ON THE DIPTERA OF NEPAL 
(STRATIOMYIDAE, THEREVIDAE 
AND DOLICHOPODIDAE) 
BY 
DAVID HOLLIS 
lately of the Department of Entomology 
British Museum (Natural History) 
Pp. 81-116 ; 48 Text-figures 
BULLETIN OF 
THE BRITISH MUSEUM (NATURAL HISTORY) 
ENTOMOLOGY Vol. 15 No. 4 
LONDON: 1964 
THE BULLETIN OF THE BRITISH MUSEUM 
(NATURAL HISTORY), instituted in 1949, is 
issued in five series corresponding to the Departments 
of the Museum, and an Historical series. 
Parts will appear at irregular intervals as they become 
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within one calendar year. 
This paper is Vol. 15, No. 4 of the Entomological 
series. The abbreviated titles of periodicals cited follow 
those of the World List of Scientific Periodicals. 
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Issued 21 July, 1964 Price Thirteen Shillings 
ON THE DIPTERA OF NEPAL 
(STRATIOMYIDAE, THEREVIDAE 
AND DOLICHOPODIDAE) 
By DAVID HOLLIS 
CONTENTS 
Page 
INTRODUCTION ........... 83 
STRATIOMYIDAE ........... 84 
THEREVIDAE ........... 84 
DOLICHOPODIDAE .......... 86 
Subfamily Chrysosomatinae ........ 88 
Subfamily Dolichopodinae ........ 89 
Subfamily Hydrophorinae ........ 93 
Subfamily Rhaphiinae ......... 93 
Subfamily Diaphorinae ......... 96 
Subfamily Campsicneminae . . . . . . . 101 
REFERENCES . . . . . . . . . . .114 
INDEX . . . . . . . . . . . .116 
SYNOPSIS 
Published and new records of Stratiomyidae, Therevidae and Dolichopodidae from Nepal 
are given. 2 new species of Therevidae ; i new genus, 22 new species (including i from Tibet) 
and i new subspecies of Dolichopodidae are described. 
INTRODUCTION 
THE dipterous fauna of Nepal is poorly known and this paper attempts to bring up 
to date both published and new records of Stratiomyidae, Therevidae and 
Dolichopodidae from this country. 
The work was stimulated by the collections of two British Museum expeditions 
to Nepal. On the first expedition, in 1954, the Diptera were collected by Mr. J. 
Quinlan. The second expedition, during the winter of 1961-62, provided the bulk 
of material studied and this collection was made by Mr. R. L. Coe. 
One new species, Hercostomus kaulbacki sp. n. from Tibet is described here because 
of the proximity of the locality and the unusual characteristics displayed by the fly. 
A full bibliography of the recorded species is not given and only references to 
taxonomic changes and new locality records are stated. 
Unless otherwise stated all type material is in the British Museum (Natural 
History) . 
The author would like to thank Mr. C. E. Dyte, of the Pest Infestation Labora- 
tories, Slough, for his valuable advice and criticism ; Mr. W. N. Ellis, of the 
Zoologisch Museum, Amsterdam, for the helpful loan of type material ; and Dr. 
Sadao Takagi, of the Entomological Institute, Hokkaido University, Sapporo, for 
his gift of type material to the British Museum. 
84 D. HOLLIS 
STRATIOMYIDAE 
Brunetti (1923), in his revision of the Oriental Stratiomyidae, records many 
species from Darjeeling and nearby districts, but in this and other relevant papers 
there is no record of Stratiomyidae from Nepal. 
In the British Museum collection one species is represented from the area. 
Ptecticus melanurus (Walker) 
Ctenophora melanura Walker, 1848 : 78. 
Ptecticus apicalis Loew, 1855 : 142. 
Sargus luridus Walker, 1856 : 8. 
Sargus leoninus Rondani, 1875 : 454. 
Sargus melanurus (Walker) Osten Sacken, 1886 : 166. 
Ptecticus aurifer Brunetti (nee Walker), 1920 : 78. 
Ptecticus melanurus (Walker) Brunetti, 1923 : 138. 
4 J, NEPAL : Taplejung Distr., Sangu, c. 6,200', mixed vegetation by stream in 
gully, ix-x.i96i ; 8 $, flying into pit latrine ; i <$, xi. 1961-1.1962 (R. L. Coe). 
THEREVIDAE 
As far as can be gathered there are no previous records of Therevidae from Nepal. 
Two apparently undescribed species of this family are present in the British 
Museum collection. 
Thereva hinu sp. n. 
(Text-figs. 1-6) 
o*. Eyes contiguous over upper frons ; lower frons grey dusted with long, straight, 
intermingled grey and black hairs. Antennae grey-black and bearing silver and black hairs and 
black bristles. Face grey, thickly covered with long silver and black hairs. Occiput grey with 
a post-ocular fringe of long black hair-like bristles and a uniform covering of long silver hairs 
ventrally. 
Dorsum of thorax grey with 3 vague brown longitudinal stripes and a uniform covering of 
thin black hairs ; 3 pre-sutural, i sutural, 2 supra-alar and i post-alar bristle present. 
Scutellum concolorous with dorsum with 4 marginal bristles and some long dark hairs. Pleurae 
grey, densely covered with long silver hairs. 
Abdomen shining dark grey-black with lateral and posterior margins of tergites grey dusted ; 
mixed pale and dark hairs dorsally but only pale hairs laterally and ventrally. Genitalia with 
lateral lamellae orange, penis S-shaped. 
Legs with coxae dark grey and long pale haired, posterior pair with i external black bristle. 
Femora grey with long silvery hairs ; posterior pair with a few black bristles ventrally. Tibiae 
orange with darkened tips and black bristles. Tarsi mainly black, basitarsi orange in basal half. 
Wings clear hyaline with a weak brown stigma, 4th posterior cell closed. Halteres dark. 
$ much more sparsely haired than <J. Eyes separated over frons which is one-third width of 
head at its narrowest and widens to almost twice this width just above antennae. Frontal 
callosities represented by a broad irregular band of darker pigment. Post-ocular fringe shorter 
than in (J and more bristle-like. 
Length, 8 mm. 
ON THE DIPTERA OF NEPAL 
FIGS. 1-7. Thereva spp., T. hinu sp. n. 1-5. <$ genitalia. i. sub-apical sternite. 
2. apical sternite. 3. lateral lamella. 4. phallobase and aedeagus. 5. apical 
tergite. 6. $ head, dorsal aspect : T. brunettii sp. n. 7. $ head, dorsal aspect. 
Holotype $, NEPAL : Taplejung Distr., damp evergreen oak forest above Sangu, 
c. 10,400', 2-26. xi. 1961, resting on stones in clearing (R. L. Coe). 
Paratypes, 7 $, i $, same data as Holotype. 
This species does not seem to be related to the known Oriental species of Thereva. 
Of the Palaearctic species, innotata Krober (1912), described from Turkestan, is 
86 D. HOLLIS 
close but may be distinguished by the orange colour of the 2nd antennal segment 
and base of the 3rd, and the brown-black thorax bearing 2 yellow-grey, longitudinal 
stripes. 
Thereva brunettii sp. n. 
(Text-fig. 7) 
$. Frons about one-sixth width of head at narrowest part and widening to over twice its 
original width above antennae ; orange-grey dusted with a trapezoid area of dark pigment 
extending to inner eye margins below ocellar triangle, light and dark haired. Antennae with 
ist segment grey, pale haired and with black bristles sub-apically ; 2nd segment and base of 
3rd reddish, remainder of 3rd segment grey-black. Face whitish-grey, long pale haired. 
Proboscis orange-brown. Occiput convex, grey, with upper post-ocular cilia black and ventrally 
with dense pale hairs ; a few black bristles below post-ocular row. 
Thorax dorsally grey-brown with short pale and dark hairs ; no definite longitudinal stripes 
but 2 vague, brown lines show laterally ; 1-2 pre-sutural, 2 sutural, 2 supra-alar and i post- 
alar bristle. Scutellum concolorous with dorsum, with 4 black marginal bristles and an even 
covering of shorter pale hairs. Pleurae grey, pale haired. 
Abdomen with tergites shining brown-black with yellow dusting on posterior margins, the 
colour widening laterally ; evenly covered with short golden hairs. 
Legs with coxae grey, white haired and with black apical bristles. Femora dark brown-black, 
white haired, posterior pair with an antero- ventral row of 4-5 short, brown-black bristles 
sub-apically. Tibiae orange-yellow, brown apically, brown bristled ; anterior pair with 2 
antero-dorsal, 2 postero-dorsal, 2 antero-ventral and 2 postero-ventral bristles. Anterior and 
middle basitarsi orange, following tarsal segments and all posterior tarsal segments brown-black. 
Wings pale yellow-brown hyaline, stigma yellow ; 4th posterior cell closed. 
cJ unknown. 
Length, 10-13 mm. 
Holotype $, NEPAL : Taplejung Distr., old mixed forest above Sangu, c. 6,200', 
25-28. x.ig6i (R. L. Coe). 
Paratypes 2 $, INDIA : Darjeeling, io.x.i92o and 8.xi.ig20, Pres. E. Brunetti. 
T. bilineata Brunetti (1920), described from N. India, is close to brunettii sp. n. 
but may be distinguished by the much wider frons, completely dark antennae and 
2 clear yellowish white stripes on the dorsum of the thorax. 
DOLIGHOPODIDAE 
To the best of my knowledge only 4 species have previously been recorded from 
Nepal : Tachytrechus crassitarsis de Meijere, 1916, recorded by Becker (1922) ; 
Sympycnus gummigutti Becker, 1922, described from Nepal ; Diaphorm mandarinus 
Wiedemann, 1830, and ThinophUus indigenus Becker, 1903. The two latter species 
were recorded by Ribeiro (1923) in his amendments to the localities in Becker (1922). 
None of these species was represented in the British Museum collections which 
included 27 species and i subspecies in 12 genera. The list of species recorded from 
Nepal now stands as : 
CHRYSOSOMATINAE DOLICHOPODINAE 
Sciopus pediformis Becker, 1922 Dolichopus angustinervis Becker, 1922 
Hercostomus ulleriensis sp. n. 
H. phollae sp. n. 
Tachytrechus crassitarsis de Meijere, 1916 
T. compositus sp. n. 
ON THE DIPTERA 
NEPAL 
HYDROPHORINAE 
Thinophilus indigenus Becker, 1903 
DIAPHORINAE 
Diaphorus mandarinus Wiedemann, 1830 
D. jeanae sp. n. 
D. sanguensis sp. n. 
Chrysotus discretus Becker, 1922 
C. pseudocilipes sp. n. 
C. kholsa sp. n. 
Argyra pseudosuperba sp. n. 
RHAPHIINAE 
Syntormon dukha sp. n. 
S. babu sp. n. 
S. ama sp. n. 
CAMPSICNEMINAE 
Micromorphus albipes (Zetterstedt, 1843) 
Sympycnus gummigutti Becker, 1922 
5. coei sp. n. 
S. arunensis sp. n. 
S. albipes nepalensis subsp. n. 
S. laetus Becker, 1922 
S. khola sp. n. 
S. turbidus Becker, 1922 
S. peniculitarsus sp. n. 
5. gauri sp. n. 
5. takagii sp. n. 
5. pahar sp. n. 
Chrysotimus anomalicems sp. n. 
Nepalomyia dytei gen. and sp. n. 
N. confusa sp. n. 
This species list cannot be taken as complete as most of the specimens studied 
were collected from East Nepal in the winter months and very little is known of 
the summer fauna and that of the more north-westerly region of the country. 
As is to be expected the Dolichopodid fauna of Nepal is mainly Oriental with 
little influence from the Palaearctic region. It is, however, interesting that many 
species, especially in the Campsicneminae, show relationships with the faunas of 
Ceylon, Indonesia, Formosa and the Philippines, and the only subspecies described 
shows a direct link through Ceylon with other islands in the Indian Ocean. The 
Himalayan Region would therefore seem to be a meeting point for several lines of 
distribution in the Oriental Region. 
The predominance of Campsicneminae in the recorded species may not be a true 
reflection of the Dolichopodid fauna of Nepal as much of the collecting was carried 
out in deep gullies with low vegetation. 
