, 1938 at Sanchez, Dominican Republic. This new species is unique in the genus so far by the character of its recoiled tergal pubescence. It is not closely allied to either davisi or millisi. The genus Lemleba is of interest zoogeographically in that the Floridian and Cuban species are obviously closely allied in body shape and special setae, and this is one of the very few known instances where faunal affinity can be shown between neotropical Florida and the West Indies, despite the relatively short distance between these areas. Genus Actium Casey Actium Casey, 1886, Bull. California Acad. Sci., 2 (6), p. 201. Type-species: Actium californicum (LeConte), 1878, Proc. Amer. Phil. Soc., 17, p. 383. California, U.S.A. (See revision of Actium by Grigarick and Schuster, 1971.) Key TO Species OF /I cftum Cuticula shining and finely punctulate visendum Parii Cuticula subopaque, with head, pronotum, and elytra finely granulated 2 PARK ET AL.: PSELAPHID BEETLES 43 2 (1) Eyes large, of many small ocular facets, and obvious from a dorsal view; frontal margin in the form of an elevated frontal roll (fig. 15). opacum Park' Eyes only slightly visible from above, small, and of only seven ocular facets; frontal roll triarcuate (fig. 16) haasi Park, n. sp.' Actium haasi Park, new species Description. — Type male: Head 0.10 long x 0.15 wide; pronotum 0.16 x 0.17; elytra 0.21 X 0.22; abdomen 0.31 X 0.23; total length 0.8. Cuticula yellowish brown, only slightly shining to subopaque as a consequence of microgranulate-punctulate surface of head, pronotum, and elytra; pubescence short and inconspicuous. Head rounded- triangular; eyes only slightly visible from above, subcircular in lateral view, small, consisting of seven facets; a pair of vertexal foveae united by a narrow sulcus; frontal margin elevated into a triarcuate roll, as a consequence of the elevated antennal acetabulae (fig. 16); occiput slightly indented medianly; ventral surface of head with a few capitate setae. Antennae 11-segmented, relatively distant, segments 1 and 2 elongate but the first not obvious as it is deeply inserted; 3 to 8 much smaller, subequal in width, third slightly obconical, others submoniliform; 9 slightly larger than eighth; 10 obviously larger than ninth; 11 largest segment, elongate-ovoidal and about as long as 3 to 10 inclusive. Pronotum with simple, convex disc; a biarcuate antebasal sulcus connecting a lateral fovea each side fully visible from above. Elytra with subhumeral indentation on each side; elytron bifoveate, with the anterior rim of each fovea elevated, entire sutural stria, flank with a subhumeral fovea beneath the subhumeral indentation each side. Abdomen with five visible tergites in relative proportions as illustrated (fig. 16); first three tergites with distinct lateral margins; a pair of very short basal abdominal carinae which are separated by slightly less than one- third of total segmental width. Seven sternites in median length ratio of 0.03/0.07/0.07/0.06/0.03/0.06/0.03 with the seventh in the form of a transverse aedeagal oval palte which is asymmetrically articulated to the right side. Third stemite bearing on each side a tumulus. Legs slender; three-segmented tarsi, the third segment bearing a single claw. Described from a unique male, the type, in the author's collection. This new species was collected by Dr. Glenn E. Haas, in whose honor this species is named. It was collected from forest floor debris near Stone Watch Tower at an elevation of 2,850 ft., in the Maricao State Forest, Puerto Rico on January 24, 1960. It is most closely related to opacum Park, from Jamaica, by the 'Figures 15 and 16 show the integumental granulation as a series of semicircles. The general effect seems too coarse and has not pleased either the author of this paper or his artist. 44 FIELDIANA: ZOOLOGY. VOLUME 68 character of its integumental ornamentation, but is quickly discriminated from this species by its rudimentary eyes and character of the frontal roll. Genus Latomelba Park Latomelba Park. 1955. Bull. Chicago Acad. Sci., 10 (7). p. HI. Type-species: Latomelba quadrisicca Park, loc. cit.. p. 112. Morce's Gap. St. Andrew Parish, Jamaica at 5.500 ft. elevation. In this genus males have seven sternites whereas females have six sternites; additionally, the males have the third sternite variously modified, and the fourth sternite bears a pair of remarkable spatulate setae on each side. Key to male.s of the species of Latomelba Lateral third of third sternite each side bearing a tumulus surmounted by a facet-like nipple, and laterad of this tumulus is a dense circlet of erect setae, the tips of which are recurved inwardly to form a structure resembling a miniature bird cage. quadrisicca Park Lateral third of third sternite each side bearing a weakly-formed, simple tumulus, and laterad of this tumulus is a weakly-formed depression sympatrica Park Genus Quadrelba Quadrelba Park, 1955, Bull. Chicago Acad. Sci., 10 (7), p. 116, 1942. Northwestern Univ. Stud. Biol. Sci. Med., p. 120. Type-species: Quadrelba parmata (Reitter), 1883, Deut. Entomol. Z., 27. p. 40 (Trimiopsis). St. Thomas, Virgin Islands, and Puerto Rico. Males with seven visible sternites, the seventh a subcircular aedeagal plate which is asymmetrically articulated to the sixth sternite; female with six visible sternites, the sixth rounded- triangular. Key to males of the species of Quadrelba 1 Vertex between the auriculate preocular angles bearing a simple, free median tubercle parmata { Reitter) Vertex not as described 2 2(1) Vertex deeply, entirely sulcate behind the thin, carinoid preocular angles; front projecting posteriorly into this sulcus medianly as a triangular field; this field bears a pair of approximate foveae, and terminates in a raised .setose angle inconspicua (Reitter) Vertex not as described; the large head has slightly reflexed preocular angles which are simply expanded ventricosa (Reitter) PARK ET AL.: PSELAPHID BEETLES 45 Genus Allomelba Park Allomelba Park, 1954, Amer. Mus. Nov., 1674, p. 16. Type-species: Allomelba antennata Park, loc. cit., p. 17. Allomelba antennata Park. South Bimini Island, Bahamas. Genus Zolium Casey Zolium Casey, 1897, Ann. New York Acad. Sci., 9, p. 560. Type-species: Zolium eggersi (Reitter) 1883, Deut. Entomol. Z., 27, p. 38 (Trimiopsis). Described from St. Thomas, Virgin Islands. Collected by and named for Herrn von Eggers. Zolium eggersi (Reitter), loc. cit., see checklist for complicated taxonomic history. Genus Trimiovillus Park Trimiovillus Park, 1954, Amer. Mus. Nov., 1674, p. 11; emended Park, 1960, Trans. Amer. Microscop. Soc, 79 (1), p. 18. Type-species: Trimiovillus bahamicus Park, loc. cit., p. 12, South Bimini Island, Bahamas. Key to species of Trimiovillus Vertexal foveae pubescent bahamicus Park Vertexal foveae nude bellingeri Park Genus Melba Casey Melba Casey, 1897, Ann. New York Acad. Sci., 9, p. 565. Type-species: Melba thoracica (Brendel), 1889, Entomol. Amer., 5, p. 196, (Trimium), Iowa, U.S.A.; designated by Bowman, 1934, p. 144. Males have seven visible stemites, the seventh a subcircular aedeagal plate asymmetrically articulated to the sixth stemite. Females have six visible stemites, the sixth rounded subtriangular. Key to We.st Indian Subgenera of Melba 1 Vertex with four free foveae; an anterior pair near the antennal bases, and a posterior pair between the eyes; no interfoveal sulcus. Melba (Rameloidea) Park Vertex not as described 2 2 (1) Vertex with three free foveae; a pair between the eyes, and a median fovea on a line between posterior ocular margins; no interfoveal sulcus. Melba (Cismelba) Park Vertex with one pair of foveae between the eyes; a more or less entire interfoveal sulcus present 3 46 FIELDIANA: ZOOLOGY. VOLUME 68 3 (2) Interantennal margin of the front gently arcuate to subtruncate between antennal bases 4 Frontal margin extended more or less above the face (more pronounced in males, but distinct in females) Melba (Frontelba) Park 4 (3) Elytral flank with an oblique line which starts near nuddle of lateral margin and extends obliquely dorso-posteriorly to apical margin Melba {Melba) Casey Elytral flank with a longitudinal line which is parallel to lateral margin from near humeral area to apical margin Melba (Perimelba) Park Melba Subgenus Rameloidea Park Rameloidea Park, 1942, Northwestern Univ. Stud. Biol. Sci. Med.. 1, pp. 117-118. 121. Type-species: Melba temporalis Raffray, 1909, Ann. Soc. Entomol. Prance, 78, p. 16. M. (Rameloidea) temporalis Raffray. Martinique (Windward Islands), type locality; Antigua (Leeward Islands) Melba Subgenus Frontelba Park Frontelba Park. 1942. Northwestern Univ. Stud. Biol. Sci. Med., 1, pp. 117-118, 120; 1952. Chicago Acad. Sci.. 9 (2). pp. 135. 140. Type-species: Melba frontalis Raffray. 1908. Ann. Soc. Entomol. France. 77, p. 35. Guadeloupe. Leeward Islands. Key to species of Melba (Frontelba) Front slightly produced above clypeus in a semicircular margin, this margin medianly roughened (dissecting binocular at 70 X, and microscope whole-mounts at 430 x show this roughening to be caused by 10 to 12 subvertical microstruts of the cuticula) clypeata (Reitter) Front produced between antennal bases, this extension bearing three tubercles in a triangular pattern .frontalis Raffray Melba Subgenus Melba Casey {sensu Casey, 1897) Key to males of species of Melba (Melba) 1 At least second and third sternites relatively simple, not bearing tubercles, or foveae, or conspicuous depressions. 2 These sternites all or in part modified by tubercles, foveae, or depressions. 4 2(1) Second to fifth sternites subequally long crassipes Raffray Seconds to fifth sternites progressively shorter 3 3 (2) Aedeagal plate (seventh stemite) irregularly and transversely rhomboidal; vertexal foveae oblong ^renadensis Raffray PARK ET AL.: PSELAPHID BEETLES 47 Aedeagal plate subcircular; vertexal foveae circular; mesotrochanters each bearing a conspicuous spine zonula Park 4 (1) Third sternites bearing a tubercle on each side specularis (Reitter) Third stemite impressed on each side, this impression medianly subtubercu- late ^ibbula (Reitter) Melba Subgenus Perimelba Park Perimelba Park. 1943. Bull. Chicago Acad. Sci., 7 (3), p. 183. Type-species: Melba granulosa Park, 1943, Bull. Chicago Acad. Sci., 7 (3), p. 184, Victoria, Tamaulipas, Mexico. Key to Species or Melba (Perimelba) Females with basal abdominal carinae of first tergite prominent, nearly half as long as segment and separated by about one-third of segmental width (male unknown). jamaicensis Park Females with these carinae very short, one-sixth of segmental length and with about the same separation (male sex with remarkable cephalic structures) (q.v. literature) hoffi Park Melba Subgenus Cismelba Park Cismelba Park, 1960, Trans. Amer. Microscop. Soc, 79 (1), p. 16. Type-species: Melba trifoveata Park, loc. cit., p. 16. M. (Cismelba) trifoveata Park. Hanover Parish, Cornwell County, Jamaica. There remains in the West Indian fauna a species unknown to the author which cannot be placed in a subgenus. This is Melba fleutiauxi (Raffray), 1890b, Bull. Soc. Entomol. France, (6) 10, pp. 204-205, collected on Guadeloupe, (Leeward Islands). Tribe BRACHYGLUTINI Brachyglutini Raffray, 1908, Genera Insectorum, 64th Fascicle, p. 190. Key to Subtribe.s of Brachyglutini 1 Ventral surface of head with a median ovate fossa. DECARTHRONINA Ventral surface of head otherwise 2 2 (1) Ventral surface of head with a median, longitudinal carina or carinoid ridge, this feature is usually strong but may be weak, and may be entire or interrupted; body usually obviously pubescent 3 Ventral surface of head medianly simple, lacking any trace of a median carina or carinoid