PROCEEDINGS OF THE UNITED STATES NATIONAL MUSEUM 
issued iMf^lVA- Sl^^l ^y ff" 
SMITHSONIAN INSTITUTION 
U. S. NATIONAL MUSEUM 
Vol.84 Washington: 1937 No. 3019 
MOTHS OF THE GENUS RUPELA (PYRALIDIDAE : 
SCHOENOBIINAE) 
By Carl Heinrich 
Bureau of Entomologij and Plant Quarantine, United States Department of 
Agriculture 
This paper is based upon specimens of pyralidid moths in the 
United States National Museum, the British Museum, the American 
iSIuseum of Natural History, and the Cornell University collection. 
I am indebted to these institutions for the loan of specimens, to 
Dr. W. T. M. Forbes for the specimens collected by him in the 
Giiianas, to Dr. H. E. Box for the gift of reared specimens from 
British Guiana and St. Lucia, and especially to W. H. T. Tams for 
giving me the correct application of the Walker and Zeller jiames. 
He made preparations of the genitalia of the Walker and Zeller 
types in the British Museum, compared them with drawings and 
slides that we submitted, and sent me photographs of the type 
slides and helpful notes on the types. I am obliged to him also 
for the loan of the British Museum material. 
Studies of the genitalia in this genus brouglit surprising results. 
What we thought were but two or three white species proved to be 
at least 18 species, sharply defined on characters of the male and 
female genitalia, but so alike in color and so variable in size and 
in Avhat ordinarily are specific differences in venation that they 
could not be separated by external characters. Needless to say, the 
specimens in the various collections were badly mixed, and no reli- 
131080—37 1 2^5 
356 PROCEEDINGS OF THE NATIONAL MUSEUM vol.84 
ance could be placed upon the references in literature to tlie older 
species. Therefore, I have omitted all but original references 
in the s^'nonymv. Distribution as given in this paper is only for 
specimens I have examined. 
Among the male specimens I recognize 20 species, and among 
the female specimens also 20. Unfortunately, in only five species 
could males and females be definitely associated. Therefore, it 
Mas necessary to give separate names to the unassociatcd males 
and females representing undescribed species. I regret having to 
do this, for eventuall}- some of the new names will have to go 
into synonymy; but it may be many years before the sexes are 
associated, and meanwhile we shall need names for the females as 
well as the males. Some temporary double naming is unavoidable. 
Thirty-one species are described as new, 2 from both sexes, 14 
from males only, and 15 from females. 
Two old names are placed in synonymy. 
Genus RUPELA Walker 
I'late 33 
Rupcla Walkep., List of the specimens of lepidopterous insects in the collec- 
tion of the British Museum, vol. 28, p. 523, 1863. — Dyak, Insecutor In- 
scitiae Menstruus, vol. 5, p. 80, 1917. (Genotype: Rupela nivea Walker.) 
Stortcria Barnes and :McDuNN0XjGn, Contr. Nat. Hist. Lepid. North America, 
vol. 2, no. 4, p. 178. 1913. — Dyar, Insecutor Inscitiae ^Menstruus, vol. 1, 
p. 105, 1913. (Genotype: Storteria unicolor Barnes and McDunnough.) 
Labial palpus upturned ; basal segment clothed beneath with long 
hairlike scales; third segment short, acuminate. Maxillary palpus 
well developed, filiform, with scales at apex slightly dilated. An- 
tenna minuteh^ serrate and pubescent, laterally flattened. Thorax 
with expanding hair tuft from tegula. Fore wing with termen 
evenly curved; 12 veins; 2 and 3 from cell before angle; 4 and 5 
from lower angle of cell, approximate, connate or stalked; 6 and 7 
from cell, separate; 10 from the stalk of 8 and 9; 11 from the cell, 
separate from, approximate to, or anastomosing with 12. Hind 
wing with 8 veins; 4 and 5 from loAver angle of cell, approximate, 
connate or stalked. Abdomen long; in female with large, expanded 
anal tuft; eighth abdominal sternite of male with several sclerotized 
areas and seventh sternite with a central sensory scale tuft on caudal 
margin (])1. 33, fig. 45) ; seventh abdominal sternite of female with 
a central longitudinal sclerotized area more or less developed (pi. 33, 
figs. 44, 46). 
Male genitalia symmetrical; uncus stout, basal part enlarged and 
variously modified; gnathos strong, with central area produced 
caudally and strongly sclerotized (beaklike) or thin and only more 
MOTHS OF THE GENUS RUPELA HEINRICH 357 
or less broadened, not produced caudally (bandlike) ; liarpe with 
basal costal process produced, cucullus weakly sclerotized and 
simple; transtilla, when distinguishable, seldom sclerotized through- 
out; anellus w^th shieldlike ventral plate (juxta) and a more or 
less sclerotized dorsal part, which is frequently armed with spines; 
aedeagus moderately long, cylindrical, straight or only slightly bent, 
penis entrance well forward of base; cornuti rarely present; vinculum 
narrow, only slightly produced beyond base of harpe; from inter- 
segmental membrane attaching to base of vinculum, a pair of fine, 
moderately long hair tufts. 
Female genitalia with bursa copiilatrix elongate, very weakly 
sclerotized. simple, with no trace of signa; ductus bursae sclero- 
tized toward genital opening; area about genital opening alwaj^s 
more or less sclerotized, often with a well-developed and deeply 
pigmented genital plate; ovipositor rods moderately long; rods of 
eighth segment collar of abdomen about twice the length of ovi- 
positor rods, strong. 
This genus, as far as I know, is confined to the New World. It 
contains all the white and two of the nonwhite American species 
formerly referred to Topeutis { = Scirpophaga). Five tropical 
American species {hivitta Moschler, perstrkdis Hlibner, rcpugnatalis 
Walker, terreUa Hampson, and irrorata Hampson) are still prop- 
erly referable to Topeutis on venational and palpal characters. 
These are all brown species or have the forewings banded with 
brown. One of the Rupela species has a brown form {tinctella 
Walker) and another {paUidula^ new species) has gray-tinted fore 
wings and fuscous hind wings. All the other species are white and 
not to be distinguished from each other except by their genitalia. 
Rupela is apparently closely allied to Topeutis^ from which it 
differs in having upturned labial palpi and vein 10 of fore wing 
from the stalk of 8-9. In Topeutis the palpi are porrect and vein 
10 is from the cell; very rarely (in a few specimens of terrella 
Hampson) is vein 10 short stalked with 8-9. These differences were 
noted by Dyar in 1913 when he removed Rupela from the synonji'my 
of Scirpophaga. where it had been placed by Hampson in 1896. 
The males divide into two distinct groups, one having a band- 
like gnathos and yellow anal tuft, the other a beaklike gnathos and 
white anal tuft. If a corresponding character can. be found in the 
female genitalia it may be possible to remove the species with the 
bandlike gnathos from Rupela and give them a separate generic 
designation ; but as yet we have no females definitely associated 
with males in this group and, therefore, are not justified in erecting 
a new oenus. 
358 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 84 
There appear to be good specific differences in the shape and size 
of the bursa copuhitrix, but, while this organ has been carefully 
figured in each case, I have not attempted to use it to define species. 
There are plenty of other more obvious characters in the female 
genitalia, and the bursa is so subject to distortion in preparation, 
so difficult to see in balsam, and subject to so much individual varia- 
tion in size or shape that the attempt to use it in classification of 
species would confuse rather than help our definitions. 