KEY TO THE GENERA KNOWN FROM THE HIMALAYAN REGION 
Those genera in parentheses are not recorded from Nepal. 
: 4th long vein forked ; thorax not longer than wide ; vertex sunken on either side of 
a prominent ocellar triangle ; abdomen long and thin, hypopygium pedunculate . 2 
- 4th long vein not forked ; thorax longer than wide ; vertex not deeply sunken 
between upper eye margins .......... 6 
t In the <J no definite joint between arista and 3rd antennal segment thus forming a 
compound structure which is flattened dorso-ventrally, shiny above and with a 
triangular patch of short hairs below ; ist antennal segment with a bulbous 
projection below ........ (Megistostylus Bigot) 
- Arista clearly demarked from 3rd antennal segment or at least not as above . . 3 
j Arista apical ......... (Chrysosoma Guerin) 
- Arista dorsal ............ 4 
I 4 scutellar bristles ; acrostical bristles at least as strong as dorso-centrals . 
(Condylostylus Bigot, sensu Becker) 
- 2 scutellar bristles ; acrosticals weaker than dorso-centrals .... 5 
5 3rd antennal segment short triangular, normally not longer than wide Sciopus Zeller 
3rd antennal segment long triangular, almost twice as long as wide (Megistostylus Bigot ?) 
88 D. HOLLIS 
6 2nd antennal segment produced thumb-like into 3rd antennal segment on inner 
surface ; pteropleura often haired * . . . . Syntormon Loew 
2nd antennal segment transverse or at most produced dorsally over 3rd antennal 
segment ............. 7 
7 ist antennal segment haired on dorsal surface ....... 8 
i st antennal segment bare on dorsal surface . . . . . . . " n 
8 Hind coxa with a vertical row of hairs on external surface ; many pro thoracic hairs 
Argyra Macquart 
- Hind coxa with a single external bristle ; i prothoracic bristle .... 9 
9 Hind basitarsus bristled on dorsal surface .... Dolichopus Latreille 
- Hind basitarsus without bristles on dorsal surface . . . . . . 10 
10 Clypeus detached from inner eye margins ; face extending almost to level of lower 
eye margins ; hind femur with a pre-apico-dorsal row of bristles Tachytrechus Walker 
- Clypeus not detached from inner eye margins ; face not extending down to level of 
lower eye margins ; hind femur with external pre-apical bristles . Hercostomus Loew 
11 Face with a complete transverse division . . . . . . . . 12 
- Face with, at most, an incomplete transverse division . . . . . 14 
12 Thorax with a pre-scutellar flattening ; acrosticals biserial ; arista sub-apical or 
apical ; hypopygium pedunculate ...... (Medetera Fischer) 
- Thorax without a pre-scutellar flattening ; acrosticals uniserial or absent ; arista 
dorsal ; hypopygium enclosed .......... 13 
13 Acrosticals uniserial ; 4 or more scutellar bristles . . . (Hydrophorus Fallen) 
- Acrosticals absent ; 2 scutellar bristles .... Thinophilus Wahlberg 
14 Thorax with a strong pre-scutellar flattening ; hypopygium free . . . . 15 
- Thorax normally without a pre-scutellar flattening ; hypopygium enclosed . . 16 
15 Acrosticals biserial ; legs and abdomen relatively long . . Nepalomyia gen n. 
Acrosticals absent ; legs and abdomen of normal length . . Micromorphus Mik 
1 6 Arista dorsal or sub-apical ; antennae placed near middle of head profile ; occiput 
concave ............. 17 
Arista dorsal ; antennae placed in upper third of head profile ; occiput convex . 18 
17 Wings wider in basal third than in apical third ; frons narrower than face ; <J with 
macrochaetae at tip of abdomen ...... Diaphorus Meigen 
- Wings oval ; face narrower than frons ; (J abdomen without macrochaetae at tip 
Chrysotus Meigen 
1 8 Bristles of head and thorax pale ...... Chrysotimus Loew 
- Bristles of head and thorax black ...... Sympycnus Loew 
Chrysosomatinae 
Sciopus pediformis Becker 
Sciopus pediformis Becker, 1922 : 204, fig. 180. 
Described from i $ from Darjeeling 6-7,000'. 
Apart from genitalia differences the $ has a wider face than the < ; tibia i with 
a single dorsal bristle in the basal quarter ; tibia 2 with 2 antero-dorsal, 2 postero- 
dorsal and i ventral bristle ; tibia 3 with i long antero-dorsal, 5 short postero-dorsal 
and 2 short ventral bristles. 
i(,i$, NEPAL : Taplejung Distr., between Sangu and Tamrang, c. 5,600', 
dense vegetation in tree shade by hill stream, 23.x. 1961 (R. L. Coe). 
ON THE DIPTERA OF NEPAL 89 
Dolichopodinae 
Dolichopus angustinervis Becker 
Dolichopus angustinervis Becker, 1922 : 9, fig. 2. 
Dolichopus angustinervis Becker ; Parent, 1934 : 3 O1 - 
Dolichopus angustinervis Becker ; Parent, 1941 : 217. 
Described from N. India and Formosa and later recorded from Honolulu (Parent 
1934) and Nankin, China (Parent 1941). 
i $, NEPAL : Katmandu, 4,500', 20. v. 1935 (F. M. Bailey] ; i $, Bakhri Kharka, 
5,500', 24.iv.i954 (/. Quintan) ; 2 $, Taplejung Distr., Sangu, c. 6,200', mixed 
vegetation by stream in gully, ix-x.igGi (R. L. Coe). 
Hercostomus ulleriensis sp. n. 
(Text-fig. 8) 
5. Frons one-third width of head, completely grey-brown dusted ; i pair of strong orbita 
bristles. Antennae mainly orange ; 3rd segment darkened at tip, as long as wide and bearing 
a pubescent arista dorsally. Face as wide as frons, silvery-white and with a convexity above 
oral margin. Palpi yellow, pale haired. Proboscis orange. Eyes short haired. Occiput 
grey ; post-ocular cilia black. 
Thorax longer than wide ; dorsum metallic green, heavily grey dusted, chaetotaxy normal. 
Scutellum concolorous with dorsum, with 2 strong marginal bristles, each with a short hair in 
front, disc bare. Pleurae heavily grey dusted. 
Abdomen metallic green, short bristled with longer bristles at posterior margins of tergites. 
Legs mainly yellow. Coxae yellow, middle pair with a longitudinal grey stripe externally, 
dark bristled. Femora yellow, middle and posterior pairs with single pre-apical bristles. 
Leg i tibia yellow with 2 antero-dorsal, i postero-dorsal and no ventral bristles ; tarsus 
yellow-brown. Leg 2 tibia yellow with 4 antero-dorsal, 2 postero-dorsal and i ventral 
bristle ; tarsus brown-black. Leg 3 tibia yellow with 3 antero-dorsal, 4 postero-dorsal 
and a row of 6-7 short ventral bristles ; basitarsus yellow, following tarsal segments brown- 
black. 
Wings pale brown hyaline ; 3rd and 4th long veins converging in their apical sections ; 
posterior cross-vein shorter than apical section of 5th vein. Squamae yellow, cilia black. 
Halteres yellow. 
Length 4 mm. 
< unknown. 
Holotype $, NEPAL : 2 miles S.W. of Ulleri, 6-7,000', i8.v.i954 (/. Quintan}. 
Of the Oriental species of Hercostomus, lucidiventris Becker, 1922, described from 
Formosa, is close but may be distinguished by its completely black antennae and 
yellow side-spots on the ist three abdominal segments. 
In keys to Palaearctic species ulleriensis sp. n. will run to novus Parent, 1927, 
described from China, but this species has an almost bare arista, mid and hind 
coxae are dark, fore-tibia has a ventral bristle and the posterior cross-vein is longer 
than the apical section of the 5th vein. 
Hercostomus phollae sp. n. 
(Text-figs. 9, 10) 
c?. Frons less than one-third width of head, completely brown dusted ; orbital bristles 
weaker than inter-ocellars. Antennae orange-brown ; 3rd segment black in apical half, as 
go D. HOLLIS 
long as wide and bearing a micro-pubescent arista dorsally. Face half width of frons, brown 
dusted. Palpi black. Proboscis brown. Occiput grey, post-ocular cilia completely black. 
Dorsum of thorax metallic green, heavily grey dusted, chaetotaxy normal. Scutellum 
concolorous with dorsum with 2 strong marginal bristles. Pleurae grey dusted. 
Abdomen metallic green with strong patches of grey dusting laterally. Hypopygium large, 
black ; external lamellae small, black, subtriangular. 
Legs mainly yellow. Coxae grey with yellow tips and black bristles. Femora yellow, hind 
pair with 4-5 pre-apical bristles. Leg I tibia yellow with i antero-dorsal, 2 postero-dorsal 
and no ventral bristles ; tarsus yellow-brown. Leg 2 tibiae missing from type. Leg 3 tibia 
yellow with 2 antero-dorsal, 4 postero-dorsal and no ventral bristles ; basitarsus yellow, 
following tarsal segments dark. 
Wings clear hyaline ; 3rd and 4th long veins slightly convergent in their apical sections ; 
posterior cross-vein shorter than apical section of 5th vein. Cilia of squamae black. Halteres 
yellow. 
$ similar to $, tibia 2 with 3 antero-dorsal, 2 postero-dorsal and i ventral bristle. 
Length 3 mm. 
Holotype <, NEPAL : Taplejung Distr., above Sangu, c. 6,500', evergreen scrub, 
5-i3.x.i96i (R. L. Coe). 
Paratypes 2 $, Dobhan, c. 3,500', shady places on shrubby slope above R. Tamur, 
21-27.1.1962 (R. L. Coe). 
H. compositus Becker, 1922, described from Satara District, 4,200', appears close 
to phollae sp. n. but has antennal segments i and 2 black, face and frons black and 
yellow fore-coxae. 
Hercostomus kaulbacki sp. n. 
(Text-figs, n, 12) 
<J. Frons metallic green, grey dusted ; orbital bristles much weaker than inter-ocellars. 
Face less than half width of frons, silvery white. Antennae completely black ; 3rd segment 
rounded at apex, as long as wide and bearing a micropubescent arista dorsally. Palpi black, 
grey dusted ; proboscis black. Post-ocular cilia black dorsally, pale laterally and ventrally. 
Thorax twice as long as wide ; dorsum metallic green, yellow-grey dusted and with a median 
longitudinal brown stripe ; acrostical bristles totally absent, other bristles much more hair-like 
than normal. Scutellum concolorous with dorsum with 2 strong marginal bristles. Pleurae 
metallic green, grey dusted. 
Abdomen metallic green, evenly short, dark bristled. Hypopygium large, black ; external 
lamellae large, yellow with a darkened rim and bearing black bristles ; penis long, barbed in 
apical third. 
Legs with coxae grey-brown, dark bristled, yellow at tips. Femora dark yellow, mid and 
hind pairs with single pre-apical bristles. Leg i tibia yellow with i antero-dorsal, 2 postero- 
dorsal and i antero-ventral bristle ; basitarsus yellow, darkened at tip, following tarsal segments 
brown. Leg 2 tibia yellow with 4 antero-dorsal, 2 postero-dorsal and no ventral bristles ; 
tarsus brown. Leg 3 tibia yellow, brown in apical fifth, with 3 antero-dorsal, 3 postero-dorsal 
and 2 short ventral bristles ; tarsus black. 
Wings pale brown hyaline ; anal lobe not developed ; costa thickened between endings of 
ist and 2nd long veins ; 2nd long vein thickened in its basal two-thirds ; 3rd and 4th long veins 
slightly convergent in their apical sections ; apical section of 5th long vein twice as long as 
posterior cross-vein. Squamae and cilia pale. Halteres yellow. 
$ unknown. 
Length 4-5 mm. 
ON THE DIPTERA OF NEPAL gi 
Holotype^, S.E. TIBET : Rong To Valley, 21^.1933, 6,500' (F. Kingdon-Ward 6- 
R. J. H. Kaulback). 
This species is not related to the Oriental fauna but in the Palaearctic region 
crassivena Stackelberg, 1934, described from China, is close but has both 2nd and 
3rd long veins thickened basally, biserial acrosticals and the middle tibia has a 
ventral bristle. 
