PROC. BIOL. SOC. WASH. 107(3), 1994, pp. 557-568 COLOLOBUS, PSEUDOPIPTOCARPHA, AND TREPADONIA, THREE NEW GENERA FROM SOUTH AMERICA (VERNONIEAE: ASTERACEAE) Harold Robinson Abstract.— ThrQQ genera of Vemonieae subtribe Vemoniinae are described as new from South America. Cololobus includes three species from the Espirito Santo area of Brazil. The type species, C. hatschbachii H. Robinson, is described as new, and combinations are made for Vernonia rupestris Gardner and V. longiangustatus. The genus is unique in the Vemonieae and most of the Ci-chorioideae in the short lobes of the disk corollas. Pseudopiptocarpha is de-scribed from Colombia, and combinations are made for the type, Vernonia elaeagnoides H.B.K., and a second species V. schultzii Karsten, which have small axillary heads, appressed and short-stalked T-shaped hairs, quadrate raphids in the achene wall, and Aynia-Xyx>t pollen. Trepadonia is named from the Andes of northern Peru for the single species V. mexiae H. Rob. The genus is similar to Vernonanthura H. Robinson but differs in the pyramidally thrysoid inflorescences with long racemiform branches and is unusual in the scandent habit. Studies of the neotropical Vemonieae have resulted in removal from Vernonia Schreb. of most species of that area that were once placed in the genus (Robinson 1980, 1987a, 1987b, 1987c, 1988a, 1988b, 1988c, 1989a, 1989b, 1990, 1992a, 1992b, 1993a, 1993b; Robinson & Funk 1987; Robinson & King 1979). Of the few species of Vernonia from South America not trans-ferred, two of the Brazilian species, V. echioides Less, and V. incana Less., have the decumbent stem bases and wholly cy-miform inflorescences of true Vernonia and are considered a peripheral element of that genus. There are other isolated species that have been studied and that are awaiting transfer to existing segregate genera. The re-maining unplaced species of Vernonia in Brazil, Colombia, and Peru that are treated here form three genera here described as new. The Brazilian species are placed here in the new genus Cololobus. The genus Tre-padonia is established for Vernonia mexiae H. Robinson (1981) from Peru. Pseudopip-tocarpha is established for two species from Colombia that have been treated at different times as part of Vernonia or, in the case of one species, as part of Piptocarpha R.Br. The pollen grains are measured in Hoy-er's solution. Cololobus H. Robinson, gen. nov. Type: Cololobus hatschbachii H. Robinson Figs. 1-9 Plantae suffruticosae vel subarborescentes ad 2.5 m altae pauce ramosae. Caules brun-nescentes teretes vix striati non fistulosi tomentelli vel in parte glabri in ramis ju-venitatis lanati, pilis biformibus interdum uniseriatis base 1-5-septatis in cellulis ap-icalibus elongatis interdum sessilis T-for-mibus longe ramosis. Folia altema, petiolis ad 3 cm longis interdum alatis; laminae sub-coriaceae oblongae vel obovatae base acu-minatae margine crenulatae vel serrulatae apice obtusae supra puberulae subtus to-mentellae et dense glandulo-punctatae, ner-
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