ridge; body usually glabrous EUPSENIINA 48 FIELDIANA: ZOOLOGY, VOLUME 68 3 (2) Each elytron beaing a long, distinct discal stria; this stria varies in length between species and genera from nearly as long as elytron to about half as long BRACHYGLUTINA Elytra usually without any trace of a dLscal stria or impression; rarely each elytron may bear a vague, short intrahumeral impression or stria which mav be from a fifth to a third of the elytral length. PSELAPTINA Subtribe PSELAPTINA Park, new subtribe Type-species: Pselaptus belfragei LeConte, 1880, Trans. Amer. Entomol. Soc, 8, pp. 184-185, Texas, U.S.A. In this subtribe are placed those genera having the following combination of structural characters: (1) ventral surface of head bearing a median, longitudinal carina which may be a sharp ridge or keel but may be a low carinoid hne; (2) antennae in the West Indian species 11-segmented; (3) antebasal pronotal and elytral foveae vary as between genera: they may be distinct and conspicuous, or very poorly developed in which case they are represented by just discernible pits, or they may be absent; (4) elytra usually lacking any discal stria or impression; on rare occasions each elytron may bear a vague, short intrahumeral impression or stria which may be from a fifth to a third of the elytral length; (5) metacoxae obviously separated by the length of a metatrochanter or more; (6) abdomen with a narrow but distinct margin on each side of the first tergite, and often on the first three tergites. This subtribe has caused considerable trouble for many years. The taxonomy is difficult. This is not because the species populations are poorly defined, but rather the early generic concepts were couched in such vague or brief generalizations that type species could not be recognized in many cases until the type specimens had been examined. Consequently much confusion arose. New species were described in unsuitable genera; often specimens in collections were masquerading under established specific names but were undescribed species. Inevitably this led to a complicated synonymy. Most of the type specimens of species in Pselaptina that were described by Schaufuss, Reitter, Raffray, and Sharp have been examined by the author in the spring of 1951. I am grateful to the late Dr. Ren6 Jeannel and M. Guy Colas of the Museum National d'Histoire Naturelle in Paris, and Dr. E. B. Britton and Mr. W. E. China of the British Museum of Natural History for placing at my PARK ET AL.: PSELAPHID BEETLES 49 disposal the facilities of their respective museums. Quite a bit of the taxonomic material which follows in this subtribe reflects this first- hand examination of European collections. Kky to Gknkra of Pski.aptina 1 Maxillary palpal segment 3 (penultimate) of exceptional size, nearly as long as segment 4, elongate-ovoidal, very flattened and with the lateral face bearing a dense mat of short pubescence (fig. 17). Bythinogaster Schaufuss Maxillar\- palpal segment 3 otherwise 2 2 (1) Maxillar>' palpal segment 4 (ultimate) with a tooth or tubercle or angle on the lateral face, this process may be rounded or acute, and may occur near base or apex of segment (fig. 18) Berdura Reitter Maxillary palpi otherwise 3 3 (2) Mesosternal process ending posteriorly in a wide, truncate to broadly rounded margin between mesocoxae; this process is usually distinctly transverse, but may be only slightly wider than long and lacks a median mesosternal carina of any kind Scalenarthrus LeConte Mesosternal process usually distinctly longer than wide, often ending in a fine, acute point and may or may not bear a median carina or keel. Pselaptus LeConte Genus Berdura Reitter Berdura Reitter, 1881, Verb. Natur. Ver. Brunn, 20, p. 187. Type-species: Berdura excisula Reitter, 1883, Deut. Entomol. Z., 27, p. 3.5, St. Thomas, Virgin Islands. Key to sprcies of Berdura Distal segment of maxillary palpus with the latero-distal area of external face rounded (fig. 18) leavitti Park, n. sp. This area of maxillary palpus acute (fig. 19) excisula Reitter Berdura leavitti Park, new species Description. - Type female: Head 0.12 long x 0.12 wide; pronotum 0.275 x 0.31; elytra 0.45 X 0.84; abdomen 0.45 X 0.45; total length 1.3. Cuticula shining yellowish brown; lightly micropunctulate; pubescence short, appressed, longer on abdomen. Head rounded-quadrate with eyes visible from above, subcircular in lateral outline, of about 24 facets; vertex convex and simple, with a pair of small vertexal foveae on a line with anterior eye margins, and united by a weak interfoveal sulcus; frontal margin slightly convex and face simple and unmodified; ventral surface of head with a median longitudinal carina, and on each side an anteriorly converging carina; on sUde mount at 430 x each mandible is seen to bear a small boss or tubercle. Maxillary palpi as illustrated (fig. 18). 50 FIELDIANA: ZOOLOGY, VOLUME 68 20 a 20 b Figs. 17-20.17. Bythinogaster simplex Schaufuss, maxillary palpus, type-species from Haiti (after Park, 1960). 18. Berdura leavitti n. sp., distal segment of maxillary palpus. 19. Berdura excisula Reitter. distal segment of maxillary palpus. 20. Pselaptus longiclai'a (Schaufuss), a, antennal segments 9-10-11 of male, b, antennal segments 9-10-11 of female. Antennae 11-segmented, distantly articulated; segments 1 and 2 large, elongate- subcylindrical; 3-7 much smaller, subequally wide, third slightly obconical, others subquadrate; 9 wider than eighth and barrel-shaped or elongate-cylindrical with slightly convex lateral margins; 10 slightly longer, and slightly bell-shai>ed; 11 largest segment, about as long as segments 7 to 10 united. Pronotum with rounded sides; a small round antebasal fovea on each flank; a median antebasal fovea not apparent at 70 x , and the two lateral foveae appear to be united by an evanescent crease, but so slightly that these foveae may be said to be "free" for practical purposes; disc simply convex. Elytra with rounded humeri and also appear to lack antebasal foveae at both 70 X and 430 x , although at 70 x there appear to be several short arcuate ridges near the anterior elytral margins; such features disappear at 430 X; sutural stria entire; no discal impression. Metathoracic wings present. Abdomen with five visible tergites in median length ratio of 0.14/0.08/0.08/ 0.07/0.1.'> with last two tergites subvertical; very narrow margins on first three tergites; ba-sal abdominal carinae very short and separated by slightly more than one- fourth of the total segmental width. Five visible sternites. Legs slender and unmodified. Tarsi .3-segmented, 1 minute, 2 and 3 very long and elongate cylindrical, the third more slender and bearing a single claw. PARK ET AL.: PSELAPHID BEETLES 51 Described from two females. The type, deposited in the Museum of Comparative Zoology, collected by Dr. P. J. Darlington on November 28, 1926 at Cienfuegos, Soledad, Cuba. The paratype, in author's collection, was collected by B. B. Leavitt, in whose honor this species is named, in Soledad, Cuba on July 1, 1932. The nearest ally is Berdura excisula Reitter, the type-species from St. Thomas, Virgin Islands. Genus Scalenarthrus LeConte Scalenarthrus LeConte, 1880, Trans. Amer. Entomol. Soc, 8, p. 135. Type-species: Scalenarthrus homi LeConte, loc. cit., Arizona, U.S.A. Key to males of the species of Scalenarthrus 1 Distal antenna] segment (11) elongate-arcuate, as long as the remaining 10 segments united, with an oblique fossa in basal half clavatus Raffray Distal antennal segment otherwise 2 2(1) Antennal segment 11 almost as wide as long, and almost divided by a deep incision which is two-thirds the width of segment, located in basal half, with apical edge straight and proximal edge oblique, the internal faces lightly pubescent jamaicensis Park Antennae otherwise 3 3 (2) Antennal segment 2 elongate but with the dorsal face tumid, segment 10 transversely asymmetrically triangular, the narrow internal face trun- cate; segment 11 about as wide as 10, unmodified, elongate acute- oviform ^uadelupensis Raffray Antennae otherwise 4 4 (3) Antennal segment 2 simply elongate-cylindrical; segment 10 transversely trapezoidal, segment 11 wider than 10, with truncate base, unmodified, elongate-oviform tomentosus Park Antennae highly modified; segment 2 much larger than 1 with distal-mesial area tuberculate; segments 7-8-9 much wider than 6, irregularly shaped and with distal-mesial margins fringed with long setae. pectinicornis Raffray Genus Pselaptus LeConte Pselaptus LeConte, 1880, Trans. Amer. Entomol. Soc, 8, p. 184. Type-species: Pselaptus belfragei: LeConte. loc. cit., p. 185. Texas, U.S.A. Key TO SPECIES of Pselaptus 1 Distal antennal segment (11) fusiform, as long as the preceding four segments united jternalis Raffray 52 FIELDIANA: ZOOLOGY. VOLUME 68 Distal antennal segment (11) fusiform, of exceptional length, with an arcuate apex, much longer than the four preceding segments united 2 2 (1) This distal segment (11) very elongate-arcuate and six times the length of the tenth segment (fig. 20a) male longiclava (Schaufuss) This segment almost four times as long as tenth segment and less arcuate (fig. 20b) female /on^i'c&ti^ (Schaufuss) Genus Bythinogaster Schaufuss Bythinogaster Schaufuss, 1887, Tijdschr. Entomol., 30, p. 111. Type-species: Bythinogaster simplex Schaufuss, loc. cit., p. 112, "Hispaniola (probably Haiti as the citation is 'Saint- Domingue')" Park, 1942. So far as known this genus is endemic to the Greater Antilles. Furthermore, the genus can be discriminated by the unique third segment of the maxillary palpus which is large, almost as long as the fourth (distal) segment, elongate-ovoidal in outline, very flattened latero-mesially, and bearing on its lateral face a dense mat of very short setae (fig. 17). The aedeagus varies greatly as between the five known species, and yet has that collective aspect of general anatomy which suggests that Bythinogaster is a natural generic aggregate. All five species have four spines (which are black-tipped for identification in the illustrations) associated with the membranous internal sac, and in each case these spines have a qualitatively different distribution. All five species have a lateral process or falx on the morphological left side; four have a flax on the morphological right side. Illustrations of the aedeagei are all drawn from the dorsal aspect with the distal (posterior) aedeagal margin toward the top of the page so that the morphological left side appears on the right side of the figure, and should be kept in mind when using the key to species. In all but one species the author dissected out the aedeagus, and so the organ could be mounted and drawn at 430 diameters. The type of B. simplex Schaufuss had the aedeagus fully exserted, and was drawn at 70 diameters magnification. The falces in this genus are very thin and it is probable that the right falx of B. simplex was present, but so amalgamated with the organ that it could not be discovered. It should be noted that the thin falces tend to have their distal ends twisted or altered after a period of years, probably as a consequence of the differential tensions set up within their own organization or as a reaction to the drying of the Canada balsam; but the four spines noted are remarkably stable. PARK ET AL.: PSELAPHID BEETLES 53 Key to species of Bythitwgaster based on aedeac.i 1 Left falx curved or inclined to the morphological right 2 Left falx curved or inclined to the morphological left 4 2 (1) Left falx suddenly arcuate, with its tip expanded as a flat, ovoidal disc (fig. 23) nymphoides Park Left falx with its tip narrowly arcuate and acute 3 3 (2) The four black-tipped spines of the internal