KEY TO HIMALAYAN Hercostomus 
1 Antennae completely black .......... 2 
- Antennae mainly yellow ........... 3 
2 Fore-coxae yellow ; acrosticals present and biserial ; in <J 2nd vein simple 
caecus Becker 
- Fore-coxae grey ; acrosticals absent ; in $ 2nd long vein thickened in its basal 
two-thirds ........... kaulbacki sp. n. 
3 All coxae grey ; 4-5 pre-apical bristles on hind femur ; 3rd and 4th long veins 
slightly convergent ; in $ face narrow, brown ..... phollae sp. n. 
All coxae yellow ; i pre-apical bristle on hind femur ; 3rd and 4th long veins 
strongly convergent ; in $ face wide, white ..... ulleriensis sp. n. 
Tachytrechus crassitarsis de Meijere 
(Text-figs. 13, 14) 
Tachytrechus crassitarsis de Meijere, 1916 : 237. 
Tachytrechus crassitarsis de Meijere ; Becker, 1922 : 32. 
Described from i < and i $ from Java and recorded from Nepal (Becker, 1922). 
The types have been examined and de Meijere's description is adequate. Figures 
are given of the $ fore-tarsus, penis sheath and ventral lobe. 
Tachytrechus compositus sp. n. 
(Text-fig. 15) 
$. Frons one-third width of head, metallic green, grey-brown dusted. Antennae mainly 
black, ist segment orange apico-ventrally ; 3rd segment rounded, with dorsal arista. Face 
metallic green, yellow-grey dusted, half width of frons just below antennae and widening to 
one and a half times its original width at clypeus, latter separated from inner eye margins and 
extending almost to level of lower eye margins. Palpi and proboscis black. Post-ocular cilia 
black above, pale laterally and ventrally. 
Dorsum of thorax dark metallic green, grey dusted, darker at bases of bristles ; chaetotaxy 
normal. Pleurae metallic green, grey dusted. 
Abdomen metallic green, heavily grey dusted laterally ; short, black bristled but posterior 
margins of tergites bear longer bristles. Hypopygium black ; external lamellae black, sub- 
triangular ; penis sheath serrate laterally. 
Legs with coxae green-grey, dark bristled. Femora green-grey, fore- and middle pairs yellow 
in apical quarter, posterior pair with a sub-apico-dorsal row of 4-5 bristles. Leg i tibia 
yellow, black apically, with 3 antero-dorsal, 2 postero-dorsal and no ventral bristles ; basitarsus 
yellow, black in apical third, following tarsal segments black, longer than wide and not flattened. 
Leg 2 tibia yellow, black apically, with 5 antero-dorsal, 3 postero-dorsal and i ventral bristle ; 
tarsal segments 1-4 each with i long and i short apico-ventral bristle, basitarsus yellow basally 
and black in the apical third, following tarsal segments dark. Leg 3 tibia yellow, black 
D. HOLLIS 
FIGS. 8-15. Hercostomus ulleriensis sp. n. 8. $ wing : H. phollae sp. n. 9. <$ wing. 
10. $ hypopygium : H. kaulbacki sp. n. n. <$ hypopygium. 12. $ wing : 
Tachytrechus crassitarsis de Meijere. 13. <J fore-tarsus. 14. ^ penis sheath and 
ventral lobe : T. compositus sp. n. 15. <J penis sheath and ventral lobe. 
ON THE DIPTERA OF NEPAL 93 
apically, with 6 antero-dorsal, 5 postero-dorsal and a row of 5-6 short ventral bristles ; 
basitarsus black but yellow in externo-basal half, following tarsal segments black, segments 1-4 
each with i long and i short apico-ventral bristle. 
Wings clear hyaline, costa thickened between h and R 1 ; 3rd and 4th long veins convergent 
apically ; posterior cross-vein convex to posterior wing margin and of equal length to apical 
section of 5th vein. Squamae pale, cilia dark. Halteres orange. 
$ similar to (J but with shorter ist antennal segment ; tibia i with 3 antero-dorsal, 2 postero- 
dorsal and 2 postero- ventral bristles ; tibia 2 with 3 antero-dorsal, 4 postero-dorsal, i antero- 
ventral and 2 postero- ventral bristles. 
Length 5 mm. 
Holotype <$, NEPAL : Taplejung Distr., Sangu, c. 6,200', yellow blooms of 
cultivated Compositae, 16-29. x. 1961 (R- L. Coe). 
Paratypes 2 $, same data as Holotype. 
Compositus sp. n. may be separated from the other species of Tachytrechus in 
the Oriental region by the following key : 
1 Femora mainly yellow ........... 2 
Femora mainly dark ........... 3 
2 Antennal segments i and 2 yellow ...... picticornis Bigot 
- Antennae completely black ; wing darkened apically between 2nd and 3rd long veins 
argentatus de Meijere 
3 Tibia 2 with more than 4 ventral bristles ........ 4 
Tibia 2 with, at most, 3 ventral bristles ........ 5 
4 Tibiae brown-black, fore-tarsus of $ simple ..... genualis Loew 
- Tibiae mainly yellow, last 4 segments of < fore-tarsus flattened laterally indicus Parent 
5 Hind basitarsus yellow in basal half on external surface ; tibia 2 with at most 3 
ventral bristles ; in fore-tarsus not flattened laterally . . compositus sp. n. 
- Hind basitarsus completely black ; tibia 2 without ventral bristles ; in $ segments 
2-4 of fore-tarsus flattened laterally ..... crassitarsis de Meijere 
Hydrophorinae 
Thinophilus indigenus Becker 
Thinophilus indigenus Becker, 1903 : 73. 
Thinophilus indigenus Becker ; Becker, 1922 : 37, fig. 27. 
Thinophilus indigenus Becker ; Ribeiro, 1923 : 336. 
Thinophilus indigenus Becker ; Frey, 1925 : 24. 
Thinophilus indigenus Becker ; Parent, 1934 : 36- 
Thinophilus indigenus Becker ; Parent, 1935 : 210 and 528. 
Thinophilus indigenus Becker ; Vaillant, 1953 : 3, figs. 1-9. 
This widely distributed species was described from Egypt (Becker, 1903) and 
recorded from Formosa (Becker, 1922), Nepal (Ribeiro, 1923), Philippines (Frey, 
1925), India (Parent, 1934), Malay Peninsula and N. Borneo (Parent, 1935) and 
Algeria (Vaillant, 1953). 
Rhaphiinae 
Syntormon dukha sp. n. 
(Text-figs. 16-18) 
cj. Frons wide, shining blue-violet. Antennae completely black ; ist segment with 1-2 
bristles dorsally ; 2nd segment haired above and produced thumb-like into inner surface of 
94 D. HOLLIS 
3rd segment, being completely covered by 3rd segment ventrally but only in its apical half 
dorsally ; 3rd segment slightly less than three times as long as wide and bearing an arista sub- 
apically which is as long as 3rd segment. Face narrow, silver pruinose. Palpi black, silver 
pollinose ; proboscis black. Post-ocular cilia uniserial and dark above, multiserial and pale 
below. 
Dorsum of thorax shining metallic green, brown dusted ; acrostical bristles uniserial, 6 pairs 
of dorso-centrals, prothoracic hairs pale. Scutellum concolorous with dorsum with 2 strong 
marginal bristles. Pleurae grey dusted, pteropleura bearing a tuft of long, pale hairs. 
Abdomen long, metallic green, tergite i with post-marginal row of very long, black bristles. 
Hypopygium enclosed. 
Legs with coxa i yellow, pale haired and with black apical bristles ; coxae 2 and 3 dark, 
latter with a single external bristle. Femora yellow, posterior pair brown apically, each with 
a dark externo-ventral streak along its entire length. Leg i tibia yellow with a weak antero- 
dorsal bristle in basal third and 2 weak postero-dorsal bristles in apical third ; basitarsus yellow 
in basal third, remainder black, following tarsal segments black, basitarsus as long as segments 
2-4 together. Leg 2 tibia yellow with 3 antero-dorsal, 2 postero-dorsal and 3 ventral bristles ; 
basitarsus yellow, black in apical fifth, following tarsal segments dark. Leg 3 tibia yellow but 
darkening to black in apical third, with 5 postero-dorsal bristles and an antero-dorsal fringe of 
bristles in apical two-thirds which become longer towards apex ; tarsus black, basitarsus 
slightly shorter than following segment and bearing an unequally bifid black spine in ventral 
third. 
Wings pale brown hyaline ; 2nd and 3rd veins divergent, 3rd and 4th veins convergent ; 
posterior cross-vein longer than apical section of 5th vein. Squamae and cilia pale. Halteres 
yellow. 
9 similar to <$ but with short 3rd antennal segment ; wide face which is convex below ; tibia 
2 with 5 antero-dorsal, 2 postero-dorsal and 3 antero-ventral bristles ; posterior femora without 
externo-ventral brown streak ; posterior tibia completely yellow and without antero-dorsal 
fringe of bristles ; simple hind basitarsus. 
Length 4-5 mm. 
Holotype <$, NEPAL : Taplejung Distr., Sangu, c. 6,200', yellow blooms of 
cultivated Compositae, 16-29. x. 1961 (R- L. Coe). 
Paratypes 2 $, mixed vegetation by stream in gully, xi. 1961-1. 1962 (R. L. Coe). 
S. dukha sp. n. is probably not related to the known Oriental species of Syntormon. 
Of the Palaearctic species pallipes (Fabricius) is close but may be differentiated 
by its apical arista, equally bifid spine on hind basitarsus, and more complete 
dorsal row of longer bristles on hind tibia. 
Syntormon babu sp. n. 
(Text-figs. 19, 20) 
$. Frons wide, shining metallic blue-violet. Antennae completely black ; ist segment bare 
above ; 2nd segment haired above and produced thumb-like into inner surface of 3rd segment ; 
latter not quite three times as long as wide and bearing an arista sub-apically which is not 
as long as 3rd antennal segment. Face narrow, silver pruinose. Palpi and proboscis black. 
Eyes short pale haired. Post ocular cilia black and uniserial above, pale and multiserial below. 
Dorsum of thorax shining metallic green, brown dusted ; acrostical bristles microscopic, 
uniserial ; 6 pairs of dorso-centrals. Pleurae grey dusted, pteropleura weakly pale haired. 
Scutellum concolorous with dorsum with 2 marginal bristles. 
Abdomen long, metallic green. Hypopygium enclosed. 
Legs with anterior coxae yellow, pale haired and with black apical bristles ; mid and 
posterior coxae dark with pale tips, posterior pair with a single external black bristle. Femora 
ON THE DIPTERA OF NEPAL 95 
completely yellow. Leg i tibia yellow with i weak dorsal bristle medially ; basitarsus yellow 
and as long as segments 2 and 3 together, segments 2-5 brown. Leg 2 tibia yellow with 3 
antero-dorsal, i postero-dorsal and 2 antero-ventral bristles ; basitarsus yellow but darkening 
apically, following tarsal segments dark. Leg 3 tibia yellow with a weak antero-dorsal fringe 
of bristles in apical half, and 3 strong postero-dorsal bristles ; basitarsus brown, shorter than 
following segment and with a simple curved spine ventro-medially, following tarsal segments 
black. 
Wings very pale brown hyaline ; 2nd and 3rd long veins divergent, 3rd and 4th long veins 
convergent ; posterior cross-vein shorter than apical section of 5th vein ; anal vein weak. 
Squamae and cilia pale. Halteres yellow. 
$ unknown. 
Length 3-5 mm. 
Holotype <, NEPAL : Taplejung Distr., between Sangu and Tamrang, mixed 
plants by damp cliff in deep river gorge, c. 5,200', i-ii.iQ62 (R. L. Coe). 
This species is related to dukha sp. n. from which it may be easily distinguished 
by the simple spine on the hind basitarsus and the more enclosed 2nd antennal 
segment. 
Syntormon ama sp. n. 