KEY TO THE SPECIES OF RUPELA 
Males 
1. Gnathos with central area caudally produced and strongly 
sclerotized (beaklike). Anal tuft white 2 
Gnathos with central area not caudally produced, thin (band- 
like). Anal tuft yellow 13 
2. Dorsal element of anellus spined 3 
Dorsal element of anellus unspined 9 
3. Uncus laterally compressed at apex 4 
Uncus not laterally compressed at apex 5 
4. Aedeagus finely scobinate on venter near apex leucatea (p. 360) 
Aedeagus sphied on lateral margins at apex segrega (p. 3G6) 
Aedeagus with pronounced lateral spur at apex pallidula (p. 365) 
5. Tegumen with projecting spur from each ventrolateral 
margin scitula (p. 374) 
Tegumen simple 6 
6. Penis bearing a line of minute cornuti liberta (p. 364) 
Penis without cornuti 7 
7. Aedeagus with ventral scobinations near apex. One pair of 
long, stout spines on anellus cornigera (p. 371) 
Aedeagus with one lateral margin near apex weakly serrate. 
Spines on anellus numerous, small, scattered albinella (p. 362) 
Aedeagus with apex smooth and labeose ; anellus with two or 
three pairs of minute .spines labeosa (p. 36.3) 
Aedeagus with sclerotized manica, otherwi.se simple ; .spines 
on anellus stout, rather short in a single cluster or in a 
pair of dense combs 8 
8. Spines of anellus a single cluster at one side of dorsal mem- 
branous part g-ibbera (p. 367) 
Spines of anellus arranged as an opposing pair of dense, dark 
combs saetigera (p. 367) 
9. Sacculus of harpe produced at apex into a long, stout spine 
(clasper) 10 
Sacculus not .so produced 11 
10. Clasper a straight spine nivea(p. 370) 
Clasper a curved spine vexativa (p. 371) 
11. Basal part of imcus scobinate and produced backward (cowllike) 12 
Basal part of uncus without spines or scobinations, not pro- 
duced backward tinctella (p. 368) 
12. Aedeagus with apical half greatly narrowed (rodlike) . Cucullus 
of harpe narrow sejuncta (p. 373) 
% 
MOTHS OF THE GENUS RUPELA — HEINRICH 359 
Aedeagus with lateral flauge at apex. Cucullus of harpe 
broad imitativa (p. 372) 
13. Penis bearing a small, serrate cornutus. Dorsal part of anellus 
consisting of a pair of strongly sclerotized, sinuous, irregu- 
larly serrate plates horridula (p. 376) 
Penis without cornutus. Doi'sal part of anellus membranous 14 
14. Apex of aedeagus produced into curved clawlike hook or 
hooks 15 
Apex or aedeagus very slightly produced, but not into hooks 
or claws 16 
15. Apex of aedeagus produced into a single, stout, curved, blunt 
hook adunca (p. 374) 
Apex of aedeagus produced into three heavy claws lumaria (p. 375) 
16. Base of uncus produced backward into forklike process with 
stubby, spined prongs monstrata (p. 377) 
Base of uncus produced backward into a broad concave plate 
bearing numerous heavy, long, curved spines spinifera (p. 377) 
Females 
1. Genital plate a narrovr semicircular band firmly joined to rods 
of eighth segment collar tinctella (p. 368) 
Genital plate not a semicircular band, nor attached to rods of 
eighth segment collar 2 
2. In area caudad of genital opening a small, sclerotized 
nipple sejuncta (p. 373) 
In area caudad of genital opening an external sclerotized pocket 3 
In area caudad of genital opening an internal sclerotized pocket 5 
From area just caudad of genital opening a strongly sclero- 
tized, projecting, hooked process 6 
Area caudad of genital opening smoothly sclerotized or rugose, 
without pockets or protruding processes 7 
3. Genital plate in the form of a st^uit, blunt, thornlike process in 
front of genital opening lara (p. 382) 
Genital plate otherwise 4 
4. Genital opening irregular, more square than circular procula (p. 384) 
Genital opening nearly circular jana (p. 381) 
5. Genital plate large, completely surrounding genital open- 
ing candace (p. 382) 
Genital plate not surrounding genital opening; appres.sed to and 
no wider than ductus bursae orbona fp. 384) 
6. Projecting process from behind genital opening truncate. Ductus 
bursae greatly expanded toward genital opening maenas (p. 383) 
Projecting process bluntly pointed. Ductus bursae not greatly 
expanded toward genital opening nereis (p. 383) 
7. Just within genital opening (from lower margin of the opening) 
a pair of short, dark, hooklike processes faustina (p. 380) 
Just within genital opening a pair of sclerotized disks gaia (p. 380) 
No hooks, disks, or other proces.ses within geriital opening 8 
8. Ductus seminalis forming a loop with ductus bursae just before 
genital opening edusa (p. 379) 
Juncture of ductus seminalis and ductus bursae otherwise 9 
9. Lower margin of genital opening sinuate leucatea (p. 360) 
Lower margin of genital opening angulate or concave 10 
360 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 84 
10. Lower margin of genital opening slightly concave 11 
Lower margin of genital opening deeply concave or angulato 12 
11. Without defined genital phate. Genital opening of moderate 
width antonia (p. 378) 
With genital phite well defined. Genital opening very wide (as 
wide as the plate) segreg'a (p. 366) 
12. With defined genital plate 13 
Without defined genital plate beiidis (p. 378) 
13. Genital plate completely surrounding genital opening; latter 
moderately wide 14 
Genital plate not surrounding genital opening ; latter as wide as 
the plate drusilla (p. .379) 
14. Genital plate rugose, especially toward caudal margin herie (p. 381) 
Genital plate smooth 15 
15. Bursa copulatrix very short, much reduced canens (p. .379) 
Bursa copulatrix normal albinella (p. 362) 
RUPELA LEUCATEA (Zeller) 
Plate 22, Figures 1-ld; Plate 30, Figure 3: > ; Plate 33, Figukes 44, 45, 48 
■ScirpopJiaga leucatea Zeller, Chilonidarum et crambidarum genera et species, 
p. 2, 1863; male and female. 
Scirpophaga lonriicornis McIschler, Abh. Senck. n;iturf, Ges., vol. 16. p. 321, 
1890 (new synonymy) ; male and female. 
Zeller described his leucatea from Brazil and St. Thoma.s. Later ^ 
he recorded it from Puerto Eico, Mexico, and Panama. I omit the 
latter reference from the synon3any, as it is quite likely that he had a 
mixed series before him. Until genitalia of all his paratypes are 
examined we can be sure only of his West Indian specimens and the 
actual type. Mr. Tams has examined the genitalia of the latter and 
checked them with my figures. I am indebted to him for the iden- 
tifications. As far as we know there is only one pure white species 
of Rupela found in the Antilles; therefore longkomk Mo.schler is 
presumably the same as leucatea. 
Male. — ^Wings shining white. Fore wing with veins 11 and 12 sep- 
arate ; 4 and 5 connate or shortly stalked. Hind wing with 4 and 5 
connate or stalked. Anal tuft white. 
Alar expanse, 22-38 mm. 
Genitalia with gnathos beaklike but having rather promin.ent lat- 
eral arms; in ventral aspect with basal half rounded (central part 
of lower margin convex) ; apical half tapering to blunt point; apex 
not appreciably upturned; inner surface near apex finely scobinate. 
LTncus stout ; basal portion broad with moderately wide dorsal 
groove; laterally compressed on dorsum at apex; from side view, 
apex broad, slanting; vicAved from beneath, triangularly pointed. 
Harpe widest just before middle, very slightly tapered to broadly 
1 Horae Soc. Ent. Itossicae, vol. 13, p. 6, 1877. 
% 
MOTHS OF THE GENUS EUPELA HEINRICH 361 
rounded apex; basal costal process triangularly produced, not 
strongly sclerotized; sacculus very slightly produced at apex. An- 
ellus consisting of ventral plate and a rugose dorsal piece, the latter 
bearing three minute spines on each of the ends attached to aedea- 
gus; ventral plate with lateral margins concave. Aedeagus cylin- 
drical, constricted somewhat at outer third, expanded laterally just 
before apex and finely scobinate on venter near apex. 
FeTrmle. — Wing color and venation as in the male except that veins 
11 and 12 are slightly anastomosed in one specimen. Anal tuft 
yellow. 
Alar expanse, 25-53 mm. 