sac (fig. 25) in an oblique row to the morphological left .parsonsi Park, n. sp. The four spines not such an arrangement (fig. 22). bisphaeroides Schaufuas 4 (1) Left falx narrowing distally to an acute and recurved apex (fig. 21). simplex Schaufuss Left falx distally biarcuate, not narrowing very much at apex which is broadly arcuate (fig. 24) diaboUcus Park Alternate key to species of Bythinogaster BASED ON THE FRONTO CLYPEAL AREA OF THE MALE SEX 1 Frontal margin on a line through dorsal antennal articulations, not medianly tuberculate 2 This interantennal frontal margin medianly bearing a tubercle which is surmounted by a pair of anteriorly inclined tufts of setae. parsonsi Park, n. sp. 2(1) Front subvertically declivous, this declivity microgranulate-alutaceous, then front terminates in a pair of apically directed, mesially setose tubercles; on each side of this overhanging margin the face extends laterally in an angular ovoidal wing which forms a part of the antennal acetabulum; beneath the median frontal margin the face is flattened, setose and extends semicircularly to the semilunate clypeus. nymphoides Park Face otherwise 3 3 (2) Frontal margin between antennae angulate and vertex just behind this margin microalutaceous; below the margin the face is laterally smooth and declivous, and medianly is extended apically as a longitudinally carinoid lobe; this lobe ends in a pair of small membranous processes; beneath this distal lobe the clypeus bears a median conical tubercle. diabolicus Park Face otherwise 4 4 (3) Interantennal line of front simple, declivous; face narrows with declivity ending in a rounded-subtriangular point which bears a tuft of setae; beneath this point the front and clypeus are deeply excavated transversely; clypeus transverse and tumid simplex Schaufuss 54 FIELDIANA: ZOOLOGY, VOLUME 68 Figs. 21-26. 21. Bythinogaster simplex Schaufuss, aedeagus (after Park, 1960). 22. Bythinogaster bisphaeroides Schaufuss, aedeagus (modified from Park, 1960). 23. Bythinogaster nymphoides Park, aedeagus (after Park, 1960). 24. Bythinogaster diabolicus Park, aedeagus (after Park, 1960). 25. Bythinogaster parsonsi n. sp., aedeagus. 26. Reichenbachia eucera (AuW), aedeagus, dorsal face. The four spines have been blackened at tips for discrimination; the dehydrated and irregular compressor muscles of the basal bulb have been stippled; the dorsal membrane of basal bulb is shown in light circular stipple. Interantennal line of front simple and declivous; the distal margin of the clypeus is narrowed into two rounded-triangular points, and between these points bears a pair of anteriorly-directed elongated, obtuse dentoid processes bisphaeroides Schaufuss Bythinogaster parsonsi Park, new species Description. - Type male: Head 0.26 long x 0.34 wide; pronotum 0.34 x 0.34; elytra 0.63 x 0..55; abdomen was damaged so that no accurate measurements could be obtained. Total length, based on other known species of the genus probably would PARK ET AL.: PSELAPHID BEETLES 55 be between 1.7 and 1.8 long, and with a maximum width probably between 0.5 and 0.6. Cuticula polished yellowish brown; subimpunctate and subglabrous. Head with large prominent eyes, subcircular in lateral view, and of 22 large facets; a pair of small vertexal foveae, each closer to its adjacent eye than to each other, and obviously smaller than an ocular facet; no interfoveal sulcus; vertex simply and gently convex; interantennal frontal margin medianly elevated into a tubercle; this tubercle surmounted by a pair of tufts of setae which are slightly inclined anteriorly; face glabrous and declivous and bearing medianly a low tubercle; face narrows anteriorly and the margins limited by a carinoid ridge to about the fronto-clypeal area, where these carinoid ridges become slightly elevated and recurved; clypeus flattened beneath these carinoid ridges, the flattened area also carinoid at the perimeter; anterior clypeal margin three-sided, the median side truncate; ventral surface of head with a short median ridge. Antennae distantly articulated, 11-segmented; segments 1 to 7 longer than wide, 1 relatively large, 2 shorter and narrower; 3 to 7 obviously smaller than 2; 3 to 5 subequal; 6 as long as 5 but slightly narrower; 7 slightly shorter than 6; 8 shorter than 7, subquadrate; 9 slightly wider than 8, subcircular; 10 obviously larger than 9, subcircular; 11 largest segment, oviform. Pronotum lacking antebasal foveae per se, a faintly darkened vestigial impression each side; disc simply convex. Elytra lacking antebasal foveae per se, but with a pair of vestigial impressions at base; a weak, entire, strioid sutural impression; humeri rounded. Metathoracic wings well-developed. Abdomen with five visible tergites and five visible stemites, but the abdomen was slightly crushed diagonally so that measurements of segmental proportions were not attempted; first three tergites with very narrow margins. Legs slender and not modified; tarsi long, slender, 3-segmented with the last bearing a single tarsal claw. Metacoxae well separated. Aedeagus as illustrated (fig. 25). Described from an unique male, the type deposited at the Museum of Comparative Zoology. It was collected at Soledad (Cienfuegos), Cuba during the period May-June, 1939 by C. T. Parsons, in whose honor this species is named. This species does not appear to have any close allies by virtue of the interantennal tubercle; geographically it is most closely allied to B. bisphaeroides Schaufuss. Oddly this latter species was not included in Raffray's Coleopterorum Catalogus (1911) and I did not know of it and so it went unlisted in my (1942) treatment of neotropical Pselaphidae. What I consider to be B. bisphaeroides Schaufuss is represented by a single male, in the Museum of Comparative Zoology, collected at light by L. L. Scaramuzza at Baragua, Cuba, August 2, 1927. Subtribe BRACK VOLUTIN A Raffray (s. str.) Brachyglutina Raffray, 1908, Genera Insectorum, 64th Fascicle, p. 108. 56 FIELDIANA: ZOOLOGY, VOLUME 68 At the present time three genera are listed in this subtribe in the West Indies. These are Achillia, Bryaxis, and Reichenbachia. Genus AchUlia (Reitter, 1890) This genus was named in honor of the distinguished French pselaphidologist, Achille Raffray (Reitter, 1890, p. 212). It is a Chilean genus, and by 1911 Raffray (pp. 87-88) listed 24 species from Chili, and Achillia excisa Schaufuss (1879, p. 494) from Cuba! Such a disjunct distribution was noted by Park (1942, p. 154). Fortunately, Jeannel (1962, p. 408) redescribed A. excisa Schaufuss from the type in the Paris Museum, and stated that it is very doubtful if the species came from Cuba, and without doubt it was a Chilean species. Therefore, I propose that this genus, Achillia be removed from the West Indian faunal list. Genus Bryaxis (Kugelann, 1794) This genus, largely of the Palaearctic Region, has had one of the most complicated taxonomic histories in the family Pselaphidae, being placed in different tribes, and fragmented and discussed by many people who have described species in the genus. In a faunal paper such as this one any further discussion would be out of place. What is relevant here is that Schaufuss described two species of Bryaxis from Cuba: Bryaxis bisinuata Schaufuss, 1887, p. 130 Bryaxis truncata Schaufuss, 1886, p. 132 Raffray (1911, p. 104) places them in Reichenbachia, but notes that he regards this generic assignment as doubtful. Park (1942, pp. 152-153) could not place either species in any group of Reichen- bachia. I do not know these two species and cannot interpret their characters from the original descriptions. Therefore, I propose that this genus Bryaxis be removed from the West Indian faunal list until authentic specimens of each can be located and redescribed. Genus Reichenbachia Leach Reichenbachia Leach, 1826, Zool. Jour., 2, p. 451. Type-species: Reichenbachia juncorum (Leach), loc. cit., p. 452, Europe and Algeria. PARK ET AL: PSELAPHID BEETLES 57 Reichenbachia is well-known and widely distributed. It is probably the largest genus in Pselaphidae, and is cosmopolitan, with the possible exception of Australia and New Zealand. For a general discussion of the neotropical fauna see Park (1942, pp. 135- 153). Key to the species of Reichenbachia This key is based on the male sex unless otherwise noted, and deals primarily with antenna 1 characteristics. 1 Antennal segment 10 abnormally large in both sexes, as wide as or much wider than the last (11) segment 2 Antennal segment 10 normal, not obviously wider or larger than segment 11 5 2 (1) Antennal 10 distinctly wider than 11, distinctly obconcial with truncate base narrower than truncate apex, and sides divergent .3 Antennal segment 10 slightly wider than 11 4 3 (2) Antennal segment 11 oblong with truncate base and acute apex; segments 7, 8, 9 very transverse (fig. 28) ma\e grenadensis Raffray Antennal segment 11 ovate, with a much narrower base; segments 7, 8, 9 less transverse female grenadensis Raffray 4 (2) Antennal segment 10 square, with mesial face rounded. male vincentania Raffray Antennal segment 10 nearly twice as long as 9, and only a little wider. female vencentania Raffray 5(1) Antennal segment 6 of male with ventral face concave, bearing a tuft of setae on its mesio-posterior angle (fig. 27) eucera (Aub^) Antennal segment 6 of male with mesial face bearing an apically setose tubercle or blunt spine 6 6 (5) Antennal segment 6 with a blunt, elongate tubercle arising from the concave ventral face and projecting from the anterior third of segment (fig. 29) darlingtoni Park, n. sp. Antennal segment 6 with a very narrow base and an irregularly swollen anterior two thirds; a short, blunt tubercle arises from the mesial face near the base of the segment (fig. 30) ^uadelupensis Raffray Reichenbachia darlingtoni Park, new species Description. - Type male: Head 0.34 long x 0.39 wide; pronotum 0.41 x 0.43; elytra 0.60 x 0.86; abdomen 0.60 x 0.77; total length 2.0. Cuticula dull reddish brown, punctulate, more obviously on the elytra; pubescence short and appressed. Head with a pair of prominent eyes, subcircular in lateral view, of about 34 large ocular facets; three vertexal foveae, a pair between the eyes and one between the 58 FIELDIANA: ZOOLOGY. VOLUME 68 antennal tubercles, each fovea set in a depression so that medianly the vertex is slightly tumid; tempora about as long as eyes; face simply declivous; ventral surface of head with the usual median carinoid ridge. Antennae as illustrated (fig. 29). Pronotum with a large pubescent antebasal fovea each side and a smaller, nude, elongate median antebasal fovea; disc simply convex. Elytra with rounded humeri; elytron bifoveate, an entire sutural stria from sutural fovea and a discal stria from discal fovea; discal stria inclines mesially to end at slightly less than distal three-fourths of elytral length; flank simple. Abdomen with five visible tergites in median length ratio of 3.8/2.0/1.3/1.2/0.8 with first three strongly margined; basal abdominal carinae short and separated by about 37 per cent of total segmental width. Five sternites medianly visible but the mount of the specimen makes measurement of proportional median lengths unsatisfactory. First stemite relatively very long, second and third much shorter; fourth almost invisible medianly; fifth stemite large and medianly flattened. Metastemum medianly concave; metacoxae well separated; mesofemora especially thicker; tarsi with a long single claw. Described from two males. The type, deposited in the Museum of Comparative Zoology, was collected by Dr. P. J. Darlington, Jr. in whose honor this species is named, in mountains north of Imias, eastern Oriente, Cuba, between 3,000 and 4,000 ft. elevation, July 25 to 28, 1936. The paratype male, in author's collection, was collected by L. C. Scaramuzza in grasses in Upper Yara Valley, Cuba, October 18, 1928. Subtribe DECARTHRONINA Park Decarthronina Park, 1951, Geol. Surv. Alabama, Mus. Pap., 31, p. 61. Genus Decarthron Brendel Decarthron Brendel, 1865, Proc. Entomol. Soc. Philadelphia, 5, p. 30. Type-species: Decarthron formiceti (LeConte) 1850, Boston Jour. Nat. Hist., 6, p. 90, Penn., N.J., and Gulf States, U.S. Designated by Bowman, 1934, p. 144. This is the only genus in the subtribe in the West Indian fauna. There are six known species, all in the subgenus Decarthron s. str. Brendel, (1865, p. 30). The genus is quickly discriminated since both sexes bear on the ventral surface of the head a conspicuous median oval fossa or fovea, and have 10-segmented antennae. At present only the male sex can be keyed out. This sex has numerous secondary modifications of the antennae, and legs, and the aedeagus is species-specific and subject to great variation in PARK ET AL.: PSELAPHID BEETLES 59 27 b 29 a Figs. 27-30. 