(Text-figs. 21, 22) 
o*. Frons wide, shining metallic blue-green. Antennae completely black, ist segment bare 
above (?), 2nd segment haired above and produced thumb-like into inner surface of 3rd segment, 
latter flask shaped and two and a half times as long as wide, arista apical and half as long as 
3rd antennal segment. Face narrow, silver pruinose. Palpi and proboscis black. Eyes short 
pale haired. Post-ocular cilia uniserial and black above, pale and multiserial below. 
Dorsum of thorax shining metallic green with longitudinal brown stripes in regions of 
acrostical and dorso-central bristles ; acrosticals biserial, 6 pairs of dorso-centrals, prothoracic 
hairs pale. Scutellum concolorous with dorsum with 2 strong marginal bristles. Pleurae grey 
dusted, pteropleura weakly pale haired. 
Abdomen dark metallic green ; hypopygium enclosed. 
Legs with anterior coxae pale, dark at base, with pale hairs and black apical bristles ; mid and 
hind coxae dark, pale at tips, hind pair with a black bristle externally. Femora yellow, hind 
pair darkened apically. Leg i tibia yellow, with i dorsal bristle in basal third ; tarsus brown, 
basitarsus as long as segments 2-4 together. Leg 2 tibia yellow, with 3 antero-dorsal, I 
postero-dorsal and no ventral bristles ; basitarsus black apically but mainly yellow, following 
tarsal segments dark. Leg 3 tibia yellow, brown-black apically, with 3 antero-dorsal, 4 
postero-dorsal and a row of short ventral bristles ; tarsus black, basitarsus simple, shorter than 
following tarsal segment. 
Wings clear hyaline ; 2nd and 3rd veins divergent, 3rd and 4th veins convergent, posterior 
cross-vein shorter than apical section of 5th vein, anal vein weak. Squamae pale, cilia brown. 
Halteres yellow. 
$ unknown. 
Length 2-5 mm. 
Holotype <, NEPAL : Taplejung Distr., Dobhan, c. 3,500', shady places on 
shrubby slope above R. Tamur, 21-27.1.1962 (R. L. Coe). 
S. frivolus Becker, 1922, is related to ama sp. n. but differs in that the acrostical 
bristles are uniserial and the hind femora are completely yellow. 
D. HOLLIS 
FIGS. 1 6-2 2. Syntormon spp., 5. dukha sp. n. 16. ^ wing. 17. <J 2nd and 3rd antennal 
segments. 18. $ hind basitarsus : 5. babu sp. n. 19. hind basitarsus. 20. ^ 2nd 
and 3rd antennal segments : S. ama sp. n. 21. ^ wing. 22. $ 2nd and 3rd antennal 
segments. 
KEY TO NEPALESE SPECIES OF Syntormon 
[ Acrostical bristles completely biserial ; arista apical ; in $ hind basitarsus simple, 
posterior cross-vein shorter than apical section of 5th vein .... ama sp. n. 
- Acrostical bristles irregularly uniserial, arista sub-apical ; in $ hind basitarsus with 
a spine ventrally ............ 2 
z Hind basitarsal spine of <$ bifid ; 3rd antennal segment not extending down dorsal 
surface of pedicel ; posterior cross-vein longer than apical section of 5th vein 
dukha sp. n. 
- Hind basitarsal spine of <$ simple ; 3rd antennal segment extended for some length 
over dorsal surface of pedicel ; posterior cross-vein shorter than apical section of 
5th vein ............ babu sp. n. 
Diaphorinae 
Diaphorus jeanae sp. n. 
(Text-fig. 23) 
<J. Frons metallic green, heavily yellow dusted, less than one-eighth width of head but hardly 
narrower than face ; orbital bristles not developed, interocellars strong. Antennae in centre 
of head profile ; ist segment yellow, bare above ; 2nd segment black with an apical crown 
of short bristles ; 3rd segment black, triangular, hardly longer than wide and bearing a 
pubescent arista sub-apically. Face grey. Palpi yellow with black bristles, proboscis brown. 
Eyes widely separated over frons, short pale haired. Occiput grey, post- vertical bristles 
developed but in same series as post-ocular cilia ; latter black and uniserial above, pale and 
multiserial below. 
ON THE DIPTERA OF NEPAL 97 
Dorsum of thorax metallic green, heavily yellow-grey dusted ; acrostical bristles biserial, 
5 pairs of dorso-centrals, rest of chaetotaxy complete. Scutellum slightly darker than dorsum 
with 2 strong marginal bristles each with a hair-like bristle in front. Pleurae metallic green, 
grey dusted. 
Abdomen with segments 1-3 mainly yellow, segments 4-5 and hypopygium metallic green ; 
2 strong macrochaetae present at tip of abdomen. Lamellae strap-like, not as long as abdomen 
is deep, yellow with black hairs. 
Legs with coxa i yellow with black bristles ; coxa 2 grey, black bristled ; coxa 3 yellow with 
a single black bristle externally. Femora yellow with double ventral fringes of black bristles 
which are slightly shorter than the femur is wide. Leg i tibia yellow without strong bristles ; 
basitarsus yellow and as long as following tarsal segments together, segment 5 with long hairs 
dorsally, pulvilli large. Leg 2 tibia yellow with 2 antero-dorsal, i postero-dorsal and i ventral 
bristle ; basitarsus yellow, following tarsal segments brown, pulvilli large. Leg 3 tibia 
yellow with 3 dorsal bristles ; tarsus brown, basitarsus slightly longer than following tarsal 
segment, pulvilli small. 
Wings clear hyaline ; 2nd vein sinuous, 3rd and 4th veins parallel in their apical sections, 
posterior cross-vein shorter than apical section of 5th vein. Squamae yellow, cilia dark. 
Halteres yellow. 
$. Unknown. 
Length 4 mm. 
Holotype^, NEPAL : Bakhri Kharka, 5,500', 24. iv. 1954 (/. Quintan). 
D. mandarinus Wiedemann, the only species of Diaphorm previously recorded 
from Nepal, is closely related to jeanae sp. n. but may be distinguished by its mainly 
yellow antennae, the much narrower face in the and the shorter genital lamellae. 
Diaphorus sanguensis sp. n. 
$. Frons metallic green, grey dusted, one-third width of head ; orbital and interocellar 
bristles well developed. Antennae in middle of head profile ; ist segment bare above, mainly 
black but with orange-yellow coloration on inner ventral aspect ; 2nd segment black with an 
apical crown of short bristles ; 3rd segment black, wider than long and bearing a pubescent 
arista sub-apically. Face as wide as frons, heavily grey dusted. Palpi yellow with black 
bristles, proboscis black. Occiput grey ; post-ocular cilia black and uniserial above, pale and 
multiserial below. 
Dorsum of thorax shining metallic green, grey dusted ; acrostical bristles biserial, 5 pairs 
of dorso-centrals. Scutellum concolorous with dorsum with 2 strong marginal bristles each 
with a short hair in front. Pleurae metallic green, grey dusted. 
Abdomen with segment i completely yellow, 2nd segment yellow ventrally and in anterior 
half laterally but green dorsally in the mid line, following segments metallic green. 
Legs with coxa i yellow, black bristled ; coxa 2 grey, black bristled ; coxa 3 yellow with a 
single black bristle externally. Femora yellow, without ventral fringes of hairs or bristles. 
Leg i tibia yellow with i antero-dorsal, 2 postero-dorsal and no ventral bristles ; basitarsus 
yellow, following tarsal segments darker. Leg 2 tibia yellow, with 3 antero-dorsal, 3 postero- 
dorsal and 3 ventral bristles ; basitarsus yellow, following tarsal segments brown. Leg 3 
tibia yellow, with 2 antero-dorsal, 4 postero-dorsal and a row of short ventral bristles ; basi- 
tarsus yellow and slightly longer than 2nd tarsal segment, following tarsal segments brown. 
Wings clear hyaline ; 2nd vein slightly sinuous, 3rd and 4th veins parallel apically, posterior 
cross-vein shorter than apical section of 5th vein. Squamae pale, cilia dark. Halteres yellow. 
cj. Unknown. 
Length 4 mm. 
Holotype $, NEPAL : Taplejung Distr., Sangu, c. 6,200', mixed vegetation by 
stream in gully, ix-x.i96i (R. L. Coe). 
D. HOLLIS 
D. sanguensis sp. n. may be separated from the other known species of Diaphorus 
in the Oriental Region by the colour combination of antennae, abdomen and legs. 
Diaphorus mandarinus Wiedemann 
Diaphorus mandarinus Wiedemann, 1830 : 212. 
Diaphorus mandarinus Wiedemann ; Becker, 1922 : 78. 
Diaphorus mandarinus Wiedemann ; Ribeiro, 1923 : 339. 
Diaphorus mandarinus Wiedemann ; Frey, 1925 : 23. 
Diaphorus mandarinus W T iedemann ; Parent, 1934 : 3 O1 - 
A widely distributed species originally described from China (Wiedemann, 1830) 
and later recorded from Formosa and India (Becker, 1922), Nepal (Ribeiro, 1923), 
the Philippines (Frey, 1925) and Central Borneo (Parent, 1934). 
FIGS. 23-25. Diaphorus jeanae sp. n. 23. < 
sp. n. 24. $ wing. 
head, from front 
25. $ antenna. 
Argyra pseudosuperba 
KEY TO HIMALAYAN SPECIES OF Diaphorus 
1 Antennae completely black ; abdomen without yellow coloration ; all coxae dark 
Mrsutipes Becker 
- Antennae partly yellow ; abdomen with yellow coloration ; fore- and hind-coxae 
yellow ............. 2 
2 First antennal segment completely yellow ; 3rd abdominal segment yellow . . 3 
- First antennal segment black above ; 3rd abdominal segment metallic green 
sanguensis sp. n. 
3 2nd and 3rd antennal segments black ; in $ frons hardly narrower than face jeanae sp. n. 
- 2nd antennal segment yellow, 3rd brown ; in $ frons at most one-third width of face 
mandarinus Wiedemann 
Chrysotus discretus Becker 
Chrysotus discretus Becker, 1922 : 89. 
Described from various localities in the Indian Himalayas. 
3 (, 2 $, NEPAL : Taplejung Distr., Sangu, c. 6,200', mixed vegetation by stream 
in gully, 
2 $, 3 , xi. 1961-1. 1962 ; 5 $, 2 , yellow blooms of cultivated 
ON THE DIPTERA OF NEPAL 99 
Compositae, 16-29. x. 1961 ' J c? * ? rotting fruits of Bhor tree on ground, 7-31. x. 
1961 ; 4 c, i $, above Sangu, c. 6,500', evergreen shrubs, 5-13. x. 1961 ; i <, i $, 
edge of mixed forest above Sangu, c. 6,500', I7.x.-i.xi.i96i ; 3 <, between Sangu 
and Tamrang, deep river gorge, c. 5,200', x-xi.i96i ; 2 <, shrubs by path, c. 5,800', 
6.xi.i96i ; i $, river banks below Tamrang bridge, c. 5,500', x-xi.i96i ; i $, Arun 
Valley, above R. Sabhaya, east shore, swept from dwarf bamboos in deep ravine, 
c. 2,000', I2.xii.i96i (R. L. Coe) ; 2 9, Bahkri Kharka, 5,500', 24.^.1954 ; i <$, 
2 miles S.W. of Rambrong, 8,000', 26.^.1954 ; i <, 5 $, 2 miles S.W. of Ulleri, 
6-7,000', i8.v.i954 ; 2 $, S.E. Sikha, 7-8,000', 23^.1954 (/. Quintan). 
Chrysotus pseudocilipes sp. n. 
cJ. Frons shining metallic green, coarsely pitted. Antennae completely black ; 3rd segment 
wider than long, subreniform with pointed tip and bearing a pubescent arista sub-apically. 
Eyes short pale haired, contiguous over face. Palpi yellow, proboscis black. Post-ocular cilia 
black above, pale laterally and ventrally. 
Dorsum of thorax shining metallic green with reddish tinges ; chaetotaxy normal. Scutellum 
concolorous with dorsum with 2 strong marginal bristles. 
Abdomen metallic green, evenly short bristled. Hypopygium enclosed. 