Genitalia without defined genital plate but with area between 
genital opening and collar sclerotized and markedly rugose; lower 
margin of genital opening sinuate; ductus bursae sclerotized only at 
genital opening. 
Tyyes.—ln British Museum {leucatea) ; Berlin Museum {Jongi- 
cortiis). 
Type localities.— Kio de Janeiro, Brazil {leucatea) ; Puerto Rico 
{longlcorms). 
Food plant.— Echinochloa polystachya. This food-plant record is 
from specimens reared by Dr. H. E. Box, St. Lucia, October 2, 1934. 
Distribution.— Jamaica; Cuba, Baragua (May), Matanzas (Au- 
gust); HisPANiOLA, Sanchez (May, June), Rio Yaque (February); 
Puerto Rico, Santa Rita (July), Mayaguez (December), Bayamon 
(June), Catano (July), Rio Piedras (December), Dorado (May), 
Desengano (May, December), Toa-Baja (January, February); 
Guadeloupe (December); Grenada, Balthazar; Martinique; St. 
Lucia, Rosseau (August, September, October) ; Antigua, Bendals 
(October) ; Mexico, Teapa (Tabasco, January) ; Guatemala, Qui- 
rigua (October); Honduras; Nicaragua; Panama, La Chorrera 
(April, May) ; Trinidad ; Venezuela, Aroa ; French Guiana, St. Jean 
Maroni; British Guiana, Georgetown (July), Kartabo (October), 
Mackenzie (June) ; Surinam, Moengo (May), Para District (April), 
Paramaribo (May); Brazil, Rio Madeira (July-August), Tapera, 
Rio Campo Bello, Rio Jurna (July), Reyes (Beni River, July), 
Prainho (November), Itatoro (Rio Madina, February), Maranhoa; 
Argentina, Villa Ana "F.C.S.F." (February, March, December) ; 
Paraguay, Villarrica (March), Sapacay (September, November), Rio 
Pacaya (July) ; Peru, Madre de Dios. 
Remarks. — One hundred and nine specimens (48 males and 61 
fem.ales) examined, from the collections of the United States National 
Museum, British Museum, Cornell University, and American Mu- 
seum of Natural History. 
362 PROCEEDINGS OF THE XATIOXAI. MUSEUM vol.84 
The species is readily identifiable by the scobinate aedeagus and 
apically compressed uncus of the male, the distinct rugositj'^ of the 
membranous area behind the gentital opening, and the sinuate mar- 
gin of the genital opening in the female. 
RUPELA ALBINELLA (Cramer) 
Plate 22, Figures 2-2(/ ; Pl.\te 29, Figxjbes 2G-26?> 
Scirpophof/a ulhineUa Cila.mer, Papilloiis oxotiques des trois parties du 
moiKle . . ., vol. 4, pi. 372, fig. D, 17S1 ; female. 
Cramer's figure would fit almost any of the white species with yel- 
low anal (female) tuft treated in this paper, and the name has been 
indiscriminately applied. Inasmuch as the type is nonexistent, I am 
fixing the name to the species apparently most abundant in the type 
localit}^ Dr. W. T. M. Forbes captured a male and female in copula, 
and we are therefore able to associate the sexes. 
Male. — Wings white. Fore wing with veins 11 and 12 separate or, 
for a short distance, closely approximate ; 4 and 5 connate or shortly 
stalked. Hind wing with 4 and 5 connate or stalked. Anal tuft 
white. 
Alar expanse. 20-34 mm. 
Genitalia with gnathos beaklike but having rather prominent lat- 
eral arms (attaching to tegumen) ; in ventral aspect basal half 
rounded (central part of lower margin convex) ; apical half tapering 
slightly to bluntly pointed apex; apex upturned (lateral view) ; inner 
surface (under high magnification) finely granulate, not spined or 
serrate. Uncus moderately stout ; basal part with central dorsal exca- 
vation; a short but prominent dorsal keel near base; apical half digi- 
tate, very slightly broadened near apex ; apex rounded. Harpe with 
lower margin indented near cucullus ; apex rounded ; basal process of 
costa not strongly sclerotized nor much produced; sacculus without 
apical projection, simple. Anellus consisting of ventral plate and a 
rugose, sclerotized dorsal piece, the latter bearing several small, 
inconspicuous spines; ventral plate w^itli lateral margins concave. 
Aedeagus cylindrical, slightly widened at apex; one lateral margin 
near apex weakly serrate. 
Female. — Wing color and venation as in the male. Anal tuft 
yellow. 
Alar expanse, 27-45 mm. 
Genitalia with genital plate well defined, sclerotized, its caudal 
margin angulate; genital opening rigid, its lower margin semicircu- 
lar; ductus bursae strongly sclerotized toward genital opening. 
There is some variation in the size of the genital opening and in the 
width of the ductus in specimens from different localities — especially 
% 
MOTHS OF THE GENUS KUPELA HEINRICH 363 
as between larger specimens from central Brazil and smaller females 
from the Guianas and Central America. I am unable, however, to 
find any characters that would seem to indicate distinct races. 
Type. — Lost. 
Type locality. — Surinam. 
Dhtribution. — Mexico, Presidio, Atoyac (Veracruz), Teapa 
(Tabasco), Misantla (May) ; British Honduras, Cayo; Guatemala, 
Quirigua (April), Tiquisati (May), Volcan Sta. Maria (March); 
Honduras, Lancetilla (June); Costa Rica (April), Port Limon 
(January), Sixola River (September); Panama, Bugaba, David, 
Tabernilla, Cabima (May), Rio Trinidad (June), Corozal (August) ; 
Trinidad, St. Augustine (November) ; French Guiana, St. Laurent 
Maroni, St. Jean Maroni; British Guiana, Georgetown (July, 
November), Mackenzie (June); Surinam, Paramaribo (April, 
June), Moengo (May), Surinam River (St. Barbara Plantation, 
April) ; Brazil, Manaos, Pernambuco, Parintins (June), Parana de 
Buyassu (January), Itacoatiara (November), Breves (January), 
Para, Urucaca (November), Rio Jurna (November), Rio Jutatie 
(January), Rio Madeira (May), Faro (April), Pariti (Rio Purus, 
October), Ponte Nova (Rio Xingu), Taperinha, Sao Paulo de 
Olivenca (November-December), Amazon River between Teffe and 
Tonantins (November) ; Colombia, Magdalena Valley, Rio Condeto 
(Choco, December) ; Ecuador (no other locality) ; Peru, Rio Ucayali 
(December). 
Remarks. — One hundred and eight specimens (33 males and 75 
females) examined, from United States National Museum, British 
Museum, and Cornell University collections. 
The species is easily identified by the serrations on the apical end 
of the aedeagus, the spining on the dorsal plate of the anellus, the 
dorsal keel on the uncus, and the peculiarly shaped female genital 
plate and genital opening. 
RUPELA LABEOSA, r.ew species 
Plate 22, FiciruEs 3-3(Z 
Male. — Wings white. Fore wing with some dark shading on 
under side in costal area above cell; veins 11 and 12 approximate 
(but nowhere touching) ; 4 and 5 closely approximate or connate. 
Hind wing with 4 and 5 closely approximate or connate. Anal tuft 
white. 
Alar expanse, 19-21 mm. 
Genitalia with gnathos beaklike but having rather long lateral 
arms (forming attachments to tegumen) ; in ventral aspect basal 
half broadly rounded, apical half narrow, tapering to apex; inner 
surface finely serrate toward apex. Uncus stout; basal part very 
364 PROCEEDINGS OF THE NATIONAL MUSEUM vol.84 
broad, somewhat bulged but not extended backward into cowllike 
lobe, its central dorsal area evenly excavate; apical half digitate; 
apex blnnt and slightly hooked. Harpe with apex rounded (cucullus 
tapering) ; basal process of costa produced (the basal processes of 
the right and left harpes fusing to form a complete, sclerotized 
transtilla) ; sacculus folded upward toward apex (an appreciable 
depression in harpe just above sacculus). Anellus consisting of 
ventral plate and a sclerotized, somewhat roughened dorsal piece, 
the latter with a very few minute spines (2 or 3) on the ends at- 
tached to aedeagus. Aedeagus cylindrical, smooth; apex flaring 
into a wide mouth (labeose). 