27. Reichenhachia eucera (AuW), male antenna, a, dorsal face, b, ventral face of segment 6. 28. Reichenhachia grenadensis Raffray, male antenna (after Raffray, 1904 and modified from Park, 1942). 29. Reichenhachia darlingtoni n. sp., antenna, a, dorsal face, b, ventral face of segment 6. 30. Reichenhachia guadelupensis Raffray, male antennal segments 5 and 6 (modified from Raffray, 1908). species details. The females lack such modifications. In the following key to males only five of the six known species can be discriminated. The sixth species, D. unifoveolatum (Schaufuss) from Cuba, is known only from the type female. [Park examined the Decarthron types in the Paris Museum and the detail in the key which follows is drawn from his examination. J.W.— M.W.S.] Key to the males of species of Decarthron Profemora with dorsal surface simply swollen in distal three-fourths of length, but the swelling evenly rounded and not marked by carinoid ridge, oval field, spine or other modifications spinosum Raffray Profemora with distal declivity of dorsal face modified: slightly compressed or bearing a carinoid ridge or an oval field or other modification 2 60 FIELDIANA: ZOOLOGY. VOLUME 68 2(1) Mesofemur excavated, the excavation, as seen from posterior face, very short (about one-eighth of femoral length and almost one-half of femoral depth); two teeth on dorsal face of mesofemur, one at proximal edge of excavation, and one behind this tooth, both teeth triangularly acute. insula re Raff ray Mesofemoral modifications otherwise 3 3 (2) Mesofemoral excavation bearing a short, wide and truncate spine at proximal margin of excavation 4 Mesofemoral modifications very different, mesofemur greatly inflated in distal three-fourths of length where it is abruptly limited at distal seven- eighths; excavation of this inflation subtriangular in profile; near upper edge of excavation the contour is interrupted by an arcuate, lamelloid, spine which continues as a long arcuate seta; aedeagus (fig. 31). arcuatum Park, n. sp. 4 (3) Profemur inflated and bearing a flat, elongate-oval granular patch that is 33 per cent as long as femur and placed obliqiely on the dorso-anterior face; aedeagus (fig. 33) vauriei Park Profemur inflated and bearing a very narrow, elongate-oval field (thus field is subgranular and three times longer than wide) that is 6 per cent as long as femur and placed on the dorsal face of the distal declivity; aedeagus (fig. 32) pectinate Park, n. sp. Decarthron arcuatum Park, new species Description. - Type male: Head 0.33 long x 0.34 wide; pronotum 0.33 X 0.34; elytra 0.52 x 0.62; abdomen 0.52 X 0.60; total length 1.7. Cuticula shining yellowish brown; head, pronotum and elytra subimpunctate, elytra punctate; pubescence longer on elytra and abdomen. Head with prominent eyes, subcircular in lateral view, each of about 28 large ocular facets; tempora short, about a sixth of lateral eye-length; a pair of nude vertexal foveae on a line through anterior third of eyes, free; face simple, longitudinally declivous, slightly tumid. Antennae distantly articulated, 10-segmented, unmodified; club of last three segments; 8 transverse, larger than 7; 9 transversely oblong, larger than 8; 10 largest segment, nearly as long as 8 and 9 united, suboviform. Pronotum simple, with a small nude median antebasal fovea. Elytra with rounded humeri; elytron bifoveate, foveae nude; entire sutural stria; weak discal impression to about median point of elytral length; flank simple. Abdomen with five visible tergites in median length ratio of 3.0/1.0/1.0/1.0/1.0 with first three laterally margined; basal abdominal carinae of first tergite slightly divergent and 0.7 as long at tips as segment and separated at tips by about 50 per cent of total segmental width. Five visible stemites in median length ratio (taken from a paratype male) of 1.1/1.0/0.4/0.4/0.8 with distal margin of fifth medianly indented. Profemur modified: dilated dorso-ventrally and bearing medianly a short, high carinoid lamella on distal declivity. Mesofemur modified: greatly inflated in distal PARK ET AL.: PSELAPHID BEETLES 61 Figs. 31-33. 31. Decarthron arcuatum n. sp., aedeagus, dorsal face. 32. Decarthron pectinate n. sp., aedeagus, dorsal face. 33. Decarthron vauriei Park, aedeagus, dorsal face (after Park, 1954). three-fourths of length where it is abruptly limited at distal seven-eighths of mesofemoral length; in profile this inflation excavated, the latter subtriangular in profile; near the upper edge of this excavation its contour is interrupted by an arcuate, lamelloid spine which is continued as a still more arcuate seta; aedeagus (fig. 31). Described from nine specimens (type male and eight para types, of which the type and four paratypes are deposited in the Museum of Comparative Zoology, and four paratypes in the collection of the author). The type was collected by B. B. Leavitt in 1932 in central Cuba; he also collected two males in central Cuba in August, 1932. One male was taken by P. J. Darlington, Jr. in Cienfuegos, Soledad, Cuba in May, 1936. Five were obtained by C. T. Parsons in Cienfuegos, Solidad, Cuba in June-July, 1939. This new species appears most closely allied to D. pectinale, also from Cuba. The aedeagi of D. pectinale (fig. 32) and vauriei, (fig. 33) are more similar in basic features than is either to D. arcuatum (fig. 31). D. vauriei was described from South Bimini, Bahamas. Decarthron pectinale Park, new species Description. - Type male: Head 0.26 long x 0.30 wide; pronotum 0.40 x 0.43; elytra 0.51 x 0.69; abdomen 0.43 x 0.69; total length 1.6. Cuticula shining reddish brown; subimpunctate; pubescence longer on elytra and abdomen. 82 FIELDIANA: ZOOLOGY. VOLUME 68 Head with prominent eyes, subcircular in lateral view, each of 28 large ocular facets; pair of free vertexal foveae on a line passing through second row of facets; tempora short, about one-sixth of lateral eye- length. Antennae distantly articulated, lO-segmented; unmodified; club of last three segments: 8 sUghtly larger than 7 and subquadrate-subcircular; 9 distinctly longer than 8, transversely subcircular; 10 oviform, nearly as long as preceding three segments united. Pronotum and elytra as described for D. arcuatum. Abdomen with five visible tergites in median length ratio of 3.2/1.0/1.0/0.07/1.0 with first three laterally margined; basal abdominal carinae divergent, a little more than half as long as first tergite and separated at tips by about 40 per cent of total segmental width. Five visible sternites in median length ratio of 3.0/0.2/0.2/0.2/0.04 with the last slightly concave and with distal margin elevated. Profemur modified: medianly moderately dilated and bearing a very narrow, elongate-oval field (this field is subgranular and three times longer than wide), and varies from 6 per cent to 10 per cent as long as femur; this narrow field is placed on the distal declivity of dorsal face. Mesofemur modified: inflated at distal three-fourths of length, this distal declivity excavated and bearing a short, truncate spine at proximal margin of the declivity. Aedeagus (fig. 32). Described from 11 males (type male and 10 paratypes, of which the type and five paratypes are deposited in the Museum of Comparative Zoology, and five paratypes in the author's collection). The type was collected by B. B. Leavitt March 1, 1932, in Soledad, Cuba. One paratype was collected by L. C. Scaramuzza, February 8, 1927, at light at Baragua, Cuba. Three were taken by C. T. Parsons at Cienfuegos, Soledad, Cuba. The other six were collected by B. B. Leavitt: two at Santa Clara, Soledad, Cuba, July to September, 1932. This new sjjecies at present appears most closely allied to D. arcuatum, also from Cuba, but the male leg modifications and aedeagi quickly separate them. The aedeagus of D. pectinale (fig. 32) appears structurally to be more allied to the aedeagus of D. vauriei (fig. 33) described from the Bahamas. Subtribe EUPSENIINA Park Eupaeniia Park, 1951, Geol. Surv. Alabama, Mus. Pap., 31, p. 61. Genus Eupsenius LeConte Eupsenius LeConte. 1850, Boston Jour. Nat. Hist.. 6 (1), p. 90. Type-species: Eupsenius glaber LeConte, 1850. loc. cit.. South Carolina, U.S.; Park (1954) lists Southern U.S., and South Bimini, Bahamas. PARK ET AL.: PSELAPHID BEETLES 63 This is the only genus of the subtribe in the West Indies and is easily discriminated. The ventral surface of the head is unmodified, lacking either the fossa of Decarthronina, or the median carinoid ridge of Brachyglutina. Furthermore, the cuticula is glabrous except for special areas. This polished, glabrous, convex body, an unusually large last antennal segment and large vertexal foveae form a habitus quickly discerned. In the key that follows one West Indian species, Eupsenius dominicanus Schaufuss (1887a, p. 108), is excluded. The authenti- city of its habitat is not in question. The species was represented by an unique specimen which lacked both antennae {teste Raffray, 1896, p. 265) and was included in the genus by Raffray (1911, p. HI) with reservation. Unfortunately, the species of Eupsenius have few secondary sex characters or other major differences, so that the antennae become of great importance in species separation. On the other hand, included in the following key is Eupsenius rufus LeConte (1863, p. 28) which is known from Florida and Alabama and may occur in the Bahamas since the type species Eupsenius glaber (1850, p. 90) is known from several southeastern states and the Bahamas (Park, 1954). Key TO THE West Indian and North American species of Eupenius 1 Visible portion of antennal segment 10 twice as large as the visible portion of antennal segment 9 .gracilis Raffray Visible portion of antennal segment 10 not this large, usually slightly wider and longer than antennal segment 9 but never twice as long as 9. 2 2(1) Distal (11) antennal segment relatively long and slender (including the closely appressed pubescence, slightly more than twice as long as its greatest width) 3 Distal (11) antennal segment relatively shorter and thicker (including the closely appressed pubescence, never twice as long as its greatest width, usually from one-fourth to two-fifths longer than wide) 4 3 (2) No median pronotal antebasal fovea or foveoid impression of any kind at center of transverse antebasal line .politus Reitter A minute median fovea at center of antebasal line .pretiosus Park 4 (2) Pronotum rather suddenly narrowed at the transverse antebasal line, the sides subparallel from this line to basal margin, and the greatest pronotal width being related to the basal margin, and the greatest pronotal width being related to the basal third width in a ratio of about 5 to 4. glaber LeConte 64 FIELDIANA: ZOOLOGY. VOLUME 68 Pronotum gradually narrower from the maximum width to basal margin, the ratio of maximum width to basal third about 5 to 4.75 (not known to occur in West Indies) rufus LeConte Tribe CTENISTINI Raffray Ctenistini Raffray. 