Legs with anterior coxae yellow, dark at base, dark bristled ; mid and hind coxae dark with 
pale tips, hind pair with a long, black bristle externally. Femora yellow, posterior pair dark 
in apical quarter and with 2-3 long bristles apico- ventrally. Leg i tibia yellow with a single 
dorsal bristle in basal third ; basitarsus yellow, almost as long as following tarsal segments 
together, these latter segments brown. Leg 2 tibia yellow with 2 long antero-dorsal, 2 
shorter postero-dorsal and no ventral bristles ; tarsus darkening towards tip. Leg 3 tibia 
yellow, darkened in apical quarter and with an irregular short ciliation on external surface ; 
tarsus brown, basitarsus slightly longer than following tarsal segment and with a short ciliation 
on external surface. 
Wings pale brown hyaline ; 3rd and 4th long veins slightly divergent ; apical section of 
5th vein 4-4 times as long as posterior cross-vein. Squamae and cilia pale. Halteres pale. 
$. Similar to <$ but eyes widely separated over face which is heavily grey dusted. 
Length 2 mm. 
Holotype^, NEPAL : Taplejung Distr., Sangu, c. 6,200', yellow blooms of cultivated 
Compositae, 16-29. x. 1961 (R- L. Coe). 
Paratypes 2 <, i $, mixed vegetation by stream in gully, ix-x.i96i (R. L. Coe). 
This species is closely related to cilipes Meigen, 1824, differing in the irregular 
ciliation of the hind tibia and tarsus, and the absence of a ventral row of bristles 
in apical third of hind tibia. 
Chrysotus kholsa sp. n. 
cj. Frons shining metallic green, coarsely pitted. Antennae completely black ; 3rd segment 
reniform and bearing a pubescent arista dorsally. Eyes short pale haired, contiguous over face. 
Palpi yellow, proboscis dark. Post-ocular cilia black above, pale laterally and ventrally. 
Dorsum of thorax shining metallic green, chaetotaxy complete. 
Abdomen metallic green, evenly short bristled. Hypopygium enclosed. 
Legs with coxae dark, pale at tips, dark bristled. Fore and mid femora broadly brown 
medially, posterior femora dark in apical quarter. Leg i tibia yellow with a single weak dorsal 
bristle in basal third ; basitarsus yellow, following tarsal segments dark. Leg 2 tibia yellow 
with 2 antero-dorsal, 2 postero-dorsal and no ventral bristles ; basitarsus yellow, following 
ioo D. HOLLIS 
tarsal segments dark. Leg 3 tibia yellow-brown, darker apically, with 3 dorsal bristles and 
a regular, short ciliation on external surface ; tarsus brown. 
Wings clear hyaline ; 3rd and 4th veins parallel, posterior cross-vein one-third length of 
apical section of 5th vein. Squamae pale, cilia dark. Halteres pale. 
$. Similar to $ but eyes widely separated over face which is metallic green and grey dusted . 
Length 2-5 mm. 
Holotype $, NEPAL : Taplejung Distr., Sangu, c. 6,200', mixed vegetation by 
stream in gully, ix-x.i96i (R. L. Coe). 
Paratypes 4 $, same data as Holotype ; i $, xi. 1961-1.1962 ; i $, yellow blooms 
of cultivated Compositae, 16-29. x. 1961 (R- L. Coe). 
C. kholsa sp. n. is similar to the previous species from which it may be readily 
distinguished by the leg coloration, the shape of the 3rd antennal segment and the 
chaetotaxy of the hind tibia. 
KEY TO NEPALESE SPECIES OF Chrysotus 
1 Post-ocular cilia completely black ; femora almost completely dark, only extreme 
tips pale .......... discretus Becker 
Post-ocular cilia pale laterally and ventrally ; anterior and middle femora, at most, 
darkened medially ........... 2 
2 Fore-coxae mainly pale ; anterior and middle femora completely pale, posterior pair 
darkened in apical quarter ....... pseudocilipes sp. n. 
- Fore-coxae mainly dark ; femora broadly dark medially . . . kholsa sp. n. 
Argyra pseudosuperba sp. n. 
(Text-figs. 24, 25) 
$. Frons a quarter width of head, metallic blue-violet with grey dusting at edges ; paired 
orbital and interocellar bristles well developed. Antennae completely black ; ist segment 
short, haired dorsally ; 2nd segment extended more than halfway over dorsal and dorso-lateral 
surfaces of 3rd antennal segment ; latter triangular, longer than wide and bearing a micro- 
scopically pubescent arista dorsally which is at least i -8 times as long as 3rd antennal segment. 
Face as wide as frons, parallel sided, silver pruinose. Palpi and proboscis black. Eyes densely 
short pale haired. Occiput metallic green, silver dusted. Post-ocular cilia black and uniserial 
above, pale and multiserial laterally and ventrally. 
Dorsum of thorax metallic green, heavily silver dusted ; acrostical bristles quadriserial, 
6 pairs of dorso-centrals, whole disc with supplementary hairs. Scutellum concolorous with 
dorsum with 4 marginal bristles and supplementary hairs on disc. Pleurae heavily grey 
dusted. 
Abdomen metallic green with tergites silver dusted at their bases laterally; densely short 
bristled, posterior margins of tergites with longer bristles. 
Legs with coxa i yellow, silver dusted and black bristled ; middle and posterior coxae dark, 
posterior pair with an externo- vertical row of dark hairs. Femora yellow, evenly covered 
with moderately long hairs, middle and posterior pairs with single pre-apical bristles. Leg i 
tibia yellow with 4-5 antero-dorsal, 3 posterior dorsal and no ventral bristles ; basitarsus 
yellow, longer than following tarsal segments together, these being brown. Leg 2 tibia 
yellow with 3 antero-dorsal, 2 postero-dorsal and 3-4 short ventral bristles ; basitarsus yellow, 
longer than following tarsal segments together, these being brown. Leg 3 tibia yellow, 
brown at extreme tip, with 4 long antero-dorsal, 4 long postero-dorsal and no long ventral 
bristles, the whole segment being covered with short hair-like bristles ; tarsus black, basitarsus 
as long as and tarsal segment. 
ON THE DIPTERA OF NEPAL 101 
Wings broad, pale brown hyaline ; 2nd and 3rd veins divergent, 3rd and 4th veins parallel 
apically, 4th vein with a strong Z-bend in apical section, posterior cross-vein sinuous and 
shorter than apical section of 5th vein, anal vein almost reaching to wing margin. Squamae 
yellow with dark tips and black cilia. Halteres orange. 
$. Unknown. 
Length 6-5 mm. 
Holotype $, NEPAL : Taplejung Distr., Sangu, c. 6,200', mixed vegetation by 
stream in gully, ix-x.i96i (R. L. Coe). 
Paratype $, INDIA : pres. E. Brunetti. 
A. superba Tagaki, 1960, described from Japan, is very close to pseudosuperba 
sp. n., differing in that the frons is more heavily silver dusted, the squamal cilia 
are pale, wing veins 2 and 3 are more divergent, the 4th wing vein has a less 
accentuated Z-bend, the arista is shorter in relation to the 3rd antennal segment 
and the chaetotaxy of the legs is different. 
Campsicneminae 
Micromorphus albipes (Zetterstedt) 
Hydrophorus albipes Zetterstedt, 1843 : 454. 
Medeterus albipes (Zetterstedt) Raddatz, 1873 : 330. 
Thrypticus bellus Strobl, 1880 : 59. 
Pseudacropsilus claripennis Strobl, 1899 : 124. 
Micromorphus albipes (Zetterstedt) Becker, 1918 : 124. 
Micromorphus albipes (Zetterstedt) ; Parent, 1929 : 194. 
Micromorphus albipes (Zetterstedt) ; Parent, 1938 : 652, figs. 904-907. 
This species is widely distributed in Europe and has been recorded from Costa Rica 
and New Zealand (Parent, 1929) and North Africa (Parent, 1938). 
i <, NEPAL : Arun Valley, below Tumlingtar, River Sabhaya, west shore, 
c. i, 800', dead leaves lying in sun on sandy shore, 22.xii.i96i (R. L. Coe). 
Sympycnus laetus Becker 
Sympycnus laetus Becker, 1922 : 94, fig. 42. 
Pycsymnus laetus (Becker) Frey, 1925 : 21. 
Pycsymnus laetus (Becker) ; Frey, 1928 : 20. 
Sympycnus laetus Becker ; Parent, i932a : 115. 
Originally described from Formosa, New Guinea, Seleo and Singapore and later 
recorded from the Philippines (Frey, 1928) and Sumbawa and Flores (Parent, I932a). 
3$, i $, NEPAL : Arun Valley, R. Arun below Tumlingtar, c. 1,800', 14-23. xii. 1961, 
evergreen shrubs bordering dry stream beds ; 2 ^, i $, above R. Sabhaya, east 
shore, swept from dwarf bamboos in deep ravine, c. 2,000', 12. xii. 1961 (R. L. Coe). 
Sympycnus turbidus Becker 
Sympycnus turbidus Becker, 1922 : 105. 
Pycsymnus turbidus (Becker) Frey, 1925 : 21. 
Sympycnus turbidus Becker ; Parent, I932a : 115. 
Described from Kurseong, Eastern Himalayas (4,700'), and recorded from the 
Philippines (Frey, 1925) and Flores (Parent, i932a). 
102 D. HOLLIS 
i <, NEPAL : Taplejung Distr., Sangu, c. 6,200', mixed vegetation by stream in 
gully, ix-x.igbi (R. L. Coe). 
Sympycnus gummigutti Becker 
Sympycnus gummigutti Becker, 1922 : 95, fig. 53. 
Described from Nepal, Burma and Darjeeling. 
Sympycnus arunensis sp. n. 
(Text-figs. 26-29) 
cJ. Frons shining metallic green-violet. Antennae with segments i and 2 yellow, latter 
with an apical crown of short bristles ; 3rd antennal segment brown, triangular, not longer 
than wide and bearing a pubescent arista dorsally. Eyes short pale haired, contiguous over 
face. Palpi and proboscis yellow-brown. Occiput grey, post-ocular cilia uniserial, black above, 
pale laterally and ventrally. 
Dorsum of thorax yellow with a broad, longitudinal, metallic green stripe medially ; acrostical 
bristles biserial, 6 pairs of dorso-centrals. Scutellum metallic green centrally but broadly 
yellow laterally and apically, bearing 2 strong marginal bristles. Pleurae yellow. 
Abdomen yellow but tergites have small, triangular, black patches dorsally at anterior 
margins. Hypopygium brown ; lamellae yellow, pale haired, pointed oval. 
Legs completely yellow, dark bristled. Fore-tibia without bristles ; fore-tarsal segments 
bear a ventral fringe of short hairs, the basitarsus bearing a longer hair in the basal third. 
Leg 2 tibia bowed, with 3 antero-dorsal, i postero-dorsal and an antero-ventral fringe of hairs 
which become longer towards tibial tip, intermingled with this fringe apically are 5 long, sinuous 
bristles ; basitarsus bears 3 sinuous bristles ventrally and a group of 3 hairs which mat at 
their tips. Leg 3 tibia with 3 dorsal bristles ; basitarsus much reduced, 2nd tarsal segment 
not as long as basitarsus and bearing a bifid lobe apico-ventrally, 3rd segment over twice as 
long as segments i and 2 together and with a ventral row of short bristles. 
Wings pale brown hyaline ; 3rd and 4th veins slightly convergent ; posterior cross-vein 
longer than apical section of 5th vein. Squamae pale with darker tips and brown cilia. Halteres 
yellow. 
$. Similar to <J but with eyes separated over face which is violet above and silver pruinose 
below ; legs not ornamented ; tibia 2 with 3 antero-dorsal, 2 postero-dorsal and 2 antero- 
ventral bristles ; 2nd segment of hind tarsus twice as long as basitarsus. 
Length 3-5 mm. 
Holotype <$, NEPAL : Arun Valley, below Tumlingtar, River Sabhaya, west shore, 
c. 1800', evergreen shrubs on sandy shore, 9-17. xii. 1961 (R. L. Coe). 
Paratype $, east shore of R. Arun below Tumlingtar, c. 1,800', 14-23. xii. 1961, 
evergreen shrubs bordering dry stream bed (R. L. Coe). 
This species is close to maculatus Parent, 1932, and is in the apicalis group, a key 
to which is given below. 
Sympycnus coei sp. n. 