Type and parafypes. — U.S.X.M. no. 51856. Paratypes also in 
British Museum. 
Type locality. — Castro, Parana, Brazil. 
Reinarl's. — Described from male type and six male paratypes from 
the type locality (four specimens collected by Wm. Schaus, three by 
E. D. Jones, no dates). 
Easily recognized by its labeose aedeagus and completely formed 
transtilla. 
Female unknown. 
RUPELA LIBEItTA, new species 
Plate 23, Fic.tniES 4-4c 
Male. — Wings white. Fore wing with veins 11 and 12 anastomos- 
ing; 4 and 5 connate. Hind wing with 4 and 5 connate. Anal tuft 
white. 
Alar expanse, 20-25 mm. 
Genitalia with gnathos beaklike; lateral arms developed; in ventral 
aspect with basal half rounded (central part of lower margin deeply 
convex) ; apical half rather narrow, not appreciably tapering; apex 
bluntly pointed: in lateral aspect gnathos distinctly upcurved. 
Uncus Avith basal part broad and stout and deeply, evenly, and rather 
widely excavate ; apical half digitate ; apex rounded. Harpe slightly 
narrowed at cucullus; apex rounded; basal process of costa pro- 
duced into short digitus; sacculus produced at a]iex as a rather 
prominent upfolded ridge. Anellus consisting of ventral plate and 
a strongly sclerotized dorsal band, which partially encircles aedeagus ; 
each extremity of dorsal band bearing a cluster of very dark, mod- 
erately long, stout spines; ventral plate with upper margin broadly 
incised and lateral margins concave. Aedeagus cylindrical, rather 
slender, of nearly equal width throughout, a few minute scobinations 
on under surface near apex ; penis bearing a thin, short line of minute 
cornuti. 
Type and paratypes.— U.S.'N.M. no. 51857. Paratype also in 
British Museum. 
% 
MOTHS OF THE GENUS RUPELA HEIXRICH 365 
Type locality.— Duviiwgo, Mexico (C. C. Hoffman, "276"). 
Remarks. — Described from type and one paratype from the type 
locality, one paratype from Colima, Mexico (Schaus, collector), one 
paratype from Jalapa, Mexico (Schaus), and one paratype from 
Cabima, Panama (A. Busck, May 20, 1911). 
The species may be recognized by the characteristic spining of the 
dorsal part of the anelliis and the line of fine cornuti on the penis. 
Only two of the species treated in this paper show any trace of 
cornuti or a cornutus. 
Female unknown. 
RUPELA PALLIDULA, new species 
Pi^iTE 23, Figures 5-5<Z 
Male. — Head and palpi as in the pure white species. Thorax 
white but with collar darker, concolorous with fore wing. Fore 
wing silvery buff, unicolorous; cilia snow white; veins 11 and 12 
separate; 4 and 5 stalked. Hind wing darker than fore wing, gray 
to grayish broAvn, cilia snow white; veins 4 and 5 stalked. Wings 
concolorous beneath, grayish broAvn. Anal tuft white. 
Alar expanse, 24-32 mm. 
Genitalia with gnathos beaklike but with lateral arms well de- 
veloped; basal, projecting part rather narrow, truncate; inner sur- 
face very finely scobinate; apex upturned. Uncus with basal part 
humped, subquadrate, rather deeply grooved posteriorly and some- 
what rugose; apical half digitate, laterally compressed on dorsum 
at apex. Harpe tapering slightly at apex; apex rounded; basal 
process of costa greatly produced, digitate (the basal processes of 
the two harpes united by a bit of membrane at tlieir apices) ; sac- 
culus produced at apex into a minute spine. Anellus consisting of 
ventral plate and dorsal membrane, the latter bearing two pairs of 
short, stout spines; ventral plate broadly incised on upper margin, 
lateral margins concave. Aedeagus witli a pronounced lateral spur 
at apex. 
Type and paratypes. — U.S.N.M. no. 51858. Paratypes also in 
British Museum. 
Type locality. — Castro, Parana, Brazil. 
Remarhs. — Described from male type and 12 male paratypes from 
the type locality and 1 male paratype from Sao Paulo, Brazil. 
The above were in the National Museum collection and British 
Museum, identified as tinctella \Yalker. The species is easily recog- 
nized by its color, the spining of the anellus, and its characteristic 
aedeagus. 
Female unknown, probably white. 
366 PROCEEDINGS OF THE NATIONAL INIUSEUM vor.. 84 
RUPELA SEGREGA, new species 
Plate 23, Figures 6-6d : Plate 30, Figure 32 
Male. — Wings shining white. Fore wing with veins 11 and 12 
separate; 4 and 5 approximate or connate. Hind wing with 4 and 
5 connate or stalked. Anal tuft white. 
Alar expanse, 26-33 mm. 
Genitalia with gnathos beaklike but with lateral arms (attaching 
to tegumen) developed; posterior margin of central basal part 
rounded; inner surface finely scobinate; apex upturned. Uncus with 
basal part slightly humped, stout, subquadrate, deeply excavate 
posteriorly and dorsally; apical two-thirds digitate, laterally com- 
pressed on dorsum at apex (as in leucatea and 'pallidula). Harpe 
simple; cucullus but slightly narrowed; basal process of costa mod- 
erately^ produced, fusing with a very weakly sclerotized transtilla; 
sacculus not i)roduced at apex. Anellus consisting of ventral plate 
and a somewhat roughened dorsal piece, tlie latter bearing one pair 
of long, stout spines and two pairs of shorter spines; ventral plate 
with upper margin incised, lateral margins deeply incised, Aedea- 
gus Avith a})ex cleft, l:iterally expanded and spined on lateral 
margins. 
Female. — "Wing color and venation as in the male. Anal tuft 
white. 
Alar expanse, 28-38 mm. 
Genitalia with genital plate Avell defined and somewhat similar 
to that of aJlmieUa but with caudal margin more acutely angled; 
genital opening almost as wide as plate, the lower (outer) margin 
concave; ductus bursae strongly sclerotized (and brown) for a short 
distance from genital opening. 
Type and paratypes. — U.S.N.M. no. 51859. Paratypes also in 
British Museum. 
Tyj)e locality. — South Bay, Lake Okeechobee, Fla. 
Remarks. — Described from male type, eight male and five female 
paratypes, the paratypes distributed as follows : Florida, Glenwood, 
one male, Fort Meade (April), two males and one female, Coconut 
Grove (E. A. Schwarz), two males, Koyal Palm State Park (F. M. 
Jones, March), one male (W. S. Blatchley, April) and one female. 
South Bay, one female, Dade City (September), one male, Biscayne 
Bay, one female; North Carolina, Havelock on Lake Ellis (F. Sher- 
man, June), one female; also one male without any locality label. 
A North American species apparently confined to the southern 
part of the United States. Specimens of segrega (as well as sejrmcta 
and white Florida females of tincteUa) have hitherto been identified 
as alhlnella Cramer. The latter as far as I know does not occur 
in the United States. 
MOTHS OF THE GENUS KUPELA HEINEICH 367 
RUPELA GIBBERA, new species 
Plate 24, Figxtres 7-7d 
Male. — Wings shining wliite. Fore wing with veins 11 and 12 
separate; 4 and 5 connate. Hind wing with -4 and 5 connate. Anal 
tuft white. 
Alar expanse, 23 mm. 