1890a. Rev. d'Entomol.. 9, pp. 140, 141. This is the first time that a member of this tribe has been reported from the West Indies. Genus Ctenisodes Raffray Ctenisodes Raffray, 1896, Ann. Soc. Entomol. France. 65, p. 274. Type-species: Ctenisodes laticeps Raffray, loc. cit., p. 275, Mexico. Ctenisodes weberi Park, new species Description. — Type male: Head including cervicum 0.41 X 0.41 wide; pronotum 0.34 X 0.41; elytra, including squamous posterior fringe 0.69 x 0.72; abdomen 0.63 X 0.72; total length 2. Cuticula shining reddish brown but obscured by the white masses of squamous pubescence. Most of the face, antennal incisures, cephalic foveae, anterior and fKJSterior comers of pronotum, pronotal and elytral foveae, posterior edges of elytra, and intersegmental abdominal conjunctivae are densely covered. The general and rather sparse scales of pronotum, elytra and abdomen are short, flat and arcuate. The amount of pubescence has been reduced intentionally in Figure 40 so as not to obscure many anatomical features. Head with very large eyes of about 24 ocular facets; three pubescent foveae on the top of the head, the usual pair between the eyes and one at base of antennal tubercle; the head narrows suddenly anterior to eyes and then expands to form an antennal tubercle; face excavated beneath the tubercle, the excavation crowded with white, massed pubescence in the upper portion, except for a very narrow, vertical lamina at center. Lower portion of the face simple and extended laterally and ventrally each side. Labrum short and very transverse. Maxillary palpi as illustrated (fig. 34). Antennae approximate on the antennal tubercle, very long, being about half as long as body (1); 11-segmented: 1 quadrate; 2 slightly shorter than 1 and quadrate; 3 to 7 subequal in width and elongate; 8 quadrate and slightly longer than 7; a poorly differentiated club of the last three segments; 9 about as long as 8. slightly wider; 10 about as long as 9 and slightly wider; 11 elongate, slightly wider than 10 at base and increasingly wider apically, slightly shorter than segments 7-10 united. Pronotum and elytra as illustrated (fig. 40). Abdomen with five normally visible tergites, the first three strongly margined. The anterior margin of the first visible tergite also with a heavily pubescent anterior margin, but in life probably is covered by the heavily pubescent posterior elytral margins; fifth tergite vertical. Seven stemites, the seventh in the form of a minute median plate which is just discernible. PARK ET AL.: PSELAPHID BEETLES 65 Figs. 34-37. 34. Ctenisodes weberi n. sp., right maxillary palpus, ventral view. 35. Neotyrus tobagoensis n. sp., ventral view of right maxillary palpus. 36. Hamotus (Hamotus) soror Raffray, mesial view of right maxillary palpus. 37. Apharus hexagonus n. sp., mesial view of right maxillary palpus. Legs slender, not modified except that the ventral surfaces of protrochanter and profemur bear erect, stiff setae. Tarsi 3-segmented, the third with a pair of strong tarsal claws. Metastemum medianly sulcate. Described on an unique male, the type, deposited in the Museum of Comparative Zoology. It was obtained by Berlese funnel by N. A. Weber, in whose honor this species is named, near Cienfuegos, Soledad, Cuba, August 6, 1920. Its only ally is Ctenisodes laticeps Raffray, (1897, p. 275) the type species and until now the only known species. C. laticeps was described from Mexico. Some time ago I obtained a female of laticeps from Montemorelos, Nuevo Leon, Mexico which was collected with ants by Dr. Charles Seevers. I compared this specimen with the type in the Paris Museum. The unique maxillary palpi of Ctenisodes (cf. Park, 1942, pp. 291-294) illustrated in the present paper (fig. 34) and general pubescent pattern are similar in laticeps and weberi but laticeps is smaller, more slender, with a relatively longer abdomen. The antennal segment proportions are also different, but in this tribe sexes frequently vary in this matter. 66 FIELDIANA: ZOOLOGY, VOLUME 68 Tribe TYRINI Raffray Tyrini Raffray, 1890a. Rev. d'EntomoL. 9. p. 162. Key to Genera of Tyrini Distal (fourth) segment of maxillary palpi bearing short, stiff rod-like setae (fig. 41b). Ephimia Reitter Distal (fourth) segment of maxillary palpi nude, or bearing minute, appressed setae if discernible Neotyrus Raffray Genus Ephimia Reitter Ephimia Reitter, 1883, Deut. Entomol. Z., 27, p. 34. Type-species: Ephimia simoni Reitter, loc. cit., St. Thomas, Virgin Islands. This genus has had a complicated taxonomic history — too complex to go into here. The major references are as follows. Raffray (1904, 1908, 1911) placed Ephimia in the tribe Hybocepha- lini. Park (1942, p. 289) followed this course, but noted that it could hardly belong in the HybocephaUni since it was the only genus in the tribe with a non-bilobed second tarsomere and having two equal tarsal claws, and consequently would key out to either the tribes Ctenistini or Tyrini. Also in the 1942 paper Park erected the genus Juxtahamotopsis (p. 303) in the tribe Tyrini. Juxtahamotopsis is a synonym of Ephimia. Jeannel (1949) erected the tribe Odontalgini and included Ephimia, and in a later classification of Pselaphidae (1955) maintained this position. But not all species of Ephimia have squamous pubescence on the sides of the head, so the genus cannot be placed in Odontalgini. Blackwelder (1944, p. 95) places Ephimia in the tribe Hybocephalini and included all of the Ctenistini. Ephimia cannot be in Ctenistini as the general body pubes- cence is aciculate, not squamous. Ephimia is placed in the tribe Tyrini in the present paper until more thorough information can be accumulated. The genus is strictly Neotropical, and the last segment of the maxillary palpus (fig. 41b) is diagnostic. Key TO SPECIES of Ephimia 1 Pubescence on sides of head squamous 2 Pubescence on sides of head not squamous 3 2 (1) Antennal segments 3 to 8 very transverse and lenticular, or transversely bean-shaped simoni Reitter PARK ET AL.: PSELAPHID BEETLES 67 Antennal segments 3 to 8 transverse-trapezoidal cubensis Park, n. sp. 3(1) Antennal segments 9 and 10 quadrate, subequally long; metasternum concave male subnitida Raffray Antennal segment 9 transverse, 10 subquadrate, nearly twice as long as 9; metasternum convex female su6mhrfa Raffray Ephimia cubensis Park, new species Description. - Type (male?): Head 0.29 long x 0.27 wide; pronotum 0.36 x 0.34; elytra 0.43 x 0.60; abdomen 0.50 x 0.60; total length 1.6 (fig. 41a). Cuticula shining reddish-brown with paler appendages; sides of head both anterior of eyes and along genal margin bearing squamous setae, antennal segments with subsquamous setae, general body pubescence bearing long, aciculate, thinly distributed setae, subimpunctate. Head only slightly longer than wide; eyes large and coarsely faceted; antennal rostrum bisected by a longitudinal sulcus; a pair of free, pubescent vertexal foveae in a shallow transverse depression between rostrum and convex vertex. Maxillary palpi with distal (4) segment as for genus (fig. 41b). Antennae 11-segmented, rather closely articulated on either side of rostrum; segments 3 to 8 transverse, trapezoidal; club well-formed, of last three; 9 quadrate with length and width subequal; 10 longer than wide in a ratio of 5/4; 11 as long as 9 and 10 united, slightly less than two times longer than wide, ovoidal. Pronotum with strongly convex, simple disc; three large free, pubescent antebasal foveae. Elytra with sloping humeri; elytron with two large pubescent antebasal foveae; entire sutural stria, the raised interval sacrified; short discal impression from each discal antebasal fovea; flank not fovea te but slightly impressed from dorsal view near humeral angle. Abdomen with five visible tergites, the first three with relative proportions as illustrated, fourth subvertical and fifth vertical and not visible from a dorsal view; first three tergites with strong lateral margins. Legs with femora, especially the anterior pair, medianly inflated; tibiae arcuate; tarsi tyrine with two large, subequal tarsal claws. Described from one specimen, the type, which is probably a male, in the Museum of Comparative Zoology. Collected by B. B. Leavitt in 1932 in Soledad, Cuba. The nearest ally of E. cubensis would appear to be E. simoni Reitter (1883) from St. Thomas, Virgin Islands but it differs, among other features, in antennal structure, as noted in the key to species of this perplexing genus. Genus Neotyrus Raffray Neotyrus Raffray, 1895, Ann. Soc. Entomol. France., 64, p. 396. 68 FIELDIANA: ZOOLOGY. VOLUME 68 Type-species: Neotyrus gibbicollis (Schaufuss), 1886, Tijdschr. Entomol., 29, p. 269 {Aplodea), Amazon Basin, Brazil. Neotyrus tobagoensis Park, new species Description. — Type male: Body proportions as illustrated (flg. 38a); total length 2.3. Cuticula shining reddish brown; head coarsely scabropunctate dorsally except for median mstro-vertexal sulcus and vertexal foveal sulci; pronotum subimpunctate; elytra and abdomen with sparse and weakly developed broad punctural impressions; pubescence reddish golden, long and appressed except for the bristling pencil of setae at apex of each genal tumulus. Head with a pair of large eyes comp>osed of very numerous, small facets; a pair of nude, shallow vertexal foveae on a line with posterior eye margins, each fovea in a free, slightly arcuate, glabrous sulcoid impression; these impressions separated by a median sulcus from a point between the vertexal foveae forward, over vertex and most of face; the fronto-rostral portion of this median sulcus becomes declivous, scabro-punctate, and terminates in an arcuate anterior margin; this margin bisects the face as a median, subvertical fronto-clypeal strip with a deep antennal acetabulum on each side; this strip ventrally joins a short, scabro-punctate clypeal margin; labrum with a narrow, glabrous median portion and is laterally subgranulate; mandibles strong, left crossed dorsal to right; maxillar\' palpi typical of genus, and with palpal segments as illustrated (fig. 35); ventral surface of head granulate-punctate with a basal gular fovea; each gena produced in a brief latero- posterior tumulus. Antennae 11-segmented, rather closely articulated on the antennal rostrum, with segmental proportions as illustrated (fig. 38b); club well-formed of the last three segments. Pronotum rounded-hexagonal; a small median, and a large lateral fovea on each side, these three foveae united by a U-shaped antebasal sulcus, which, in turn, lies in a .shallow U-shaped impression; disc convex, becoming elevated just above the median fovea in a prominent, subacute tumulus. Elytra with rounded humeri; elytron bifoveate, an entire sutural stria from sutural fovea, and a long, broad discal impression from discal fovea with the mesial wall of this impression rather abruptly raised; flank not foveate. Abdomen with five tergites but from a dorsal view the very large first tergite comprises almost all of the dorsum; first two tergites with lateral margins obvious; second and third tergites declivous, fourth tergite vertical, fifth tergite inclined ventro-anteriorly. Seven stemites, but the first obscured by pubescence; median length ratio of second to seventh 2.5/0.8/0.8/0.8/1.0/0.6 with the seventh in the form of a small operculum which fits into the posterior margin