(Text-figs. 30, 31) 
cJ. Frons wide, shining metallic blue-green. Antennae with segments i and 2 completely 
yellow, latter with an apical crown of short bristles ; 3rd segment brown-black, triangular, not 
longer than wide and bearing a pubescent arista dorsally. Eyes short pale haired, contiguous 
over most of face. Palpi yellow, proboscis brown. Occiput grey dusted ; postocular cilia 
uniserial, black above, pale laterally and ventrally. 
ON THE DIPTERA OF NEPAL 
103 
28 
31 
33 
FIGS. 
28. 
30. 
26-33. Sympycnusspp.,S.arunensis8p.n. 26. <^wing. 27. <$ fore-basitarsus. 
$ mid tibia and basitarsus. 29. <$ hind tarsal segments i 3 : S. coei sp. n. 
(J wing. 31. (J hind tarsus : 5. &#o/a sp. n. 32. <J hypopygium. 33. cj wing. 
io 4 D. HOLLIS 
Dorsum of thorax metallic green centrally, broadly yellow at anterior and lateral margins ; 
acrostical bristles absent, 6 pairs of dorso-central bristles. Scutellum metallic green on disc 
but with completely yellow margins and bearing 2 strong marginal bristles. Post-scutellum 
and post-notum with a median longitudinal brown stripe. Pleurae yellow with a dark triangular 
spot at wing base. 
Abdomen with venter completely yellow, tergites triangularly black-brown dorsally except 
tergite i which is completely yellow. Lamellae yellow, pale haired, pointed oval, ventral 
lobes chitinised. 
Legs with coxae yellow, dark bristled. Femora yellow, posterior pair with a few long hairs 
at the base ventrally and a single pre-apical bristle. Leg i tibia yellow, without bristles ; 
tarsus not ornamented, basitarsus as long as tibia and as long as following tarsal segments 
together. Leg 2 tibia yellow with 3 antero-dorsal, i postero-dorsal and 2 ventral bristles ; 
basitarsus yellow and as long as following tarsal segments together, these being darker. Leg 3 
tibia yellow with no long bristles although all bristles are slightly longer than those on the anterior 
and middle legs ; basitarsus yellow, very short and with 2 long, pale hairs ventrally, 2nd 
tarsal segment shorter than basitarsus and with a bilobed ventral appendage, 3rd tarsal segment 
longer than segments 4 and 5 together and with a ventral row of short bristles, tarsal segments 
4 and 5 brownish, the 4th four times as long as the 5th. 
Wings as long as fly, anal corner not developed ; hyaline but with an apical brown spot ; 
posterior cross- vein shorter than apical section of 5th vein. Squamae pale with dark tips 
and brown cilia Halteres yellow. 
$. Similar to $ but eyes separated over face which is silver pruinose (brown from below) ; 
hind tarsus simple, with 2nd segment twice as long as basitarsus and almost twice as long as 
3rd segment ; wings without apical brown spot. 
Length 4-4-5 mm. 
Holotype $, NEPAL : Taplejung Distr., Sangu, c. 6,200', mixed vegetation by 
stream in gully, xi. 1961-1. 1962 (R. L. Coe). 
Paratypes 25 <$, 27 $, same data as Holotype ; 2 ^, 4 $, above Sangu, mixed 
vegetation in dried up ravine, c. 6,800', 16.^1962 ; i $, edge of mixed forest above 
Sangu, c. 6,500', I7.x-i.xi.i96i ; i $, below Sangu, c. 4,000', mixed vegetation on 
sheltered slopes above river, 3^.1962 ; i $, Dobhan, east bank of River Tamur, 
c. 3,500', mixed vegetation by stream in deep gully, i-ii.i962 (R. L. Coe). 
Four paratypes (2 <$, 2 $) bearing same data as Holotype are deposited in the 
Zoologisch Museum, Amsterdam. 
5. coei sp. n. clearly belongs to the apicalis group, the species of which may be 
separated by the following key : 
KEY TO apicalis GROUP OF Sympycnus 
1 Antennae completely yellow ....... gummigutti Becker 
Antennae partly, or completely dark ........ 2 
2 Antennae completely black ; acrostical bristles uniserial ; in <$ wings with 3 small, 
dark spots apically, legs i and 3 ornamented . . thienemanni (Stackelberg 1931) 
Antennae in part yellow ; acrostical bristles biserial or absent .... 3 
3 Acrosticals biserial ............ 4 
- Acrosticals absent ............ 5 
4 3rd antennal segment twice as long as broad ; dorsum of thorax broadly green ; in 
c? tibia 2 without special hairs or bristles .... acuticornis Frey 
3rd antennal segment as long as broad ; dorsum of thorax narrowly green ; in 
tibia 2 with a complete row of special hairs ventrally .... arunensis sp. n. 
ON THE DIPTERA OF NEPAL 105 
5 Wing with an apical brown spot () ......... 7 
Wing clear hyaline ($)........... 6 
6 Posterior cross-vein shorter than apical section of 5th vein . . coei sp. n. $ 
Posterior cross-vein as long as apical section of 5th vein . . apicalis de Meijere $ 
7 All pairs of legs modified in some way ; posterior cross- vein as long as apical section 
of 5th vein ............. 
- Only hind leg modified ; posterior cross-vein shorter than apical section of 5th vein 
coei sp. n. $ 
8 Tibia 2 without longer hairs at the base ventrally ; tibia 3 ventrally with 4 sub-apical 
hairs standing at 90 to tibia ....... apicalis de Meijere $ 
- Tibia 2 with longer hairs at the base ventrally ; tibia 3 without hairs ventrally 
maculatus Parent $ 
Sympycnus khola sp. n. 
(Text-figs. 32, 33) 
(J. Frons shining metallic green, weakly grey dusted. Antennae brown-black ; 3rd segment 
triangular, not longer than wide and bearing a pubescent arista dorsally. Face narrow, silver 
pruinose but appearing black from front. Proboscis and palpi brown. Eyes short pale 
haired, narrowly separated over face. 
Dorsum of thorax shining metallic blue-violet, yellow on anterior and lateral margins ; 
acrostical bristles biserial, 6 pairs of dorso-centrals. Scutellum blue-violet on disc but yellow 
at margins and bearing 2 strong marginal bristles. Pleurae mainly yellow, pteropleura with a 
dark spot below squamae, metapleura brown. 
Abdomen with venter yellow, segments 1-4 brown-black laterally and dorsally, segment 5 
and hypopygium yellow ; latter enclosed ; lamellae yellow, lateral appendages chitinised. 
Legs with coxae yellow, dark bristled, middle pair with a dark spot externally in the postero- 
basal corner, hind pair with i external bristle. Femora yellow, middle and posterior pairs 
with single external bristles. Leg i tibia yellow, without bristles ; basitarsus yellow and as 
long as tarsal segments 2 and 3 together, segments 2-5 brown. Leg 2 tibia yellow with 
3 antero-dorsal, 2 postero-dorsal, 2 antero-ventral and i postero-ventral ; tarsus yellow-brown, 
simple. Leg 3 tibia yellow with i antero-dorsal, 2 postero-dorsal and 3 short ventral bristles ; 
basitarsus yellow and shorter than segment 2 which is simple, segments 2-5 brownish. 
Wings hyaline ; 3rd and 4th veins slightly divergent ; apical section of 5th vein not twice 
as long as posterior cross-vein. Squamae pale with brown cilia. Halteres pale yellow. 
$. Similar to <$. 
Length 2 mm. 
Holotype <, NEPAL : Taplejung Distr., between Sangu and Tamrang, x-xi.igGi, 
mixed shrubs in deep gorge, c. 5,200' (R. L. Coe). 
Paratypes i <, same data as Holotype ; i <f>, mixed plants by damp cliff in deep 
river gorge, c. 5,200', 1-11.1962 (R. L. Coe}. 
S. laetm Becker, 1922, has a superficial resemblance to khola sp. n. but differs in 
that the ist and 2nd antennal segments are yellow and, in the $, the hind tarsus is 
slightly modified. 
Sympycnus gauri sp. n. 
(Text-fig. 34) 
cJ. Frons shining metallic blue-violet. Antennae completely black ; 3rd segment triangular, 
longer than wide and bearing a pubescent arista dorsally. Eyes short pale haired, contiguous 
over face. Palpi and proboscis dark. 
io6 D. HOLLIS 
Dorsum of thorax metallic green ; acrostical bristles irregularly uniserial, 5 pairs of dorso- 
centrals. Scutellum concolorous with dorsum with 2 strong marginal bristles. Pleurae 
metallic green-black, grey dusted. 
Abdomen mainly metallic green, ist segment yellow basally. Hypopygium enclosed. 
Legs with coxae dark, pale at tips, dark bristled, posterior pair with i bristle externally. 
Femora mainly dark but pale basally and apically, anterior and middle pairs with ventral 
fringes of long, dark hairs. Leg i simple, yellow-brown, tibia without bristles or hairs. 
Leg 2 tibia yellow with 2 antero-dorsal, i postero dorsal and no ventral bristles ; tarsus 
brown, simple. Leg 3 tibia yellow with i antero-dorsal, 2 postero-dorsal and no ventral 
bristles ; tarsus simple, basitarsus yellow, shorter than segment 2, segments 2-5 brown. 
Wings hyaline ; 3rd and 4th veins parallel, apical section of 5th vein almost twice as long as 
posterior cross-vein. Squamae brown, cilia black. Halteres pale yellow. 
$. Unknown. 
Holotype <$, NEPAL : Taplejung Distr., river banks below Tamrang Bridge, 
c. 5,5oo', x-xi.ig6i (R. L. Coe). 
Paratype <J, same data as Holotype. 
This species is similar to residuus Becker, 1922, described from Formosa, but the 
latter species may be distinguished by the absence of acrostical bristles, yellow 
coxae and femora, the latter without ventral hair fringes, and the relatively shorter 
posterior cross-vein. 
Sympycnus albipes nepalensis subsp. n. 
(Text-figs. 35-37) 
Sympycnus albipes Lamb, 1926 : 548, figs. 7-10. 
6*. Frons shining metallic blue-green, weakly pale dusted. Antennae mainly brown-black ; 
3rd segment yellowish at base, triangular, not longer than wide and bearing a pubescent arista 
dorsally. Eyes short pale haired, contiguous over face. Palpi and proboscis yellow. Occiput 
grey dusted ; post-ocular cilia uniserial, black. 
Dorsum of thorax shining metallic green, acrostical bristles uniserial, 6 pairs of dorso-centrals. 
Scutellum metallic green on disc but lateral and posterior margins broadly yellow, bearing 
2 strong marginal bristles. Pleurae metallic green above, yellowish below. 
Abdomen with ist segment green, 2nd segment yellow, remaining segments and hypopygium 
green. 
Legs with coxae yellow, dark bristled, posterior pair with a single bristle externally. Femora 
yellow, anterior pair with a few scattered long hairs ventrally, posterior pair with single pre- 
apical bristles. Leg i tibia yellow, without bristles but with a row of 5-6 long hairs ventrally 
in the apical half ; basitarsus yellow, longer than following segment and with a row of short 
hairs ventrally, 2nd segment brown, longer than 3rd segment which is as long as segments 
4 and 5 together, 4th segment with some long hairs dorsally, 4th and 5th segments whitish. 
Leg 2 tibia yellow with 2 weak antero-dorsal and 3 weak ventral bristles ; basitarsus yellow, 
following tarsal segments brown. Leg 3 tibia mainly yellow, broadly brown apically, with 
2 antero-dorsal, i postero-dorsal and no ventral bristles ; tarsus brown, basitarsus short with 
i bristle ventro-medially, 2nd segment shorter than basitarsus and with a long, thin, apico- 
ventral lobe, 3rd segment longer than segments 4 and 5 together. 
Wings hyaline ; 3rd and 4th veins parallel apically, apical section of 5th vein over twice 
as long as posterior cross-vein, anal vein weak. Squamae pale, cilia dark. Halteres yellow. 
$. Similar to $ but with eyes separated over face which is blackish ; legs simple, 2nd seg- 
ment of hind tarsus longer than basitarsus. 
Length 2-2-5 rnm. 