Genitalia with gnathos beaklike the lateral arms developed ; central 
part rather broad and stubby, tapering slightly to bluntly pointed 
apex; inner surface evenly and markedly granulate. Uncus heavy; 
basal part subquadrate, humped, deeply excavate on dorsum, the 
margins of the excavations rugose ; apical part digitate, apex bluntly 
pointed. Harpe with costal process short, the latter fusing into a 
partially sclerotized but appreciable and complete transtilla; cucul- 
lus very slightly tapered; apex rounded: sacculus produced at apex 
into an upturned, rounded, sclerotized. plateiike protuberance. An- 
ellus consisting of ventral plate and a membranous dorsal part: from 
one side of the latter projects a curved sclerotized band which bears 
at its extremity a single cluster of stout, short spines. Aedeagus 
with greatly elongated, dark, rigid manica {Ma., pi. 24, fig. 75) ; 
otherwise simple. 
Type. — In Cornell University collection. 
Type locality . — Moengo, Boven, Cortica Kiver, Surinam (W. T. M. 
Forbes, May 23, 1927). 
Bemarks. — Described from one male. May be really identified by 
the uncus, the single lateral spine cluster on the anellus, and the 
enlarged and stronglv sclerotized manica of the aedeagus. 
Female unknown. 
RUPELA SAETIGERA, new species 
Platk 24, Figures S-Sc 
Male. — Wings shining white. Fore wing with veins 11 and 12 
separate; 4 tnd 5 connate. Hind wing with 4 and 5 connate. Anal 
tuft white. 
Alar expanse, 23 mm. 
Genitalia with gnathos beaklike ; lateral arms developed : central 
part with truncate lower margin ; apical part tapering, aj^ex pointed ; 
inner surface with few and very weak granulations. Uncus heavy; 
basal part much enlarged, subquadrate, only slightly grooved on 
dorsum and with lateral-dorsal angles pointed and slightly produced; 
apical part rather short (in comparison to other species), tapering 
slightly, a central dorsal ridge running its entire length; apex blunt. 
Harpe constricted at cucullus: cucullus small, bluntly pointed; basal 
costal process slightly produced; sacculus broad, surface concave, 
368 PROCEEDINGS OF THE NATIONAL MUSEUM vol.84 
apex bluntly triangular and very slightly produced, free edge mi- 
nutely serrate. Anellus consisting of ventral plate and dorsal mem- 
brane; the latter bearing an opposing pair of long, dense, very dark 
spine combs; spines numerous and moderately stout; lateral margins 
of ventral plate concave. Aedeagus slender, very slightly tapering; 
fused manica sclerotized for a short distance; otherwise simple. 
T'y;?^.— U.S.N.M. no. 518G0. 
Type locality. — Castro, Parana, Brazil (W. Schaus). 
Rejnarks. — Described from one male. ^lay be at once recognized 
by the bluntly triangular apex of the sacculus, the spine combs on 
the anellus, and the shape of the basal part of the uncus. 
Female unknown. 
RUPELA TINCTELLA (Walker) 
Plate 24, FicrnES 9-9c; Plate 32, Figlt.es -12, 4.3; Pl\te 33. Figuke 4G 
Salapola tincteUa Walker, List of the specimens of lepidopterous insects in the 
collection of the British INIusenm, vol. 28, p. 526, 1863 ; female. 
Scirpophaga selleri Moschler, Verh. zool.-bot. Ges. Wien, vol. 31, p. 435, 1882 
(new synonymy) ; female. 
Scirpophaga holophaealis Hampson, Ann. Mag. Nat. Hist., ser. 7, vol. 14, p. 181, 
1904 (new synonymy) ; male. 
Storteria unicolor Baisnes and McDunnough, Contr. Nat. Hist. Lepid. North 
America, vol. 2, no. 4, p. 178, 1913 ; male. 
Rupela holophaealis (Hampson) Dyar, Insecutor Inscitiae Menstruus, vol. 5, 
p. 80, 1917. 
I am indebted to Mr. Tarns for the identification of this species. 
He sent me a description and a photograph of the genitalia of 
Walker's type (a female) and compared a slide and a specimen I 
submitted. jNIoschler's species we know only from his description, 
but that leaves little doubt as to what he had before him. He says 
that the fore wing is "gelblich angehauctes Weiss." This (since 
his type is a female) could apply only to the female of fhicteUa. 
Hampson's JwlophaealL'^ I associate on the evidence of distribution 
and a female of tinctellu in the National Museum collection, which 
matches in color the paler fu.scous males of Iwlophaeolis. Dyar 
established the synonymy of holophaealis and miicolor. I have 
examined the genitalia of the Barnes and McDunnough type. 
The species is extremely variable in color and venation and nor- 
mally the males and females are sharply contrasted, the nuiles being 
brownish ocherous and the females white. But this dimorphism 
is not constant. I have before me two males from Argentina 
(British Museum collection) that are almost pure white and not to 
be distinguished from the other wdiite species except l)y genitalia. 
We have also one pale brown female from French Guiana. The 
genital characters are constant. 
MOTHS OF THE GENUS RUPELA — HEINRICH 369 
Male. — Head, thorax, fore and hind wings, and dorsum of abdo- 
men pale brown or brownish ocherous (rarely Avhite), some speci- 
mens paler than others and some with hind wing slightly darker 
than fore wing, normally with fore and hind wings concolorous. 
In the brownish specimens the dark shading extends to the palpi 
and legs. Fore wing with veins 11 and 12 separate; 10 sometimes 
very short stalked with 8 and 9; 6 more or less approximate to 7 
at base (in other species 6 and 7 usually well separated and par- 
allel) ; 4 and 5 separate or connate. Hind wing with 4 and 5 sepa- 
rate, connate or shortly stalked. Anal tuft very pale ocherous or 
white. 
Alar expanse, 20-34 mm. 
Genitalia with gnathos beaklike; lateral arms rather long; cen- 
tral part with truncate lower margin; apical part rather short, not 
tapering; apex bluntly rounded; inner surface finely granulate. 
Uncus with basal part very little widened and not at all humped 
(rather flattened) ; apical part tapering; apex blunt. Harpe simple; 
apex bluntly rounded; basal projection of costa produced, the pro- 
jectioFiS of the two harpes joining at their apices to form a narrow 
transtilla. Anellus consisting of ventral plate and slightly rough- 
ened dorsal membrane; ventral plate elongate, its lateral margins 
notched. Aedeagus with basal half enlarged; without spines or 
serrations. 
Female. — Head, thorax, and wings sordid white to pure white 
(rarely pale brownish ocherous) ; the fore wing frequently with a 
yellowish tint. Venation as in the male. Anal tuft yellow; the 
underlying scales black-brown and wavy (in other species the under- 
lying scales are often dark brown or blackish but nearly always 
straight). 
Alar expanse, 25-42 mm. 
Genitalia with genital plate strongly sclerotized, forming a semi- 
circular band attached firmly to the rods of the eighth segment collar. 
Ductus sclerotized only at genital opening. 
Types. — In British Museum {tinctella and holophaealis) ; Berlin 
Museum (selleri) ; United States National Museum (unicolor). 
Type localities.— Venezu(iU {tinctella) ; Paramaribo, Surinam 
(zeileri) ; Abaco, Bahamas {holophaeaUs) ; Everglades, Fla. 
{unicolor) . 
Distrilut ion. —Visited States, Florida, Everglades (April), 
Miami, Coconut Grove, Dade City, Panacea (August), St. Peters- 
burg (April, July), Fort Meade (April), Grove (May); Mexico, 
Huasteca^ (Veracruz) ; Cuba, Matanzas (April) ; British Guiana, 
Georgetown (July) ; French Guiana, St. Laurent Maroni, St. Jean 
Maroni; Surinam, Geldersland (Surinam Eiver), Kartabo (No- 
370 PROCEEDII^GS OF THE NATIONAL MUSEUM vol.84 
vember) ; Trinidad (June) ; Brazil, Castro (Parana) ; Paraguay, 
El Gran Chaco (November) ; Argentina, Villa Ana (February, 
March). 
Remarks. — Ninety-two specimens (54 males and 38 females) ex- 
amined, from the collections of the United States National Museum, 
British Museum, and Cornell University. 
