ROSSER W. GARRISON., Los Angeles County Agricultural Commissioner's Office, U.S.A. A REVISION OF THE NEW WORLD GENUS ERPETOGOMPHUS HAGEN IN SELYS (ODONATA: GOMPHIDAE) Contents Garrison, R. W., 1994. A revision of the New World genus Erpetogomphus Hagen in Selys (Odonata: Gomphidae). -Tijdschrift voor Entomologie 137: 173-269, figs. 1-230, tabs. 1-4 [issn 0040-7496]. Published 15 December 1994. This revision of adult Erpetogomphus includes a phylogenetic assessment of all 21 species using outgroup comparison and parsimony algorithm, descriptive biogeography, keys to both sexes, synonymies, descriptions, type designations, and illustrations, including distribution maps of all species. Six new species are described: E. agkistrodon, E. leptophis, E. elaphe, E. liopeltis, E. bothrops, E. heterodon. Erpetogomphus coluber is considered a junior synonym of E. compositus, E. natrix is considered a subspecies of E. lampropeltis, and a neotype is designated for E. cophias. Phylogenetic assessment of 41 mostly somatic characters shows Erpetogomphus to be partitioned into three monophyletic groups: 1) six dark green species (E. constrictor, E. sabaleticus, E. tristani, E. agkistro-don, E. schausi, E. ophibolus) with mostly allopatric or parapatric distributions along the eastern coast of Mexico south into northern Colombia and Venezuela, 2) two species {E. leptophis, E. eu-tainia) with distributions from southern Texas south through Mexico and up the west coast to Michoacan states, and 3) 13 remaining species (E. elaphe, E. elaps, E. liopeltis, E. bothrops, E. vi-perinus, E. désignants, E. sipedon, E. lampropeltis, E. crotalinus, E. heterodon, E. compositus, E. boa, E. cophias) with distributions in the centrai United States south through Mexico to Costa Rica. Derived characters States were gleaned mostly from primary and secondary genitalic characters (head structure, penis, hamules, caudal appendages, vulvar lamina); but satisfactory resolution of terminal clades, especially of the third group, is difficult due to apparent character reversals. Dr. R. W. Garrison, Los Angeles County Agricultural Commissioner's Office, 3400 La Madera Ave., El Monte, CA 91732, USA Key words. -Odonata; Gomphidae; Erpetogomphus-, systematica; keys; cladistics; phylogeny; biogeography. Erpetogomphus agkistrodon sp. n 199 Erpetogomphus schausi Calvert 200 Erpetogomphus eutainia Calvert 201 Erpetogomphus leptophis sp. n 204 Erpetogomphus elaphe sp. n 205 Erpetogomphus elaps Selys 207 Erpetogomphus liopeltis sp. n 210 Erpetogomphus bothropssp. n 212 Erpetogomphus viperinus Selys 215 Erpetogomphus désignants Hagen /'«Selys .... 217 Erpetogomphus sipedon Calvert 22 1 Erpetogomphus lampropeltis Kennedy 223 E. I. lampropeltis Kennedy 225 E .1. natrix Williamson and Williamson stat. n 226 Erpetogomphus crotalinus (Hagen in Selys) .. 227 Erpetogomphus heterodon sp. n 230 Erpetogomphus compositus Hagen in Selys ... 232 Erpetogomphus boa Selys 236 Erpetogomphus cophias Selys 23 8 Acknowledgements 240 References 241 Illustrations 247 Introductory part Introduction 174 History 174 The status of the name Erpetogomphus menetriesii (Selys) 174 Biology 176 Phylogenetics and biogeography Phylogenetic analysis 177 Cladospecies 180 Distribution patterns 181 Descriptive biogeography 182 Characters 182 Erpetogomphus species groups 185 Systematic part Introduction 185 Genus Erpetogomphus Hagen /«Selys, 1858 186 Keys 188 Erpetogomphus constrictor Ris 193 Erpetogomphus sabaleticus Williamson 195 Erpetogomphus tristani Calvert 196 Erpetogomphus ophibolus Calvert 197 173
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