ON THE DIPTERA OF NEPAL 107 
Holotype <, NEPAL : Arun Valley, east shore of R. Arun below Tumlingtar, 
c. 1,800', 14-23. xii. 1961, evergreen shrubs bordering dry stream-beds (R. L. Coe). 
Paratypes i <$, same data as Holotype ; i $, above River Sabhaya, east shore, 
swept from dwarf bamboos in deep ravine, c. 2,000', 12. xii. 1961 (R. L. Coe}. 
S. albipes albipes Lamb, 1926, was described from Rodriguez Island and recorded 
from the Seychelles and Ceylon. The new subspecies may be separated from it by 
the following points :- partly yellow antennae, fewer and longer acrostical bristles, 
much less darkened hind tibial tip and, in the $, the more numerous and shorter 
ventral hairs on the fore-tibia. 
In Becker's key, 1922, to the Oriental species of Sympycnus both subspecies will 
run to strenuus Becker, 1922, but this species has a reddish 3rd antennal segment 
and an unmodified fore-tibia and tarsus. 
Sympycnus peniculitarsus sp. n. 
(Text-fig. 38) 
<J. Frons one-third width of head, shining metallic green-blue, weakly pale dusted. Antennae 
with first and second segments black ; segment 3 brownish yellow, triangular, not longer than 
wide and bearing a pubescent arista dorsally. Eyes short pale haired, contiguous over face. 
Palpi and proboscis dark. Occiput grey ; post-ocular cilia uniserial, black above, pale laterally 
and ventrally. 
Dorsum of thorax shining metallic green with bluish and bronze tinges ; acrostical bristles 
uniserial, 5 pairs of dorso-centrals. Scutellum concolorous with dorsum with 2 strong marginal 
bristles. Pleurae metallic green, grey dusted. 
Abdomen mainly green-black, segment 2 and anterior margin of segment 3 yellow dorsally, 
segments 3 and 4 yellow ventrally. Hypopygium enclosed. 
Legs with coxae pale, dark bristled, posterior pair with i bristle externally. Femora yellow, 
middle and posterior pairs with single pre-apical bristles, posterior pair dark at tip. Leg i 
tibia yellow, without bristles but with a complete ventral fringe of pale hairs ; tarsus yellow, 
2nd segment with a ventral extension in basal two-thirds forming a notch in which is situated 
a group of short, chitinised hairs. Leg 2 tibia yellow, with 3 antero-dorsal, i postero-dorsal, 
i antero-ventral and no postero-ventral bristles ; tarsus brown, simple. Leg 3 tibia yellow, 
with 3 antero-dorsal, 2 postero-dorsal and 1-2 short ventral bristles ; tarsus brown, simple, 
basitarsus shorter than following segment. 
Wings hyaline ; 3rd and 4th veins parallel apically, apical section of 5th vein almost 3 times 
as long as posterior cross-vein, anal vein weak. Squamae pale, cilia dark. Halteres yellow. 
9- Similar to <$ but with eyes separated over face and simple fore-leg. 
Length 1-5-2 mm. 
Holotype , NEPAL : Arun Valley, below Tumlingtar, River Sabhaya, west shore, 
c. i, 800', evergreen shrubs on sandy shore, 9-17. xii. 1961 (R. L. Coe). 
Paratypes 3 <$, same data as Holotype ; 13 $, 9 $, above R. Sabhaya, east shore, 
swept from dwarf bamboos in deep ravine, c. 2,000', 12. xii. 1961 ; 20 <$, 17 $, east 
shore of R. Arun below Tumlingtar, c. 1,800', 14-23. xii. 1961, evergreen shrubs 
bordering dry stream-beds (2 $ and 2 $ of this series deposited in the Zoologisch 
Museum, Amsterdam) ; 2 <$, 4 $, swept from Ricinus communis L., 23. xii. 1961 ; 
13 cJ, 24 <j>, Taplejung Distr., below Sangu, c. 4,000', mixed vegetation on sheltered 
slopes above river, 3.1.1962 (2 $ and 2 $ of this series are deposited in the Entomo- 
logical Institute, Hokkaido University, Sapporo) ; 3 $, shady places on shrubby 
io8 D. HOLLIS 
slope above R. Tamur, 21-27.1.1962 ; i <, 3 $, Dobhan, c. 3,500', mixed vegetation 
in dry gully on wooded slope, 29.1.1962 ; i $, i $, evergreen trees overhanging 
stream in deep gully, 30.1.1962 ; 8 <$, n $, east bank of River Tamur, c. 3,500', 
mixed vegetation by stream in deep gully, 1-11.1962 (R. L. Coe). 
S. turbidus Becker, 1922, is close to peniculitarsus sp. n. but may be distinguished 
from the latter by its black 3rd antennal segment, hind tibia with 3 ventral bristles 
and, in the <$, the simple fore-leg. 
Sympycnus takagii sp. n. 
(Text-figs. 39, 40) 
cj. Frons metallic blue-violet, grey dusted. Antennae black ; 3rd segment triangular, 
almost twice as long as wide, long haired and bearing a pubescent arista dorsally. Face very 
wide, half width of frons, metallic green but heavily grey dusted. Palpi and proboscis dark. 
Eyes short pale haired, widely separated below antennae. Occiput grey ; post-ocular cilia 
uniserial, black above, pale laterally and ventrally. 
Dorsum of thorax metallic blue-green, heavily yellow-brown dusted ; acrostical bristles 
absent, 5 pairs of dorso-centrals. Scutellum concolorous with dorsum with 2 strong marginal 
bristles. Pleurae metallic green, grey dusted. 
Abdomen long, completely metallic green, hypopygium enclosed. 
Legs with anterior coxae yellow, dark bristled ; middle and posterior pairs dark, pale at 
tips, hind pair with single bristles externally. Femora yellow, posterior pair darkened dorsally 
in the apical half. Leg i tibia yellow, without bristles ; basitarsus yellow, following tarsal 
segments brown. Leg 2 tibia yellow with 2 antero-dorsal and i antero- ventral bristle ; 
tarsus yellow-brown. Leg 3 tibia yellow with i antero-dorsal, 3 postero-dorsal and 3 short 
ventral bristles ; tarsus brown, basitarsus shorter than following segment. 
Wings pale brown hyaline ; 3rd and 4th veins parallel in their apical sections, apical section 
of 5th vein over twice as long as posterior cross-vein, anal vein weak. Squamae and cilia 
brown. Halteres brown. 
$. Unknown. 
Length 2 mm. 
Holotype <$, NEPAL : Taplejung Distr., Sangu, c. 6,200', mixed vegetation by 
stream in gully, xi. 196 1-1.1962 (R. L. Coe). 
Paratype $, same data as Holotype. 
This species is distinct from other Oriental species of Sympycnus by virtue of the 
extremely wide face in the <$. S. residuus Becker, 1922, described from Formosa, 
is perhaps closest but may be separated by the completely yellow coxae, the 
relatively shorter 3rd antennal segment and the narrower face in the $. 
Sympycnus pahar sp. n. 
(Text-fig. 41) 
<J. Frons metallic blue- violet, grey dusted. Antennae with segments i and 2 black ; 
3rd segment brown, triangular, as long as wide and bearing a pubescent arista dorsally. Face 
narrow, dark. Palpi and proboscis dark. Eyes short pale haired, narrowly separated over 
face. Occiput grey ; post-ocular cilia uniserial, black dorsally, pale laterally and ventrally. 
Dorsum of thorax metallic green, heavily grey dusted and with a median longitudinal brown 
stripe ; acrostical bristles absent, 5 pairs of dorso-centrals. Scutellum concolorous with 
log 
dorsum with margins narrowly yellow and bearing 2 strong bristles. Pleurae metallic green- 
black, grey dusted. 
Abdomen long, metallic green-black. Hypopygium enclosed. 
Legs with anterior coxae pale, dark at tips and with dark bristles ; middle and posterior 
coxae dark, pale at tips, posterior pair with i external bristle. Femora yellow, hind pair 
without pre-apical bristles. Leg i tibia yellow, without bristles ; tarsus yellow. Leg 2 
tibia yellow with 2 antero-dorsal, i postero-dorsal and no ventral bristles ; tarsus simple, 
yellow. Leg 3 tibia yellow with 2 postero-dorsal and 3 short ventral bristles ; tarsus brown, 
basitarsus shorter than following tarsal segment. 
Wings hyaline ; 2nd and 3rd veins strongly divergent, 3rd and 4th veins parallel in their 
apical sections, apical section of 5th vein over twice as long as posterior cross-vein, anal vein 
weak. Squamae and cilia brown. Halteres yellow. 
$. Similar to <$ but with wider face. 
Length 2 mm. 
Holotype <$, NEPAL : Taplejung Distr., Sangu, c. 6,200', mixed vegetation by 
stream in gully, xi. 1961-1.1962 (R. L. Coe). 
Paratypes i <, same data as Holotype ; i <, below Sangu, by stream in shady 
ravine, c. 6,000', 30. x. 1961 ; i$, between Sangu and Tamrang, mixed plants by 
damp cliff in deep river gorge, c. 5,200', 22. xi. 1961 (R. L. Coe). 
This species is close to takagii sp. n. from which it may be distinguished by the 
completely yellow hind femur and, in the <$, the narrower face and shorter 3rd 
antennal segment. 
KEY TO NEPALESE SPECIES OF Sympycnus 
1 Thorax with pleurae and sides of dorsum broadly yellow ..... 2 
Dorsum of thorax completely metallic green ....... 6 
2 Antennae completely dark ; acrostical bristles biserial ; in <$ legs simple khola sp. n. 
Antennae at least in part yellow ; in $ at least hind basitarsus modified . . 3 
3 Antennae completely yellow ; in <$ fore-tibia with a hair-like bristle apically 
gutnmigutti Beck 
Antennae with i or more segments dark ........ 4 
4 Acrostical bristles present and biserial ........ 5 
- Acrostical bristles absent ; abdomen with segments triangularly black dorsally ; $ 
wing with apical brown spot and only hind basitarsus modified . . . coei sp. n. 
5 Abdomen mainly yellow ; 3rd and 4th wing veins slightly convergent ; posterior 
cross-vein as long as apical section of 5th vein ; in <$ all pairs of legs ornamented 
arunensis sp. n. 
- Abdomen mainly brown ; 3rd and 4th long veins parallel ; apical section of 5th vein 
twice as long as posterior cross-vein ; in $ only hind basitarsus slightly modified 
laetus Beck 
6 All coxae dark, femora dark ; acrostical bristles irregularly uniserial ; in ^ anterior 
and middle femora with long hairs ventrally ..... gauri sp. n. 
- At least anterior coxae pale .......... 7 
7 Only anterior coxae pale ; acrostical bristles absent ; in <J face wide . . 8 
All coxae pale ; acrostical bristles present ....... 9 
8 3rd antennal segment much longer than wide ; hind femora brown in apical half 
dorsally ; anterior coxae completely yellow ..... takagii sp. n. 
3rd antennal segment not longer than wide ; hind femora completely yellow ; 
anterior coxae darkened at tips ....... pahar sp. n. 
9 Scutellum broadly yellow around posterior margin ; in <$ fore-tibia with long hairs 
ventrally and posterior basitarsus modified . . albipes nepalensis subsp. n. 
no D. HOLLIS 
- Scutellum completely green ; in <$ fore-tibia and posterior basitarsus simple . 10 
10 Hind tibia with 1-2 ventral bristles ; in 2nd tarsal segment of fore-leg with basal 
two-thirds extended ventrally and forming a notch which carries a brush of short, 
stout hairs .......... peniculitarsus sp. n. 
Hind tibia with 3 ventral bristles ; in <$ fore-tarsus simple . . turbidus Beck 
Chrysotimus anomalicerus sp. n. 
(Text-figs. 42-44) 
cj. Frons wide, metallic blue-green, weakly pale dusted. Antennae with segments i and 2 
yellow, former bare above, latter with an apical crown of short bristles ; 3rd segment brown- 
black, long haired, rounded basally and with elongate apical region, forming an appendage 
which is thicker and longer than arista ; latter short, only as long as width of frons and borne 
dorsally on 3rd antennal segment. Face narrow, silver dusted. Palpi and proboscis pale. 