The species is easily identified by male and female genitalia. In 
all the species I have seen there is none except tinctella that has the 
genital plate of the female fused to rods of the collar. 
RUPELA NIVEA Walker 
Pl.\te 25. Figures 10-lOc 
Riipchi nivea Waikee, List of the specimens of lepidopteroiis insects in the 
collection of the British Museum, vol. 28, p. 524, 1863; male. 
This species, though quite distinct in genital characters from any- 
thing else in the genus, has long been listed as a synonym of alhlneiJa 
Cramer. I am indebted to Mr. Tams for examining genitalia of 
Walker's type and giving me the correct identification. 
Male. — Wings pure white. Fore wing with veins 4 and 5 closely 
approximate, connate or stalked; 11 and 12 closelj^ approximate or 
anastomosing. Hind wing with 4 and 5 closely approximate, con- 
nate or stalked. Anal tuft white. 
Alar expanse, 24-37 mm. 
Genitalia with gnathos beaklike, heavy, smooth, broad at base (in 
ventral aspect), tapering to bluntly pointed apex. Uncus broad at 
base and tapering to apex, stout; basal part extended backward into 
bulbous lobe ; from lateral view widest and slightly humped at middle 
(decidedly humped in specimen from Castro, pi. 25, fig. 10c). Harpe 
with apex tapering and rounded; basal process of costa {Clh) greatly 
extended, digitate ; sacculus produced into clasper (6^/) ; the latter 
a long, stout, straight spine. Anellus consisting of a rigid ventral 
plate and a dorsal membrane which attaches to the aedeagus ; u.pper 
(caudal) margin of plate deeply incised; membrane rugose, the 
wrinkles appreciably sclerotized. Aedeagus smooth; cylindrical; 
tapering slightly from beyond middle to apex. 
Type. — In British Museum. 
TyjJe locality. — Para (Santarem), Brazil. 
Distribution. — Panama, Porto Bello (April, May) ; Brazil, Castro 
(Parana) ; Argentina, Gran Chaco (October), Villa#Ana (January, 
February), Goya. 
liejiiarhs. — Twelve specimens examined. These are all males and 
are from the United States National Museum and British Museum 
collections. The Castro male (from the British ISIuseum collection) 
is somewhat abnormal. Tlie uncus is more appreciably humped than 
MOTHS OF THE GEKUS RUPELA HEII^RICH 371 
in any of the other specimens and the clasper had a short secondary 
spine branching from its base. These differences are probably but 
individual abnormalities and, in my opinion, do not justify any 
separate name (varietal or otherwise) for the Castro specimen. 
The female is unknown. 
RUPELA VEXATIVA, new species 
Plate 25, Figubes 11-llc 
Male. — Wings pure white. Fore wing witli veins 4 and 5 connate ; 
11 anastomosing with 12. Hind wing with 4 and 5 very shortly 
stalked. Anal tuft white. 
Alar expanse, 27 nun. 
Genitalia wdth gnathos beaklike, triangular, cleft from apex to 
middle. Uncus broad at base, tapering sharply to pointed and 
slightly hooked apex; basal part extended backward as a bulbous 
lobe (cowllike) ; dorsal surface of lobe with a few weak, scattered 
scobinations. Harpe with cucullus somewhat broadened toward 
apex; apex rounded; basal process of costa long, digitate; saccuius 
produced into clasper, the latter, a strong, stout spine, curved up- 
Avard to costa. Anellus consisting of a rigid ventral plate and dorsal 
membrane; plate somewhat constricted at middle, lateral margins 
deeply incurved; membrane but slightly rugose, weakly sclerotized. 
Aedeagus smooth, bulging laterally just beyond middle, thence tap- 
ering rather abruptly toward apex ; apex spatulate. 
Type. — In Britisli Museum. 
Type locality. — Quirigua, Guatemala. 
Remarks. — Described from one male collected bv Wm. Schaus 
(April). 
This species is easily recognized by its peculiar aedeagus, its cleft, 
triangular gnathos, and its strong, curved, hooklike clasper. 
Female unknown. 
RUPELA CORNIGERA, new species 
Plate 25, Fi(iui:Es Vl-Vlh 
Male. — Wings white. The wings are not so bright as in most of 
the other species, but their dullness may be due to the condition 
of the specimens. Under side of fore wing faintly dark shaded 
in costal area; veins 11 and 12 separate; 4 and 5 connate or shortly 
stalked. Hind wing with 4 and 5 connate or shortly stalked. Anal 
tuft w^iite. 
Alar expanse, 27-30 mm. 
Genitalia with gnathos beaklike, tapering abruptly to middle, and 
gradually from middle to bluntly rounded apex. Uncus stout, basal 
part very broad and with central rib; apical two-thirds digitate; 
131080—37 2 
372 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 84 
apex bluntly rounded. Harpe approximately rectangular; basal 
process of costa somewhat produced but not strongly sclerotized, 
fusing into membranous transtilla; saceulus folded upward toward 
apex, not otherwise produced. Anellus consisting of ventral plate 
and a more strongly sclerotized U-shaped dorsal band; dorsal margin 
of ventral plate slightly concave ; dorsal band bearing a pair of very 
long, stout spines. Aedeagus bent beyond middle; apical area ven- 
trally compressed and slightly concave; the hind margin of this 
concave area armed with short thornlike scobinations ; apex truncate. 
Ti/pe.—U.S.^M. no. 51861. 
Paratypes. — In British Museum. 
Type locality. — Castro, Parana, Brazil. 
/?e??i«.r7^.— Described from male type (Schaus, collector) and two 
male paratypes (E. D. Jones, collector) from the type locality. In 
addition to the above I have before me two small males belonging 
to the British Museum and collected at Obydos, Brazil, by E. E. 
Austin, February 2, 1896. They are smaller (18.5-20 mm) than 
the type series, and their genitalia are about half the size of those 
of typical carnigei^a. Otherwise there appears to be no difference. 
Possibly they represent a race. On the other hand, they may be 
merely stunted, aberrant specimens. Therefore I am not including 
ihem among the paratypes. 
Female unknown. 
RUPELA IMITATIVA, new species 
Plate 26, Figures 13-13c 
Male. — Wings shining white. Fore wing with veins 11 and 12 
approximate; -1 and 5 approximate. Hind wing Avith 4 and 5 con- 
nate or very shortly stalked. Anal tuft white. 
Alar expanse, 36-38 mm. 
Genitalia with gnathos beaklike; lateral arms developed; lower 
margin slightly convex; apical part stout, tapering to bluntly 
pointed apex; apex upturned; inner surface with a weak, central, 
longitudinal ridge. Uncus with basal part ovoid, produced back- 
■svard (cowllike), finely and densely scobinate; apical part stout 
and but slightly tapered; apex bluntly pointed. Harpe with saceulus 
produced at apex into a strongly sclerotized fold; basal projection 
of costa produced, prominent; cucullus not narrowed, apex bluntly 
rounded. Anellus consisting of ventral plate and a slightly rough- 
ened dorsal membrane, ventral plate with upper margin concave, 
lateral margins widely and deeply concave. Aedeagus with basal 
part somewhat enlarged; apical half narrowing to blimt apex; 
apex with lateral flange. 
Type. — In British Museum. 
Paratype.—V.^.'^M. no. 51862. 
MOTHS OF THE GENUS RUPELA — HEINRICH 373 
Type locality.— Castro, Parana, Brazil (E. D. Jones). 
Remarks. — Described from male type and one male paratype from 
the type locality. May be recognized at once by the flanged aedeagus. 
Female unknown. 
RUPELA SEJUNCTA. new species 
Plate 26, Figures 14-1 4c; Plate 32, Figlre 39 
Male. — ^Wings shining white. Fore wing with veins 11 and 12 
anastomosing; 4 and 5 connate. Hind wing with 4 and 5 connate 
or very shortly stalked. Anal tuft white. 
Alar expanse. 28-33 mm. 
