A revision of the spider genus Cocalus (Araneae: Salticidae) F. R. Wanless Department of Zoology, British Museum (Natural History), Cromwell Road, London SW7 5BD Introduction Cocalus Koch 1846 is a small genus of oriental and Australian Salticidae comprised of four known species, one of which is described here as new. Although the type species, Cocalus concolor Koch, is only known from a single adult male which has lost both palps, it is thought the low elevation on the carapace, shared by all species is sufficiently characteristic to unite the species here included in Cocalus. In the only known female (C. murinus Simon) the epigyne is dark with a median orifice and is not especially characteristic when compared with epigynes of related genera. However, the male palps (in the two species, C. gibbosus sp.n., and C. limbatus Thorell, for which these are known) are distinctive, the sinuous finger-like protuberance of the tibiae being, as far as I am aware, unique within the Salticidae. Opisthoncus Koch from Australia is the only other genus known to the author to have a similar elevation on the carapace. It does not, however, appear to be closely related to Cocalus as the eye patterns are different, the elevation is situated between the posterior lateral eyes and the palps are less complex. The biology of Cocalus is unknown. The standard abbreviations and measurements are those used by Wanless (1978a), but for the leg spination the system adopted is that used by Platnick and Shadab (1975). Genus COCALUS Koch Cocalus Koch, 1846: 180, 230. Type species Cocalus concolor Koch, by subsequent designation (Simon, 1901:408). Koch, 1850:48. Marschall, 1873:393. Scudder, 1882:72. Peckham & Peckham, 1885:266, 288. Simon, 1901:405, 407, 408. Sherborn, 1922:1375; 1932:345. Petrunkevitch, 1928 : 181. Neave, 1939, 1 : 778. Roewer, 1954 : 934. Bonnet, 1956 : 1 173. Cocala: Simon, 1 864 : 327, [lapsus calami]. Waterhouse, 1912:61. Neave, 19391: 778. DEFINITION. Medium to large spiders ranging from about 1'5 to 8'6 mm in length. Sexual dimorphism not marked. Carapace: characteristic of genus, profile generally as in Figs 2 A, B; 3A, B; moderately high, longer than broad, widest between coxae II-III, lateral margins usually with membraneous margin; fovea moderately long, situated just behind posterior lateral eyes; positioned centrally within posterior ocular quadrangle a small bump (arrowed in Fig. 2 A, B, E). Eyes: set on moderately well developed tubercles, with black surrounds that are normally covered in hairs; arranged in three transverse rows comprised of anterior medians and anterior laterals, posterior medians, and posterior laterals; anteriors subcontiguous with apices strongly procurved in frontal view and recurved in dorsal view; anterior medians largest, anterior laterals about half diameter of anterior medians; posterior medians relatively large, positioned more or less midway between anterior laterals and posterior laterals and outside optical axis of anterior laterals; posterior laterals about as large as anterior laterals, set close to lateral margins of carapace when viewed from above; quadrangle formed by posterior median and posterior lateral eyes broader than long and widest posteriorly; entire quadrangle (measured from between bases of anterior medians to posterior margins of posterior lateral eyes) occupying between 45 and 47 per cent of carapace length. Clypeus: between 32 and 64 per cent of diameter of anterior median eyes, usually clothed in hairs. Chelicerae: robust, more or less parallel, slightly inclined anteriorly; Bull. Br. Mus. nat. Hist. (Zool.) 41 (5): 253-26 1 Issued 1 7 December 198 1 253
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