of the last tergite. Metasternum glabrous; deeply medianly concave, the concavity margined laterally by a carinoid ridge each .side, this ridge erected medianly as a large, triangular tooth; ba.se of metasternum with a large, circular, pubescent fovea. Ix?gs well-developed; protrochanters with a ventro-distal tooth; profemora medianly erected into a glabrous, high carinoid ridge; mesotrochanters with a strong ventro-di.Htal tooth; mesofemora with an obliquely flattened, glabrous, elongate-oval field; tarsi with two claws, one of which is slightly longer and thinner, both acute. PARK ET AL.: PSELAPHID BEETLES 69 Based on a unique male, the type in author's collection. Collected by the Rev. E. J. Pearce on April 4, 1961 in a rotten log near Scarborough, Tobago. The nearest known ally is Neotyrus harem Park (1942, p. 301) from within the nest of the termite Coptotermes niger Snyder, on Barro Colorado Island, Panama Canal Zone. N. harem differs from the above description of tobagoensis in a number of ways: (1) the median cephalic sulcus continues to be glabrous over the face; (2) the vertex and occiput are sparsely and weakly granulate; (3) the vertexal foveae lie in short, oblique sulci which are directed laterally; (4) the clypeus is but weakly granulate. These two species are separated by about 1,300 miles of longitude and tobagoensis should occur in the large Venezuelan fauna; it is wholly different from the Brazilian gibbicollis (Schaufuss) (1886, p. 269) in which the antennae have segments 1 through 9 longer than wide, and 10 obcordate; it differs markedly from coptocolus Park (1942, p. 299) known from the termitarium of Coptotermes niger Snyder on Barro Colorado Island, Panama Canal Zone. N. coptocolus differs from both tobagoensis and harem in having the entire integument opaque and coarsely granulate-asperate, with short, stiff pubes- cence. Tribe HAMOTINI Park Hamotini Park, 1951, Geol. Surv. Alabama, Mus. Paper., 31, p. 67. Key to Gener.ji and Subgenera of Hamotini 1 Distal segment (4) of maxillary palpi with the mesial face bearing a sulcus for distal half only (fig. 37) Apharus Reitter Distal segment (4) of maxillary palpi with mesial face bearing entire sulcus from apex to base (fig. 36) Hamotus Aube 2 2 (1) Pronotum with three free antebasal foveae Subgenus Hamotus A\ib6 Pronotum with the three antebasal foveae united by a sulcus. Subgenus Hamotoides Schaufuss Genus Apharus Reitter Apharus Reitter, 1882, Deut. Entomol. Z., 26 (1), p. 129. Type-species: Apharus mulleri Reitter, loc. cit., p. 130, Sao Paulo, Brazil. Apharus hexagonus Park, new species Description. - Type male: Head 0.26 long x 0.33 wide: pronotum 0.34 x 0..38; elytra 0.50 x 0.79; abdomen 0.46 x 0.64; total length 1.56. 38 a 41 a 70 PARK ET AL.: PSELAPHID BEETLES 71 Cuticula shining dark reddish brown; subimpunctate; pubescence reddish golden; the setae long but longer on elytra, semi-appressed and aciculate. Head rounded trapezoidal with a pair of prominent eyes of fairly large facets; vertex medianly bearing a small, weak, circular impression; a pair of nude, free vertexal foveae each closer to its adjacent eye than to each other; antennal tubercle short, broad and bisected by a broad, longitudinal sulcus; front narrow and vertical; clypeus rounded obtriangular; labrum short and transverse; mandibles strong, left crossed dorsal to right; gular field flattened, glabrous, with a basal gular fovea and each lateral margin extended as a strong, arcuate lamina; maxillary palpi typical of genus, last two palpal segments illustrated (fig. 37). Antennae 0.77 long, 11-segmented; 1 elongate; 2 shorter and quadrate; 3 to 8 transverse sublenticular, shorter than second; club of last three with 9 and 10 trapezoidal, 10 larger than 9; 11 largest, seven times longer than 10, subovoidal. Pronotum rounded-hexagonal, disc convex, a small, free, nude antebasal fovea; the usual pair of lateral foveae are apparently absent. Elytra with rounded, slightly elevated humeri; elytron bifoveate, the sutural at origin of entire sutural stria, the discal at origin of a short, weak discal impres.sion, and the flank simple and not foveate. Abdomen with five visible tergites in median length ratio of 2.6/1.0/1.0/1.0/1.8 with last vertical and rounded rhomboidal; first three margined; five visible sternites (first hidden by pubescence and visible only laterally), the second to sixth in a median length ratio of 1.4/0.6/0.2/0.2/0.25 and slightly flattened medianly. Legs with protrochanters each bearing a ventrodistal spine; profemora each with a tooth, which is inclined anteroventrally, at center of length; tarsi with two claws. Described from a unique male, the type specimen in author's collection. Collected by Rev. E. J. Pearce on April 1, 1961, on the Courland Estate, near Scarborough, Tobago. The nearest ally known is A. armipes Raffray (1891, p. 318) from Caracas, Venezuela. Among other features these two species differ in the male metasternum. In hexagonus the metasternum is medianly sulcoid and the lateral walls are simply tumuloid, whereas in armipes the median sulcus is bounded each side by a large tuberculate tooth. Genus Hamotus Aube Hamotus Aub^, 1844, Ann. Soc. Entomol. France, (2) II, p. 92. Type-species: Hamotus lateritius Aube, loc. cit., p. 92. Venezuela; Colombia. Opposite: Fics. 38-41. 38. Neotyrus tohagoensis n. sp.. a, dorsal view, b, antenna. 39. Fustiger jamaicaensLs n. sp., a, dorsal view, b, outline of antenna, c. truncate tip of last antennal segment .showing pube.scence and presumed sensory organs. 40. Ctenisodes weheri n. sp., dorsal view. 41. Ephimia cuhensis n. sp., a, dorsal view, b, maxillary palpus segments 3 and 4. 72 FIELDIANA: ZOOLOGY, VOLUME 68 This large, chiefly neotropical genus has a characteristic distal segment of the maxillary palpi bearing an entire, longitudinal sulcus on the mesial face (fig. 36). For a general discussion see Park (1942, pp. 308-332). Hamotus Subgenus Hamotus sensu Aub^ Hamotus Raffray. 1904, Ann. Soc. Entomol. France, 74, p. 388; Genera Insectorum, 64th Fascicle, p. 398. Hamotus (Hamotus) soror Raffray, 1904, loc. cit., p. 403. Described from Caracas, Venezuela; also known from Trinidad and Tobago in the West Indies. Hamotus Subgenus Hamotoides Schaufuss Hamotoides Shaufuss, 1887, Berlin. Entomol. Z., 31, p. 298. Hamotus (Hamotoides) hirtus Raffray, 1904, Ann. Soc. Entomol. France 74, p. 408. Described from Grenada (Windward Islands); also now known from St. Vincent (Windward Islands) and Guadeloupe (Leeward Islands). Subfamily CLAVIGERINAE Clavigerinae Redtenbacher, 1849, Fauna Austriaca, pp. 58, 647. Tribe FUSTIGERINI Fustigerini Jeannel, 1949, Mem. Mus. Nat. d'Hist. Nat., N. S.. 29. p. 31. Key to genera of Fustigerini Third (distal) antennal segment with a transverse suture near middle of length. Pseudofustiger Reitter Third (distal) antennal segment entire, lacking such a suture Fustiger Brendel Genus Pseudofustiger Reitter Pseudofustiger Reitter, 1884, Deut. Entomol. Z., 28. p. 167. Type-species: Pseudofustiger stricticornis (Reitter), 1883, Deut. Entomol. Z., 27. p. 33 (ArticeriLs). St. Thomas, Virgin Islands. Key to speciE-S of Pseudofustiger Pronotum wider than long stricticornis (Reitter) Pronotum longer than wide barroi Bierig PARK ET AL: PSELAPHID BEETLES 73 Genus Fustiger Brendel Fustiger Brendel, 1866, Proc. Entomol. Soc. Philadelphia, p. 189. Type-species: Fustiger fuchsii Brendel, loc. cit., p. 190. Tennessee, U.S.A. Kky to species of Fustiger 1 Mesofemora unarmed 2 Mesofemora bearing a very long, arcuate, blunt spine which is about one- third as long as femur male haytiana Mann 2 (1) Pronotum quadrate, as wide as long 3 Pronotum subcorneal, longer than wide male smithi Raffray 3 (2) Head almost three times longer than wide female schwarzi Mann Head only slightly more than two times longer than wide. female jamaicaensis Park, n. sp. Fustiger jamaicaensis Park, new species Description. — Type (female?): Head (including cervicum) 0.48 long X 0.22 wide; pronotum 0.31 X 0.31; elytra 0.55 x 0.77; abdomen 0.72 x 0.78; total length 2. Cuticula shining reddish brown with elytra paler yellowish brown; head weakly and irregularly scabropunctate, rest of body subimpunctate; general body pubescence sparse, short and inconspicuous. Head with proportions as illustrated (fig. 39a); eyes of about 22 rather large facets, slightly subreniform in lateral view as a consequence of a triangular piece of the gena intruding at the ventro-posterior eye margin; a pair of small, free vertexal foveae on a line through posterior eye margins. Antennae 3-segmented as illustrated (fig. 39b); segment 3 (distal) 0.64 mm. long, very elongate-obconical, gradually increasing in diameter to af)ex with an arcuation near distal three- fourths. The truncate end of this segment warrants special notice: this surface is depicted in Figure 39c. The surface is not flush, but concave. When observed directly from above the .scanty, thin, general pubescence is seen around its perimeter; on the edge of the area are about two dozen rod-shaped special setae; scattered over the concavity are numerous small, round to oval bodies, the surfaces of which seem to be dotted, as by small pores. The rod- like setae may be tactile receptors; the oval bodies may be chemoreceptors. Pronotum with a large, gradually deepening median fovea. Elytra with rounded humeri; elytron with no basal foveae apparent, but with an entire sutural stria; no discal impression, and flank unmodified. Abdomen dorsally almost wholly comprised of the large tergal field composed of the first three visible tergites, the margins of which are visible laterally; this tergal field convex for distal three- fourths, and then becoming deeply concave in proximal fourth, beneath the posterior elytral margins, and glabrous; first tergite bearing a pair of trichomes at base (fig. 39a). The golden trichomal brush of .special setae lies on the mesial margin of a cojnplex base which arises from the margin of the segment, each 74 FIELDIANA: ZOOLOGY, VOLUME 68 side. Fourth tergite barely visible from above; fifth (terminal) tergite rounded- triangular. Six visible sternites in median length ratio of 1.0/3.2/1.0/0.8/0.4/1.4 with the second transversely deeply concave at base. Metastemum strongly convex, with a few setae on the crest of the convexity. Legs slender and unmodified by spines or teeth; typical clavigerine tarsi of three segments, the first two very short and the last long, slender and subcylindrical and bearing a single tarsal claw. This new species is based on a single specimen, the type, in the collection of the Illinois Natural History Survey. The sex is in some doubt but is assumed to be a female. It was collected by Milton W. Sanderson by beating and sweeping vegetation in Hardwar Gap, Jamaica on June 25, 1958. The nearest ally to F. Jamaicaensis is also in doubt but Jamaicensis has some affinities with schwarzi Mann from Cayamas, Cuba. CHECKLIST FARONINAE (Jeannel. 1949, p. 15) FARONINI (Raffray. 1890a. p. 82) Coccoplectus Sharp (1887. p. 22) bellingeri Park (1955, p. 101) Jamaica PYXIDICERINI (Raffray. 1903. p. 504) Bythinoplectus Reitter (1881. p. 195; 1883, p. 37) acutangulus Raffray (1904, p. 503) Grenada bahamicus Park (1954, p. 3) Bahamas *foveatus Reitter (1883, p. 37) St. Thomas laminatus Park, n. sp Tobago PSELAPHINAE (Redtenbacher, 1849, p. 57) JUBININI (Raffray, 1903, p. 507) Balega Reitter (1883, p. 43) *elegans Reitter (1883, p. 43) St. Thomas intermedia Park (1960, p. 5) Jamaica longiceps Park, n. sp Dominican RepubHc Stratus Schaufuss (1872, p. 246, 452) dominicanus Park, n. sp Dominican Republic hirsutus Park, n. sp Dominican Republic Jubus Schaufuss (1872, p. 455) clavatus Raffray (1903. p. 511) Grenada insularis Raffray (1908. p. 31) Guadeloupe TANYPLEURINI (Jeannel. 