Genitalia with gnathos beaklil^e; lateral arms developed; lower 
margin broadly rounded; apical part abruptly tapering to bluntly 
pointed apex; apex upturned and notched; outer (under) surface 
slightly roughened. Uncus with basal part ovoid, produced back- 
ward (cowllike), covered with short scobinations ; apical part some- 
what short in proportion to basal, tapering to pointed apex. Harpe 
with narrowed cucullus; apex bluntly rounded; sacculus produced 
into a very short spine at apex; basal projection of costa greatly 
produced. Anellus consisting of ventral plate and unsclerotized 
dorsal membrane; ventral plate with upper margin deeply angidate- 
emarginate, lateral margins widely and rather deeply concave. 
Aedeagus with apical half and extreme base greatly narrowed; 
apical half rodlike, curved at apex; manica prominent, but weakly 
sclerotized. 
Female. — ^^Ving color and venation as in the male. Anal tuft 
white. 
Alar expanse, 25-30 mm. 
Genitalia with genital plate large, completely surrounding genital 
opening, strongly sclerotized, its lateral areas granulate; lower mar- 
gins of genital opening deeply rugose, black-brown and very strongly 
sclerotized; at caudal margin a short, hollow, outwardly projecting 
nipple. 
Type and faratypes. — U.S.N.M. no. 51863. Paratypes also in 
British Museum. 
Type locality. — Harris County, Tex. (May). 
Remarhs. — Described from male type, six male and six female 
paratypes; the paratypes distributed as follows: Texas, Han-is 
County, one male and one female; Georgia, one male and one female; 
Florida ("Allen Kiver to Deep Lake", April 12, 1912), one male, 
Everglades, one female; Alabama, Selma (E. A. Schwarz, Septem- 
ber 1880), one male; "Staten Island" ("26-VI-Ol"), one female; 
also two males and two females from the Zeller collection (1880) 
in the British Museum, without locality label but presumably from 
Texas. 
374 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 84 
The species as far as I know is limited to the United States. 
Hitherto specimens have been identified as albinella Cramer. It can 
be identified at once by its peculiar aedeagus and female genital 
plate. 
RUPELA SCITULA, new species 
I'LATB 26, Figures 15-15c 
Male. — Wings shining Avliite. Fore wing with veins 11 and 12 
separate; 4 and 5 approximate or connate. Hind wing with 4 and 
5 stalked. Anal tuft white. 
Alar expanse, 21-29 mm. 
Genitalia with gnathos beaklike ; lateral arms well developed ; basal 
projecting part truncate; apical part tapering to pointed apex; in- 
ner surface sparsely and very finely scobinate. Uncus with basal 
part bilobed but rather small as compared with other species, the 
lobes bearing several short spines on dorsum; apical part long, 
tapering to pointed apex. Harpe with basal process of costa slightly 
produced; apex bluntly rounded; sacculiis produced into a narrow, 
elongate fold at apex, otherwise simple. Anellus consisting of 
ventral plate and a partially sclerotized dorsal band, the latter 
bearing one pair of very long, stout spines and a pair of much 
thinner and shorter spines; lateral margins of ventral plate nar- 
rowly but deeply excurvate. Aedeagus with apex deeply l)ut nar- 
rowly excavate on under side and bearing several short, stout^, 
ventral spines. Tegumen vpith sharp, projecting spur {Tgsp) from 
each inner lateral margin. 
Tijpe and paratype. — U.S.N.M. no. 51864. Paratypes also in Cor- 
nell University collection and British Museum. 
Type locality. — Tucuman, Argentina, 
Remarlcs. — Described from male type and seven male paratjqies, 
the paratypes distributed as follows: On boat from Mexico (Quar- 
antine no. Phila. 27764), one; Brazil, Obydos (E. E. Austin, Febru- 
ary 2, 1896), one, Rio Janunda ("11-4-47"), two; British Guiana, 
Mackenzie. Demerara (W. T. M. Forbes, June 23-24, 1927), tlu-ee. 
^fay be identified at once by the shape of the uncus, the spining 
of the dorsal part of the anellus, and the spurs on the tegumen. 
Female unknown. 
RUPELA ADUNCA, new species 
Plate 27, Figures 16-16c 
Male. — Wings white. Fore wing with veins 11 and 12 separate; 
4 and 5 shortly stalked. Hind wing with 4 and 5 very shortly 
stalked. Anal tuft a'cUow. 
% 
MOTHS OF THE GENUS RUPELA HEINKICH 375 
Alar expanse, 39 mm. 
Genitalia with gnatlios bandlike, the central portion widely ex- 
panded (apronlike). Uncus with base enlarged, bulbous, covered 
with small, papillate protuberances and with a narrow dorsal groove ; 
apical part laterally compressed (knifelil^e), slightly humped at 
middle; apex pointed. Harpe with basal projection from costa 
greatly produced and fusing with membranous, minutely scobinate 
transtilla; sacculus produced at apex into an upcurved sclerotized 
lip from Avhich a sclerotized ridge extends upward to the costa; at 
center of ridge a slight projection. Anellus consisting of ventral 
plate and dorsal membrane; ventral plate with upper margin 
broadly and deeply angulate and lateral margins deeply excavate. 
Aedeagus heavy; basal part swollen; apex produced into a stout, 
blunt hook. 
Type.—V.S.'NM. no. 51865. 
T]/j}e locality. — Bolivia. 
Bemarks.—DQ^QvW,^^ from one male from the National Museum 
collection labeled as follows: "17-46-55 S. Lat., 63-5-34 Long." 
A large species easily identified by its peculiar aedeagus and uncus. 
Female unknown. 
RUPELA LUMARIA, new species 
Plate 27, Figures 17-17c 
Male. — Wings white. Fore wing with veins 11 and 12 separate; 
4 and 5 closely approximate. Hind wing with 4 and 5 closely 
approximate. Anal tuft yellow. 
Alar expanse, 30 mm. 
Genitalia with gnathos bandlike, slightly expanded at middle. 
Uncus with basal part a broadly triangular, hoodlike, rugose pro- 
jection with deep central, dorsal groove; apical part abruptly taper- 
ing to pointed apex. Harpe with basal costal projection strongly 
produced and fusing with finely scobinate, membranous transtilla; 
sacculus produced into an elongately triangular, coarsely scobinate 
projection with apex sharply pointed; apex of harpe bluntly 
rounded. Anellus consisting of ventral plate and membranous dorsal 
part, the latter faintly rugose. Aedeagus stout, cylindrical; apex 
expanded and produced into three heavy claws. 
Type.—V.^.^M. no. 51866. 
Type locality.— CaviWo, Costa Eica (W. Schaus, March). 
/?ewarA,'.s.— Described from single male. Can be identified by 
basal modification of uncus, the 3-clawed aedeagus, and the heavy, 
triangular, scobinate projection of the sacculus. 
Female unknown. 
376 PROCEEDINGS OF THE NATIONAL I^IUSEUM vol.8* 
RUPELA HORniDULA, new species 
Pi.ATE 27, Figures 18-18/ ; Plate 33, FiomiE 47 
Male. — Wings white. Fore wing with veins 11 and 12 separate; 
4 and 5 connate or closely approximate. Hind wings with 4 and 5 
connate or shortly stalked. Anal tuft yellow. 
Alar expanse, 22-32 mm. 
Genitalia with gnathos bandlike, simple. Uncus with basal part 
bifid, produced as two subtriangular, laterally flattened, jagged 
lobes. Apical part smooth except for a slight dorsal keel near apex ; 
apex hooked. Harpe broad ; apex bluntly pointed ; basal projection 
of costa considerably produced and fusing into membranous trans- 
tilla; sacculus produced at apex" into a short, broad, blunt, up- 
curved spur; from this a sclerotized ridge extends to the basal pro- 
jection of the costa. Anellus consisting of ventral plate and a pair 
of strongly sclerotized, narrow, sinuous, irregularly serrate dorsal 
plates; ventral plate with upper (caudal) margin deeply angulate 
and lateral margins excavate. Aedeagus with from one to three 
ventral thornlike teeth toward apex ; penis bearing a small, flattened, 
serrate coniutus. 
