PROCEEDINGS OF THE BIOLOGICAL SOCIETY OF WASHINGTON 111(2):320-371. 1998. Systematics of the Raninidae (Crustacea: Decapoda: Brachyura), with accounts of three new genera and two new species Annette B. Tucker Department of Geology, Kent State University, Kent, Ohio 44202, U.S.A. Abstract. — Reexamination of the Raninidae reveals revised relationships of raninid genera, both fossil and Recent. Symethis Weber is removed from the Raninidae and placed in the newly erected Symethidae under the Raninoidea. One subfamily is reestablished, Palaeocorystinae, and several subgenera are elevated to generic status: Notopocorystes McCoy, Eucorystes Bell, and Cre-tacoranina Mertin within the Palaeocorystinae. Lysirude Goeke, within the Lyreidinae, is distinguished as a discrete genus rather than as a subgenus of Lyreidus De Haan. Additionally, three new genera are described: Macroacaena, within the Lyreidinae and Carinaranina and Quasilaeviranina within the Ran-inoidinae. Two new raninid species, Laeviranina goedertorum and Carinarani-na marionae, from the Eocene Hoko River Formation of Washington, U.S.A., are established. Descriptions of three species previously described by Rathbun are emended based upon new fossil material: Carinaranina willapensis (Rath-bun) new combination, Laeviranina lewisanus (Rathbun) and L. vaderensis (Rathbun). The description of Eumorphocorystes sculptus Binkhorst is emend-ed. Phylogenetic relationships within the Raninidae are explored using parsi-mony analyses. A hypothetical phylogeny is established for the Raninidae, including fossil and extant genera. One result of these analyses is the impor-tance of using character states from the oldest recognized species for fossil genera, while continuing to use character states of the type for extant genera. Reexamination of the Raninidae was ini-to Eumorphocorystes Binkhorst, 1857, were tiated as a result of an investigation of fossil incorrectly placed. In order to resolve this decapods recovered from the Eocene Hoko problem, it was necessary to reexamine Eu-River Formation, Olympic Peninsula, morphocorystes and emend the original de-Washington, U.S.A. Two new species of scription. raninids were discovered and are described In order to make complete comparisons from this locality. In addition, many new of fossil raninids from Washington State, it specimens of fossil raninids described by was found essential to examine many other Rathbun (1926) also were collected, adding extant and fossil forms. That effort dem-greatly to the understanding of those spe-onstrated the need to provide an arrange-cies. Three of Rathbun's descriptions are ment that would include fossil and Recent emended herein, those of Carinaranina wil-species. To accomplish this, species were lapensis (Rathbun, 1926) new combination, studied employing traditional systematic Laeviranina lewisanus (Rathbun, 1926), procedures, and were arranged in genera and L. vaderensis (Rathbun, 1926). It has defined by mutually exclusive characteris-been recognized for some time that speci-tics. The generic-level and subfamily-level mens from the Pacific coast of North Amer-arrangements were tested using cladistic ican, which Rathbun (1926, 1932) referred methods.
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