1949. p. 79) DALMODIINA (Park, 1951, p. 61) Insulomodes Park, n. g. *tobagoensis Park, n. sp Tobago, Grenada, Trinidad excavatus Park, n. sp Tobago Euharmophola Park (1960, p. 23) 'carinata Park (1960, p. 23) Jamaica Dalmonexus Park (1942, p. 272) tobagoensis Park, n. sp Tobago TANYPLEURINA s. str. (Jeannel, 1949, p. 79) Buns Fletcher (1928, p. 227) ensipes (Raffray) (1891, p. 316) Venezuela, Trinidad, Antigua Bythinophysis Raffray (1908, p. 266) humilis (Raffray) (1908, p. 39) Guadeloupe EUPLECTINI s. lat. (Park, 1942, p. 63) RHINOSCEPSINA (Bowman, 1934, p. 144) •Type-species: if West- Indian in range 75 76 FIELDIANA: ZOOLOGY. VOLUME 68 Rhinoscepsis LeConte (1878. p. 382) inaularis Park (1955. p. 105) Jamaica TROGASTRINA (Jeanne!. 1949. p. 75) Eurhexius Sharp (1887. p 41) incertus Park. n. sp Tobago Rhexinia Raffray (1890a. p. 104, 106) versicolor Raffray (1908. p. 37) Guadeloupe PANAPHANTINA (Jeannel. 1950. p 76) Thesiectus Park ( 1960. p. 8) 'probus Park (1960, p. 9) Jamaica Thesium Casey (1884. pp. 94, 117) affabilis Park (1960. p. 6) Jamaica antennalis Park (1955. p. 106) Jamaica pearcei Park. n. sp Tobago cornutum Park (1960. p. 8) Jamaica frontalis Park (1955, p. 108) Jamaica BIBLOPORINA (Park, 1951, p. 64) Biblomimus Raffray (1903, p. 62) 'minutus Raffray (1903, p. 62) „ Guadeloupe impressa Raffray (1908, p. 36) St. Vincent EUPLECTINA s. str. (Jeannel, 1949, p. 45) Thesiastes Casey (1893, p. 444, 457) liliputanus Raffray (1904, p. 543) Grenada hopensis Park, n. sp Jamaica Euplectus Leach (1817, p. 80) exiguus Raffray (1903. p. 542) Grenada insularis Raffray (1908, p. 35) Guadeloupe illepidus Raffray (1903, p. 542) St. Vincent gouyavensis Park, n. sp Grenada episcopalis Park, n. sp Tobago TRIMIINA (Jeannel, 1950. p. 139) Malleoceps Park (1954, p. 1) pearcei Park, n. sp Puerto Rico * darlingtoni Park (1954, p. 1) Dominican Republic Hanfordia Park ( 1960, p. 10) 'absoluta Park (1960, p. 10) Jamaica Dalmomelba Park (1954. p. 6) 'cazieri Park (1954. p. 6) Bahamas Lemelba Park (1953. p. 1) hirsuta Park. n. sp Dominican Republic millsi Park. n. sp Cuba Haasiella Park. n. g. 'medicina Park. n. sp Puerto Rico Actium Casey (1886. p. 201) visendum Park (19.55. p. 109) Jamaica opacum Park (1960. p. 14) Jamaica haasi Park. n. sp Puerto Rico Melba Casey (1897. p. .565) M (Rameloidea) Park (1942. p 14) 'temporalis Raffray (1909. p. 16) Martinique PARK ET AL.: PSELAPHID BEETLES 77 M. (Cismelba) Park (1960, p. 16) 'trifoveata Park (1960, p. 16) Jamaica M. (Frontelba) Park (1942, p. 120) clypeata (Reitter) (1883, p. 40) St. Thomas 'frontalis (Raffray) (1908, p. 35) Guadeloupe M. (Melba) s. str. (Casey) (1897, p. 565) crassipes Raffray (1908, p. 34) Guadeloupe grenadensis Raffray (1903, p. 535) Grenada zonula Park (1954, p. 13) Bahamas specularis (Reitter) (1883, p. 38) St. Thomas, Water Island gibbula (Reitter) (1883, p. 39) St. Thomas M. (Perimelba) Park (1943, p. 183) jamaicensis Park (1955, p. 114) Jamaica hoffi Park (1960, p. 15) Jamaica Latomelba Park (1955, p. Ill) 'quadrisicca Park (1955, p. 112) Jamaica sympatrica Park (1960, p. 12) Jamaica Quadrelba Park (1942, p. 120) *parmata (Reitter) (1883, p. 40) Puerto Rico, St. Thomas inconspicua (Reitter) (1883, p. 41) St. Thomas ventricosa (Reitter) (1883, p. 39) Puerto Rico, St. Thomas Ramelbida Park (1942, p. 112) *quadrifoveata (Raffray) (1903, p. 537) St. Thomas Trimiovillus Park (1954, p. 11) *bahamicus Park (1954, p. 12) Bahamas bellingeri Park (1960, p. 17) Jamaica Sandersonella Park, n. g. * transversa Park, n. sp Haiti Trimiosella Raffray (1898, p. 236) *anguina (Reitter) (1883, p. 42) St. Thomas Hispanisella Park, n. g. *haitiana Park, n. sp Haiti hirsuta Park, n. sp Haiti AUomelba Park (1954, p. 16) *antennata Park (1954, p. 17) Bahamas Zolium Casey (1897, p. 560) *eggersi (Reitter) (1883, p. 38) St. Thomas Trimiopsis eggersi, Reitter 1883, p. 38 Zolium eggersi, Casey 1897, p. 562 Melba eggersi, Raffray 1903, p. 562 Allomelba eggersi. Park 1955, p. 116 BRACHYGLUTINI Raffray (1904, p. 108) DECARTHRONINA (Park, 1951, p. 61) Decarthron Brendel (1865, p. 30) spinosum Raffray (1904, p. 192) Grenada insulare Raffray (1904, p. 189) Grenada arcuatum Park, n. sp Cuba vauriei Park (1954, p. 20) Bahamas pectinate Park, n. sp Cuba 78 FIELDIANA: ZOOLOGY, VOLUME 68 EUPSENIINA (Park. 1951. p. 61) Eupsenius LeConte ( 1850. p. 90) gracilis Raffray (1904. p. 197) Grenada politus Reitter (1883. p. 36) St. Thomas pretiosus Park (1960. p. 18) Jamaica *glaber LeConte (1850. p. 90) U.S.A.. Bahamas dominicanus Schaufuss (1877. p. 108) Dominican Repubhc BRACHYGLUTINA s. str. (Raffray. 1904. p. 108) Reichenbachia Leach (1826, p. 451) grenadensis Raffray (1904. p. 168) Grenada vincentania Raffray (1904. p. 169) St. Vincent eucera (AuW) (1844. p. 120) Puerto Rico darlingtoni Park. n. sp Cuba guadelupensis Raffray (1908, p. 38) Guadeloupe PSELAPTINA (Park, new subtribe) By thinogaster Schaufuss ( 1887. p. 1 1 1 ) nymphoides Park (1960. p. 22) Jamaica parsonsi Park. n. sp Cuba bisphaeroides Schaufuss (1887. p. 297) Cuba 'simplex Schaufuss (1878, p. 112) Hispaniola diabolicus Park (1960, p. 22) Jamaica Berdura Reitter (1881, p. 187; 1883, p. 35) leavitti Park, n. sp Cuba 'excisula Reitter (1883, p. 36) St. Thomas Scalenarthrus LeConte (1880, p. 135) clavatus Raffray (1904, p. 127) Grenada jamaicensis Park (1960, p. 19) Jamaica guadelupensis Raffray (1912, p. 289) Guadeloupe tomentosus Park (1960. p. 20) Jamaica pectinicomis Raffray (1904. p. 127) St. Vincent Pselaptus LeConte (1880, p. 184) sternalis Raffray (1904, p. 208) Grenada longiclava (Schaufuss) (1887, p. 126) Cuba CTENISTINI (Raffray. 1890a, pp. 140, 141) Ctenisodes (1896, p. 247) weberi Park. n. sp Cuba TYRINI (Raffray. 1890a, p. 162) Ephimia Reitter (1883, p. 34) 'simoni Reitter (1882-1883, p. 34) St. Thomas cubensis Park, n. sp Cuba subnitida Raffray (1904. p. 312) Grenada Neotyrus Raffray ( 1895, p. 396) tobagoensis Park, n. sp Tobago HAMOTINI (Park. 1951. p. 67) Apharus Reitter (1882. p. 129) hexagonus Park. n. sp Tobago Hamotus Aub^ (1884. p. 92) H. {Hamotus) s. str. Aub^ (1884. p. 92) Boror Raffray (1904. p. 403) Venezuela. Trinidad. Tobago PARK ET AL.: PSELAPHID BEETLES 79 H. (Hamotoides) Schaufuss (1887, p. 298) hirtus Raffray (1904, p. 408) Grenada, St. Vincent, Guadeloupe CLAVIGERINAE (Redtenbacher, 1849, pp. 58, 647) FUSTIGERINI (Jeannel, 1949, p. 31) Pseudofiistiger Reitter (1884, p. 167) *stricticornis (Reitter) (1883, p. 33) St. Thomas barroi Bierig (1945, p. 15) Cuba Fustiger Brendel (1866, p. 189) haytiana Mann (1915. p. 163) Haiti smithi Raffray (1904, p. 455) St. Vincent schwarzi Mann (1918, p. 105) Cuba jamaicensis Park, n. sp Jamaica TYPE DEPOSITIONS OF NEW SPECIES Park - Orlando Park collection at Field Museum of Natural History INHS - Illinois Natural History Survey MCZ = Museum of Comparative Zoology Bythinoplectus laminatus Park, male TYPE: Park Balega longiceps Park. TYPE male: MCZ Stratus dominicanus Park, TYPE male & 2 PARATYPES: MCZ: 2 PARATYPES: Park Stratus hirsutus Park, TYPE male: MCZ Insulomodes tobagoensis Park, TYPE male & 5 PARATYPES: Park Insulomodes excavatus Park, TYPE male & 9 PARATYPES: Park Dalmonexus tobagoensis Park, TYPE male & 1 PARATYPE female: Park Eurhexius incertus Park, TYPE male: Park Thesium pearcei Park, TYPE male: Park Thesiastes hopensis Park, TYPE female & 2 PARATYPES: INHS; 1 PARATYPE female: Park Euplectus gouyavensis Park, TYPE male: Park Euplectus episcopalis Park, TYPE male: Park Malleoceps pearcei Park, TYPE male & 2 PARATYPES: Park HispaniseUa haitiana Park, TYPE male & 12 PARATYPES: INHS; 2 PARA- TYPES: Park HispaniseUa hirsuta Park, TYPE male: INHS Haasiella medicina Park, TYPE male: Park Sandersonella transversa Park, TYPE male: INHS Lemelba miUsi Park, TYPE male: INHS Lemelba hirsuta Park, TYPE female: MCZ Actium haasi Park, TYPE male: Park Berdura leavitii Park, TYPE female: MCZ; PARATYPE female: Park Bythinogaster parsonsi Park, TYPE male: MCZ Reichenbachia darlingtoni Park, TYPE male: MCZ; PARATYPE male: Park Decarthron arcuatum Park, TYPE & 4 PARATYPES: MCZ; 4 PARATYPES: Park Decarthron pectinale Park. TYPE & 5 PARATYPES: MCZ; 5 PARATYPES: Park Ctenisodes weberi Park. TYPE male: MCZ Ephimia cubensis Park. TYPE: MCZ Neotyrus tobagoensis Park, TYPE male: Park Apharus hexagonus Park, TYPE male: Park Fuatiger Jamatcensis Park, TYPE: Park 80 SUMMARY The pselaphid beetle fauna of the West Indies is poorly known. This report attempts to summarize our knowledge of the described species together with the descriptions of 30 new species, four new genera, and one new subtribe. Of the 113 species known to date, 25 species, representing 17 genera or subgenera, are recorded from Jamaica which has the largest known pselaphid fauna. This may be a reflection of more intensive Berlese sampling on Jamaica than on any of the other Greater Antillean Islands. All three known subfamilies, Faroninae, Pselaphinae, and Cla\dgerinae, including 19 tribes and subtribes are represented in the West Indies. Endemicity in the West Indies is very high, type species of 21 or 40 per cent of the 53 West Indian genera and subgenera having been designated from this area. With much collecting remaining to be done, especially on Cuba and Hispaniola, it is premature to speculate on the faunal origins of the genera and tribes in the West Indies. However, as the analysis stands at present, more genera are shared in common with Central and South America than with Mexico and the United States. Shared with Central and South America, each 20 genera; Mexico, 17 genera; United States, 16 genera. 81 REFERENCES AUBE. C. 1844. Revision de la famille des Ps^laphiens. Ann. Soc. Entomol. France, (2), Part II. pp. 73-160. BlERIG, A. 1945. Nuevos Clavigeridae de Costa Rica y Cuba. Univ. de Costa Rica, San Jose, (Eklidon especial), pp. 4-15, illus. Blackwelder, R. E. 1936. Morphology of the Coleopterous Family Staphylinidae. Smithson. Misc. ColL, 94(13), pp. 1-102, illus. 1944. Checklist of the Coleopterous Insects of Mexico, Central America, The West Indies, and South America, Part I. Bull. 185, Smithson. Inst., pp. i-xii, 1-188. Bowman, J. R. 1934. The Pselaphidae of North America. Pittsburgh, Pa. (privately publ.), pp. i-vi, 1-149. Brendel, E. 1865. On some new species of Pselaphidae. Proc. Entomol. Soc. Philadelphia, pp. 28-32. 1866. Descriptions of some new species of Pselaphidae. Proc. Entomol. Soc. Philadelphia, July 1866, pp. 189-194. 1889. Descriptions of Scydmaenidae and Pselaphidae. Entomol. Amer., 6, pp. 193- 197. Casey, T. L. 1884. Contributions to the Descriptive and Systematic Coleopterology of North America. Part I, pp. 1-60. Part II, pp. 61-198. Philadelphia (privately publ.). 1886. Descriptive notices of North American Coleoptera, Part I. Bull. Calif. Acad. Sci.. 2 (6). pp. 157-264. 1887. On some new North American Pselaphidae. Bull. 8, Calif. Acad. Sci., Vol. II. pp. 455-482. pi. 16. 1893. Coleopterological Notices, V. Ann. New York Acad. Sci., 7, pp. 281-606. 1897. Coleopterological Notices, VII. Ann. New York Acad. Sci., 9, pp. 285-684. PARK ET AL.: PSELAPHID BEETLES 83 Fletcher, F. C. 1928. Pselaphidae collected by Dr. Alfons Dampf in Central America. Ann. Entomol. Soc. Amer., 21, (2), pp. 203-231, figs. 33. Grigarick, a. a. and R. O. Schuster 1971. A revision of Actium Casey and Actiastes Casey (Coleoptera: Pselaphidae). Univ. Calif. Press, Berkeley, pp. 1-56. Grouvelle, a. and A. Raffr.ay 1908. Supplement a la Liste des Coleopteres de la Guadeloupe. Ann. Soc. Entomol. France, 77. pp. 33-40. 1912. Supplement a la Liste des Coltopteres de la Guadeloupe. Ann. Soc. Entomol. France, 81, pp. 289-290. Jeannel, R. 1949. Les Ps^laphides de I'Afrique Orientale. Mem. Mus. Nat. d'Hist. Nat., N.S., 29 (1), pp. 1-226, figs. 103. 