Eyes short pubescent, narrowly separated over face. Head bristles yellow, i pair of orbitals, 
i pair of inter-ocellars, post-ocular cilia uniserial. 
Dorsum of thorax shining metallic green, pale dusted. Bristles yellow, acrostical bristles 
uniserial, 5 pairs of dorso-centrals. Scutellum concolorous with dorsum with 2 strong marginal 
bristles. Pleurae metallic blue-green, grey dusted. 
Abdomen metallic green, evenly short, pale bristled. Hypopygium enclosed. 
Legs completely yellow, yellow bristled. Posterior coxae with a single external bristle. 
Femora without bristles. Leg i tibia without bristles ; basitarsus shorter than tibia but as 
long as tarsal segments 2 and 3 together. Leg 2 tibia with 2 antero-dorsal, i postero-dorsal, 
no ventral bristles and a crown of bristles apically ; tarsus simple. Leg 3 tibia with rows of 
short dorsal and ventral bristles ; basitarsus shorter than segment 2. 
Wings hyaline, pointed at tip ; 3rd and 4th veins slightly divergent, apical section of 5th 
vein over twice as long as posterior cross-vein, anal vein weak. Squamae and cilia pale. 
Halteres yellow. 
$. Similar to $ but with short 3rd antennal segment, longer arista and wider face. 
Length 2 mm. 
Holotype <$, NEPAL : Taplejung Distr., below Sangu, by stream in shady ravine, 
c. 6,000', 30. x. 1961 (R. L. Coe}. 
Paratype $, Sangu, c. 6,200', mixed vegetation by stream in gully, ix-x.i96i 
(R. L. Coe). 
This species is only tentatively placed in the genus Chrysotimus as it differs from 
other known species in the extraordinary form of the 3rd antennal segment in the $. 
This structure is very similar to that of a few species in the South American genus 
Kophosoma van Duzee, 1926. 
There are no previous records of Chrysotimus occurring in the Oriental region and 
the well defined uniserial acrostical bristles and the absence of yellow coloration in 
anomalicerus sp. n. render it unlike the known Palaearctic species of the genus. 
NEPALOMYIA gen. n. 
(Text-figs. 45-48) 
Head higher than wide ; frons wide ; i pair of orbital bristles and i pair of inter-ocellar 
bristles, both pairs well developed ; post-vertical bristles not developed ; post-ocular bristles 
uniserial. Antennae placed above middle of head profile, shorter than head ; ist segment short, 
bare above ; 2nd segment short, transverse, and with a crown of apical bristles ; 3rd segment 
ON THE DIPTERA OF NEPAL 
FIGS. 34-44. Sympycnus gauri sp. n. 34. 
35. <J wing. 36. <$ hypopygium. 37. $ 
sp. n. 38. (J 2nd segment of fore-tarsus 
40. (J antenna : 5. pahar sp. n. 41. $ 
sp. n. 42. (J wing. 43. <$ antenna. 44. 
J wing : S. albipes nepalensis subsp. n. 
fore-tibia and tarsus : S. peniculitarsus 
S. takagii sp. n. 39. <J hypopygium. 
hypopygium : Chrysotimus anomalicerus 
9 antenna. 
H2 D. HOLLIS 
triangular, as long as wide and bearing a pubescent arista dorsally. Face wide in both sexes, 
with an incomplete transverse division and not reaching to lower eye margins. Eyes short 
haired. Palpi and proboscis well developed. Occiput concave. 
Thorax longer than wide with a well developed pre-scutellar flattening in posterior third ; 
acrostical bristles biserial, 5 pairs of dorso-centrals, i prothoracic, i humeral, i internal post- 
humeral, 2 notopleurals, i weak sutural, 2 supra-alars and i post-alar. Scutellum semicircular 
with 2 strong marginal bristles. 
Abdomen long with 5 unmodified segments, only tergite i with longer posterior marginal 
bristles ; hypopygium sessile but not enclosed, highly complex. 
Legs long and fragile, bristles weak. Posterior coxa with a single external bristle. Femur 
3 with or without pre-apical bristles. Tibia 2 with an apical crown of bristles. Posterior 
basitarsus shorter than following tarsal segment. 
Wings with 2nd and 3rd veins divergent, 3rd and 4th veins parallel, posterior cross-vein 
slightly before middle of wing and shorter than apical section of 5th vein, anal vein present and 
reaching almost to posterior wing margin. 
Type species : Nepalomyia dytei sp. n. 
The affinities of this genus are obscure but probably the closest known genus is 
Xanthochlorus Loew, 1857. The two genera show some similarities especially in the 
form and complexity of the <$ genitalia (see Becker, 1918 : 129). 
Xanthochlorus differs from Nepalomyia in the following characters : a basal 
arista, absence of acrostical bristles, hind basitarsus longer than the following tarsal 
segment and reduction or absence of the anal vein. 
N. dytei sp. n. 
(Text-figs. 45, 46) 
cj. Frons about one-third width of head, dark green, grey dusted. Antennae black ; 
ist segment short, bare above ; 2nd segment shorter than ist and with a crown of apical 
bristles ; 3rd segment triangular, as long as wide and bearing a pubescent arista dorsally. 
Face half width of frons, concave except above mouthparts, metallic green and grey dusted 
and with an incomplete transverse division. Palpi and proboscis black. Occiput concave, 
dark metallic green ; post-ocular bristles black and uniserial ; post- verticals not developed. 
Dorsum of thorax metallic green, grey dusted, with a well developed pre-scutellar flattening 
in posterior third ; acrostical bristles biserial, 5 pairs of dorso-centrals. Scutellum concolorous 
with dorsum with 2 strong marginal bristles. Pleurae metallic green, grey dusted. 
Abdomen dark metallic green, grey dusted ; evenly short bristled but ist tergite with longer 
posterior marginal bristles. Hypopygium large, sessile but free ; appendages chitinised at 
tip, dorsal pair with hooked tips. 
Legs mainly brown, only trochanters yellow. Posterior coxae with single external bristles. 
Posterior femora with or without pre-apical bristles. Leg i tibia without bristles ; basitarsus 
as long as tibia, 2nd tarsal segment shorter than basitarsus but longer than 3rd segment, latter 
longer than segments 4 and 5 together, these being of equal length. Leg 2 tibia with 2 antero- 
dorsal, 2 postero-dorsal and no ventral bristles ; basitarsus shorter than tibia but as long as 
segments 2 and 3 together. Leg 3 tibia with i antero-dorsal, i postero-dorsal and no ventral 
bristles ; basitarsus two-thirds length of following segment. 
Wings very pale brown hyaline ; 2nd and 3rd veins divergent, 3rd and 4th veins parallel 
apically, posterior cross-vein three-fifths length of apical section of 5th vein, anal vein complete 
almost to posterior wing margin. Squamae brown, cilia black. Halteres pale yellow. 
?. Similar to <J but fore-basitarsus shorter than tibia and the 4th and 5th segments of the 
fore-tarsus are as long as the 3rd segment. 
Length 2-5-3 mm. 
ON THE DIPTERA OF NEPAL 
45 
FIGS. 45-48. Nepalomyia spp., N. dytei sp. n. 45. <$ hypopygium. 
N. confusa sp. n. 47. <J hypopygium. 48. <$ antenna. 
46. 
wing 
Holotype $, NEPAL : Taplejung Distr., between Sangu and Tamrang, spray- 
splashed rocks in deep gorge, c. 5,200', 6-28. xi. 1961 (R. L. Coe). 
Paratypes 2 $, 7 $, same data as Holotype ; 4 $, 1-14.11.1962 ; i $, mixed plants 
by damp cliff in deep river gorge, c. 5,200', 22.xi.i96i ; i <j>, x-xi.i96i, mixed shrubs 
in deep gorge, c. 5,200 (R. L. Coe). 
(2 $, bearing same data as Holotype, deposited in the Zoologisch Museum, 
Amsterdam). 
This species is denned in the key below. 
H4 D - HOLLIS 
N. confusa sp. n. 
(Text-figs. 47, 48) 
(J. Frons one-third width of head, dark green. Antennae yellow-brown ; ist segment 
short, bare above ; 2nd segment shorter than ist and with a crown of apical bristles ; 3rd 
segment triangular, as long as wide and bearing a pubescent arista dorsally. Face one-third 
width of frons, dark metallic green, flat with an incomplete transverse division. Palpi and 
proboscis black. Eyes short haired. Occiput dark, shining ; post-ocular bristles black, 
uniserial ; post- verticals not developed. 
Dorsum of thorax green-brown, posterior calli and lateral margins of scutellum yellowish ; 
pre-scutellar flattening well developed ; acrostical bristles biserial, 5 pairs of dorso-centrals. 
Pleurae metallic green, grey dusted. 
Abdomen brownish-green, venter yellow ; tergites evenly short bristled but tergite i with 
longer posterior marginal bristles. Hypopygium large, sessile but free, appendages not 
chitinised, dorsal pair not hooked. 
Legs mainly yellow, only mid and posterior coxae brown in basal two-thirds. Coxae dark 
bristled, posterior pair with single external bristles. Femora short haired, posterior pair 
without pre-apical bristles. Leg i tibia without bristles ; basitarsus shorter than tibia, 
tarsal segments of decreasing lengths, segment 4 longer than segment 5. Leg 2 tibia with 
2 antero-dorsal, i postero-dorsal and no ventral bristles ; basitarsus shorter than tibia but as 
long as segments 2 and 3 together. Leg 3 tibia with i antero-dorsal, i postero-dorsal, i 
ventral bristle and i longer sub-apical bristle externally ; basitarsus one-third length of 2nd 
tarsal segment. 
Wings hyaline ; 2nd and 3rd veins divergent, 3rd and 4th veins parallel, posterior cross-vein 
half length of apical section of 5th vein, anal vein complete almost to posterior wing margin. 
Squamae yellow, cilia black. Halteres pale yellow. 
?. Similar to 6*. 
Length 2-5-3 mm - 
Holotype <$, NEPAL : Taplejung Distr., Sangu, c. 6,200', mixed vegetation by 
stream in gully, ix-x.i96i (R. L. Coe). 
Paratype $, between Sangu and Tamrang, spray-splashed rocks in deep river 
gorge, c. 5,200', 6-28. xi. 1961 (R. L. Coe). 
The two known species of the genus Nepalomyia may be distinguished as 
follows : 
i Legs mainly brown ; antennae black ; in <J anterior leg with basitarsus as long as 
tibia, tarsal segment 3 longer than segments 4 and 5 together, these being of equal 
length ; hypopygium with appendages chitinised apically and dorsal pair with 
hooked tips ........... dytei sp. n. 
- Legs mainly yellow ; antennae yellow-brown ; in <$ anterior leg with basitarsus 
shorter than tibia, segment 3 slightly longer than segment 4, latter i^- times as 
long as segment 5 ; hypopygium with appendages not chitinised and dorsal pair 
not hooked apically but with a few hairs basally .... confusa sp. n. 
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ON THE DIPTERA OF NEPAL 115 
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n6 
INDEX 
albipes (Zett.) (Micromorphus) 101 
albipes nepalensis subsp. n. (Sympycnus) 106 
ama sp. n. 95 
angustinervis Becker 8y 
anomalicerus sp. n. no 
arunensis sp. n. 102 
babu sp. n. 94 
brunettii sp. n. 86 
coei sp. n. 102 
compositus sp. n. 91 
confusa sp. n. 114 
crassitarsis de Meijere 91 
discretus Becker 98 
dukha sp. n. 93 
dytei sp. n. 112 
gauri sp. n. 105 
gummigutti Becker 102 
hinu sp. n. 84 
indigenus Becker 93 
jeanae sp. n. 96 
kaulbacki sp. n. 90 
khola sp. n. 105 
kholsa sp. n. 99 
laetus Becker 101 
mandarinus Wiedemann 98 
melanurus (Walker) 84 
Nepalomyia gen. n. no 
pahar sp. n. 108 
pediformis Becker 88 
peniculitarsus sp. n. 107 
phollae sp. n. 89 
pseudocilipes sp. n. 99 
pseudosuperba sp. n. 100 
sanguensis sp. n. 97 
takagii sp. n. 108 
turbidus Becker 101 
ulleriensis sp. n. 89