1950. Faune de France, N. 53. Coleopteres Pselaphides, Lechevalier, Paris, pp. 1- 421, figs. 169. 1955. Les Pselaphides de I'Afrique Australe. Mem. Mus. D'Hist. Nat., 9, pp. 1-196, figs. 226. 1962. Les Pselaphides de la Paleantarctide Occidentale. Biol, de L'Amerique Australe, Centre National Investigaciones Cientificas y Technicas, Vol. 1, pp. 295-479. Kugelann, J. G. 1794. Verzeichniss derin einigen Gegenden Preussens bei jetz entdekten. Kafer- Arten. Schneider's Mag., 1 (5), pp. 513-582. Leach, W. E. 1817. On the stirpes and genera comix)sing the Family Pselaphidae; with the names of the British species. The Zool. Misc.; being descriptions of new or interesting animals, 3, pp. 80-87. 1826. On the stirpes and genera composing the Family Pseladphidae; with descriptions of some new species. Zool. Jour., 2, pp. 445-453. LeConte, J. L. 1850. On the Pselaphidae of the United States. Boston Jour. Nat. Hist., 6(1), pp. 64-110. [Sept. 1849 date on original paper. J.W.-M.W.S.] 1863, 1866. Descriptions of new species of North American Coleoptera, Smithson. Misc. Coll., Part I, pp. 1-86 (1863), pp. 87-168 (1866). 1878. Additional descriptions of new species. Proc. Amer. Phil. Soc., 17, pp. 373-434. 1880. Short studies of North American Coleoptera. Trans. Amer. Entomol. Soc, 8, pp. 613-218. 84 FIELDIANA: ZOOLOGY. VOLUME 68 Mann. W. M. 1915. Some Myrmecophilous insects from Hayti. Psyche, 22 (5). pp. 161-166. Illus. 1918. Myrmecophilous insects from Cuba. Psyche, 25 (5), pp. 104-106. Park. O. 1942. A study in Neotropical Pselaphidae. (Book). Northwestern Univ. Stud. Biol. Sci. Med., no. 1, pp. x, 403, pLs. 21. 1943. A preliminary study of the Pselaphidae (Coleoptera) of Mexico. Bull. Chicago Acad. Sci., 7 (3). pp. 171-226, pis. 3. 1951. Cavernicolous Pselaphid beetles of Alabama and Tennessee, with observations on the taxonomy of the family. Geol. Surv. Alabama, Mus. Paper 31, pp. 1-107, figs. 18. 1952a. A revisional study of Neotropical Pselaphid Beetles. Part I. Tribes Faronini, Pyxidicerini and Jubinini. Chicago Acad. Sci., Spec. Pub., no. 9, pp. 1-49, pis. 3. 1952b. A revisional study of Neotropical Pselaphid beetles. Part II. Tribe Euplectini sensu latiore. Chicago Acad. Sci., Spec. Pub., no. 9, pp. 50-150, pis. 7. 1953. A new genus of Pselaphid beetles from the Everglades. Nat. Hist. Misc., no. 121, p. 4. 1954a. The Pselaphidae of South Bimini Island, Bahamas, British West Indies (Coleoptera). Amer. Mus. Nov., no. 1674, pp. 1-25, figs. 22. 1954b. A new genus of Pselaphid beetles from the Antilles. Nat. Hist. Misc., no. 138, pp. 1-4, figs. 3. 1955. Contributions to the Pselaphid beetle fauna of Jamaica. Bull. Chicago Acad. Sci., 10 (7), pp. 101-120, figs. 5. 1960. Pselaphid beetles of Jamaica. Trans. Amer. Microscop. Soc, 79 (1), pp. 5-25, figs. 18. 1964. Observations upon the behavior of Myrmecophilous Pselaphid Beetles. Pedobiologia, Bd. 4 (3). pp. 129-137. 1965. Revision of the genus Batriasymmodes (Col. Psel.), Trans. Amer. Microscop. Soc, 84 (2), pp. 184-201. Raftray, a. 1890a. Etude sur les Ps^laphides: Genera et descriptions d'esp^ces nouvelles. Rev. d'Entomol., 9, pp. 1-28, 81-172, 193-219, 264-265. 1890b. Stance du 26 Novembre 1890. (Description d'une nouvelle espece de pe^laphide). Bull. Soc. EntomoL France, ser. 6, Vol. 10, pp. cciv-ccv, 204-205. 1891. Voyage de M. E. Simon au Venezuela (D^cembre 1887-Avril 1888). 10 M^moire. Ps^laphides. Ann. Soc. Entomol. France, ser. 6, Vol. 10, pp. 297-330, illus. 1893. Essai Monographique sur la tribu des Faronini (Ps6laphiens). Rev. d'Entomol., 12, pp. 157-196, figs. 65. 1895. Revision du genre Tryopsis Saulcy (Aplodea Reitter) et description de deux genres nouveaux du m£me groupe. Ann. Soc. Entomol. France, 64, pp. 391-400. PARK ET AL.: PSELAPHID BEETLES 85 1896. Nouvelles Etudes sur les Pselaphides et les Clavigerides. Ann. Soc. Entomol. France, 65, pp. 227-284, pis. 2. 1897. Revision des Batrisus et genres voisins de TAm^rique centrale et m^ridionale. Ann. Soc. Entomol. France, 66, pp. 431-517. lUus. 1898. Notes sur les Pselaphides. Revision g^n^rique de la tribu des Euplectini. 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Beitrag zur Kenntniss der Clavigeriden, Pselaphiden und Scydmaeniden von Westindien. Deut. Entomol. Z., 27, pp. 33-46. 1884. Ueber die bekannten Clavigeriden-Gattungen. Deut. Entomol. Z., 28, pp. 167- 168. 1890. Coleopterologische Notizen XXXVIII. Wien. Entomol. Zeit., 9, pp. 210-213. 1897. Zehnter Beitrag zur Coleopteren Fauna von Europa und den angrenzenden Landern. Wein. Entomol. Zeit., 16, pp. 203-206. Roberts, W. A. 1940. The Caribbean. The story of our destiny. Bobbs-Merrill, Indianapolis, Indiana. IX. 361 pp. SCHAUFUSS, L. W. 1872a. Beschreibung einiger Pselaphiden. Nunquam Otiosus, Dresden, 2, pp. 259- 274. 1872b. Ueber Pselaphidengattungen. Nunquam Otiosus, Dresden, 2, pp. 450-460. 86 FIELDIANA: ZOOLOGY. VOLUME 68 1886. Beschreibung neuer Pselaphiden aus der Sammlung des Museum Ludwig Salvator. Ein Beitrag zur Fauna Brasiliens, der Kgl. Niederlaindischen Besitzungen in Indien und NeuhoUands. Tijdschr. Entomol., 29, pp. 241-296. Illus. 1887a. Beschreibung neuer Pselaphiden aus der Sammlung des Museum Ludwig Salvator. Ein Beitrag zur Fauna Brasiliens, der Kgl. Niederlandischen Beatzungen in Indien und NeuhoUands. Tijdschr. Entomol., 30, pp. 91-165. lUus. 1887b. Uber Pselaphiden und Scydmaeniden des Kbnigl. zoologischen Mus. zu Berlin und verwandte Arten. Berlin. Entomol. Z.. Bd. 31. Heft II, pp. 287-320. Sharp, D. 1887. Pselaphidae. Biologia Centrali-Americana. Coleoptera, 2, Part I, pp. 1-46, pi. 1. TANNER, V. M. 1927. A preliminary study of the genitalia of female Coleoptera. Trans. Amer. Entomol. See, 53, pp. 5-50. Wagner, J. A. 1975. Review of the Genera Euplectus, Pycnoplectus, Leptoplectus, and Acolonia (Coleoptera: Pselaphidae) Including Nearctic Species North of Mexico. Entomol. Amer.. New York, 49 (2), pp. 125-207, figs. 1-74. Index to the Subfamilies. Tribes, Subtribes and Genera FOR THE Systematic Section. Keys, and Figures Major text references are in boldface, and figure references are in italics. Achillia, 56 Actium, key9-29 and 42. 37, 42 AUomelba, key-29. 38. 45 Apharus, 65, key -69. 69 Aplodea, 68 Balega, 7, key-8. 8 Berdura, key-49. 49, 50 Biblomimus, key-24, 24 Bibloporina. key-19, 24 Brachyglutina, key-48, 55 Brachyglutini, 6, keys-7 and 47, 47 Bryaxis, 6 Buns, key-18. 19 Bythinogaster, 6. keys-49 and 53, 50, 52 Bythinophysis, key-18, 19 Bythinoplectus, 5 Caccoplectus , 4 (Cismelba), key-45.47 Clavigerinae, key -4. 72 Coptotermes, 69 Ctenisodes, 64,65,70 Ctenistini. key-7, 64. 66 Ephimia, key-66, 66, 70 Euharmophola, key-13, 17 Euplectina, keys-19 and 24, 24 Euplectini, keys-7 and 19, 19 Euplectus, keys-24 and 26, 26 Eupseniina, 47, 62 Eupsenius, 62, key-63 Eurhexius, key-20, 20 Faroninae, 4 Faronini, 4 (Fronte/6a),key-46,46 Fustiger, 70, keys-72 and 73, 73 Fustigerini, key-72, 72 Haasiella, key-28, 35, 37 Hamotini, keys-7 and 69, 69 (Hamotoides), key -69, 72 Hamotus, 65, key -69, 71 Hanfordia, 19, key-28, 40 Hispanisella, keys-29 and 32, 32, 35, 37 Hybocephalini, 66 Insulomodes, key-13, 13, 13 Dalmodes, 19 Dalmodiina, keys- 12 and 13 Dalmomelba, key-28, 35 Dalmonexus, key-13, 17 Dalmosella, 36 Decarthron, 58, key-59, 61 Decarthronina, key-47, 58 Jubinini, keys-7 and 8, 7 Jubus, 7, 8, key-12, 12 Juxtahamotopsis, 66 Latomelba, 19, keys-29 and 44, 32, 38, 44 Lemelba, key8-28 and 40. 36. 37, 40 87 88 FIELDIANA: ZOOLOGY. VOLUME 68 Malleoceps, key8-28 and 29. 29, 31 Melba. keys-29 and 45. 45 (Melba),A% Rhexinia. key-20. 21 Rhinoscepsina, key-19, 20 Rhinoscepsis, 20 Neotyrus, 65. key-€6. 67, 70 Odontalgini, 66 Panaphantina. keys- 19 and 22, 21 iPerimelba), keys-46 and 47. 47 Pselaphinae, key8-4 and 6, 4 Pselaptus, 48. keys-49 and 51. 50, 51 Pselaptina, keys-48 and 49, 48 Pseudofustiger, key-72. 72 Pycnoplectus, 27 Pyxidicerini, key -4, 5 Quadrelba, keys-29 and 44, 38, 44 Ramelbida, key-29, 31, 40 {Rameloidea), key-45, 46 Reichenbachia, 54, 56, key-57 Sandersonetla, key-29, 37, 38 Scale narthrus, keys-49 and 51, 51 Stratus, 7, 10, key-10 Tanypleurina, keys-12 and 18, 18 Tanypleurini, keys-7 and 12, 12 Thesiastes, keys-24 and 25, 24 Thesiectus, Viey-22,22 T/icsjum,key-22,22, 23 Triangusella, 32 Trimiina, keys-20 and 28, 28 Tnmiosella, key-29, 32, 35 Trimiovillus, keys-29 and 45, 45 Trogastrina, keys- 19 and 20, 20 Tyrini, keys-7 and 66, 66 Zo/ium, key-29, 45 Index to Species for the Systematic Section, Keys, and Figures Major text references are in boldface, and figure references are in italics. absoluta Park (Hanfordia), 40 acuta Park ( Triangusella ). 32 acutangulus Raffray (Bythinoplectus), key-5 affabilis Park (TAesium), key-22 anguina ( Reitter) (Trimiosella ), 35 antennaiis Park {Thesium), key-22 antennata Park {AUomelba), 45 arcuatum Park {Decarthron), key-60, 60,61 armipes Raffray (Apharus), 71 bahamicus Park {Bythinogaster), key-5 bahamicus Park {Trimiovillus), key-45 barroi Bierig (Pseudofustiger), key-72 belfragei LeConte [Pselaptus ), 48, 51 bellingeri Park (Caccoplectus ), 5 bellingeri Park (Trimiovillus), key-46 bisinuata Schaufuss [Bryaxis), 56 bisphaeroides Schaufuss (Bythinogaster), key-53, 54 carinata Park (Euharmophola), 17 cazieri Park (Dalmomelba) , 35 clavatus Raffray (Jubus), 12 clavatus Raffray (Scalenarthrus), key-5 1 clypeata Reitter (Melba), key-46 coptocolus Park (Neotyrus), 69 PARK ET AL.: PSELAPHID BEETLES 89 comutum Park ( Thesium ), key -29 crassipes Raffray {Melba), key-46 cubensis Park (Ephimia), key-67, 67, 70 guadelupensis Raffray (Reichenbachia) , key-57, 59 guadelupensis Raffray {Scalenarthrus ), key-51 darUngtonia Park (Reichenbachia), key-57, 57, 59 davisi Park (Lemelba), key -40, 40 diabolicus Park (Bythinogaster), key-53, 54 dominicanus Park {Stratus), key-10, 10 dominicanus Schaufuss {Eupseninus), eggersi ( Reitter ) (ZoUum), 45 elegans ( Reitter) (Balega), key -8 ensipes (Raffray) {Buris), 19 episcopalis Park (Euplectus) , key-26, 27,28 eucera Aube {Reichenbachia), 54, key-57, 59 excavatus Park {Insulomodes), key-14, 16 excisa Schaufuss (i4cAj7i'a), 56 excisula Reitter (flendura), key-49, 49, 50 exiguus Raffray {Euplectus), key-26 fleutiauxi (Raffray) {Melba), 47 foveatus Reitter {Bythinoplectus), key-5 frontalis Raffray {Melba), key-46 frontalis Park {Thesium), 22 gibbicoUis (Schaufuss) (Neotyrus), 68, 69 gibbula Reitter {Melba), key-47 glaber LeConte {Eupsenius ), key-63, 62 gouyavensis Park {Euplectus), key-26, 26 gracilis Raffray {Eupsenius), key-63 grenadensis Raffray {Melba), key-46 grenadensis Raffray {Reichenbachia), key-51, 59 haasi Park {Actium),37, key-43, 43 haitiana Park {Hispanisella) , key -32, 33,55 hambletoni Park {Eurhexius), 21 harem Park (Neotyrus), 69 haytiana Mann {Fustiger), key-73 hexagonus Park (i4pAarus), 65, 69, 71 hirsuta Park {Hispanisella), key-32, 34,37 hirsuta Park {Lemelba), 37, key-40, 41 hirsutus Park (Stratus), key-10, 11, i3 hirtus Raffray (Hamotus), 72 hoffi Park (Me/6a), key-47 hopensis Park {Thesiastes), key-25, 25 humilis (Raffray) {Bythinophysis), 19 illepidus Raffray {Euplectus), key-29 impressa Raffray {Biblomimus), key-24 incertus Park {Eurhexius), 20, 21 inconspicua (Reitter) {Quadrelba), key-44 insulare Raffray (Decarthron) , key -60 insularis Park {Rhinoscepsis), 20 insularis Raffray {Euplectus), key-26 insufans Raffray {Jubus), 12 intermedia Park {Balega), key-8, 9 jamaicaensis Park {Fustiger), 70, key-73, 73 jamaicaensis Park {Melba), key-47 jamaicaensis Park {Scalenarthrus), key-51 laminatus Park {Bythinoplectus), key-5, 5 laticeps Raffray (Ctenisodes), 64, 65 teoL-itri Park (fl^rdura), key-49, 49, 50 liliputanus Raffray {Thesiastes), key-25 Publication 1240