Type and paratypes. — U.S.N.M. no. 51867. Paratypes also in 
collections of Cornell University, British Museum and Harold E. 
Box. 
Type locality. — Campo Bello, Rio de Janeiro, Brazil (Zikan, 
collector). 
Food plant. — Andropogon hicornis. This food-plant record is 
from specimens submitted by Dr. H. E. Box, San Jose, British 
Guiana, April 1936 (Mj^ers no. 5328). 
Remarks. — Described from male type and 25 male paratypes, the 
paratypes distributed as follows: Brazil, Campo Bello (Rio de 
Janeiro), two. Organ Mountains (near Tijuca, Rio de Janeiro), one, 
Ponte Nova (Rio Xingu), three; Surinam, Zanderij (Boven, Para 
District, April), one; French Guiana, St. Jean Maroni, five; 
British Guiana, San Jose (Pupunimi District, April), two, George- 
town, one, Kartabo (Bartica District, October, November), four, Mac- 
kenzie (June), two, Rio Demerara, one; Trinidad, (Dyar collection, 
B. M. no. 1923-361), two, (Saunders collection, B. M. no. 94-68), one. 
This species is easily recognized by the structure of its anellus and 
uncus. There is considerable variation in the size and spining of 
the basal lobes of the uncus in different specimens, but between the 
extreme forms (shown in pi. 27, figs. 18fZ and 18e) there is every 
possible intergrade so that no distinct varieties or races can be 
established. 
Female unknown. 
% 
MOTHS OF THE GENUS RUPELA HEINRICH 377 
RUPELA SPINIFERA, new species 
Plate 28, Figukes 19-19c 
Male. — Wings shining white. Fore wing with veins 11 and 12 
separate: 4 and 5 connate. Hind wing with 4 and 5 connate or 
shortly stallved. Anal tuft yellow. 
Alar expanse, 39-45 mm. 
Genitalia with gnathos bandlike, simple. Uncus with basal part 
produced backward into a broad, rounded, concave, and bifurcate 
plate armed along its outer margin with long, heavy spines; apical 
part long, its base clothed with long, slender hairs on dorsum, its 
apex sharply hooked. Harpe broad; cucullus abruptly narrowed; 
apex bluntly pointed ; basal projection of costa long, digitate, fusing 
into membranous transtilla; sacculus produced at apex into a wide 
flange which is extended in a sclerotized ridge to basal projection 
of costa. Anellus consisting of ventral plate and dorsal membrane; 
upper margin of ventral plate angulate, lateral margins broadly 
excavate. Aedeagus long, tapering from slightly enlarged base to 
produced apex. 
Type.—\J.^.'^M. no. 51868. Paratype in British Museum. 
Type locality. — Castro, Parana, Brazil. 
Remarks. — Described from male type (W. Schaus, collector) and 
one male paratj'pe (E. D. Jones, collector) without dates and both 
from the type locality. 
A large species distinguished by its long slender aedeagus and 
heavily spined uncus. 
Female unknown. 
RUPELA MONSTRATA, new species 
PLATE 2S, Figures 20-20c 
i¥aZe.— Wings white. Fore wing with veins 11 and 12 separate; 
4 and 5 closely approximate. Hind wing with 4 and 5 closely approx- 
imate. Anal tuft yellow. 
Alar expanse, 46 mm. 
Genitalia with gnathos bandlike, simple. Uncus with base only 
slightly widened, produced backward into a stout, forked process 
with heavy, ribbed stem and thick, stubby prongs, the latter bearing 
one or two spines; apical part greatly extended, digitate; apex 
abruptly tapering and slightly hooked. Harpe broad; cucullus 
abruptly narrowed; apex bluntly pointed; basal projection of costa 
normally produced but not strongly sclerotized (transtilla mem- 
branous and not well defined) ; sacculus produced at apex into a 
shallow flange whose upper extremity is slightly notched. Anellus 
consisting of ventral plate and dorsal membrane; the upper and 
378 PROCEEDINGS OF THE NATIOX.VL MUSEUM vol.84 
lateral margins of the ventral plate are so deeply angulate that the 
upper half of the plate appears as two diver<Tent bands. Aedeagus 
long; basal half slightly enlarged; apical half not appreciably taper- 
ing ; apex truncate, with one or two short teeth at ventral extremity. 
Type.—U.S.l^M. no. 51869. 
Type Jocallty. — Castro, Parana, lirazil (W. Schaus). 
Remarks. — Described from one male. The largest of the male 
llii))elas, easily identified by its anelliis and greatly elongated digitate 
uncus. 
Female unknown. 
UNASSOCIATED FEMALES 
RUPELA ANTONIA, new species 
Plate 30, Figube 31 
Wi]igs white. Fore wing with veins 11 and 12 separate; 4 and 
5 connate. Hind wing with 4 and 5 stalked. Anal tuft yellow. 
Alar expanse, 38-43 mm. 
Genitalia as in Jeucatea except that the lower margin of the genital 
opening is concave rather than sinuate and the sclerotization of the 
ductus bursae extends farther back from the genital opening. 
Type and y^am/y/^e.— U.S.N.M. no. 51892. 
Type locality. — 'Sixola River, Costa Rica (April, September). 
Described from two females, both from the type locality. 
RUPELA BENDIS, new species 
Plate 29, Figure 21 
Wings Avhite. Fore ^Ying with 11 and 12 separate ; 4 and 5 connate 
or shortly stalked. Hind wing with 4 and 5 connate or shortly 
stalked. Anal tuft yellow. 
Alar expanse, 28-33 mm. 
Genitalia similar to those of alhinella but without defined genital 
plate; area about genital opening very weakly sclerotized. Lower 
margin of genital opening concave and with a small notch in center. 
Type and paratype. — U.S.N.M. no. 51893. Paratype also in British 
Museum. 
Type locality. — Aroa. Venezuela. 
Remarhs. — Described from three fenuiles, the paratypes distributed 
as follows: Venezuela, Aroa, one; Brazil, Parapanema (Sao Paulo), 
one. 
There are also two rather doubtful specimens in the British Mu- 
seum from Brazil (Parana de Buyassu and Rio Cararauca). In one 
of these the central notch in the lower margin of the genital opening 
is obsolete and in the other nearly so. 
MOTHS OF THE GENUS RUPELA HEINRICH 379 
RUPELA CANENS, new species 
Tlate 21), Figure 23 
Wings white. Fore wing with veins 11 and 12 anastomosing; 4 and 
6 connate. Hind wing with 4 and 5 shortly stalked. Anal tuft yellow. 
Alar expanse, 26-38 mm. 
Genitalia similar to those of hendis, but area about genital opening- 
more appreciably sclerotized; a genital plate faintly indicated. 
Lower margin of genital opening deeply conca^'e (somewhat angu- 
late). Bursa copulatrix very small, not reaching length of rods of 
eighth segment collar. Ductus bursae sclerotized for some distance 
from genital opening. 
r//;>e.— U.S.N.M. no. 51894. Paratype in British Museum. 
Type locality. — Sao Paulo de Olivenca, Brazil. 
Remarks. — Described from two females, the paratype from Parin- 
tins, Brazil. 
In some characters the genitalia are more similar to those of 
alhinella than to those of hendis. However, they seem to indicate 
a species distinct from either. 
RUPELA DRUSILLA, new species 
Plate 29, Figure 24 
Wings white. Fore wing with veins 11 and 12 separate; 4 and 5 
connate. Hind wing with 4 and 5 stalked. Anal tuft white. 
Alar expanse, 28 mm. 
Genitalia with genital opening wide. Genital plate reduced to 
area back of genital opening (not completely surrounding the open- 
ing), triangular. Lower margin of genital opening angulate. Duc- 
tus bursae laterally broadened and well sclerotized toward genital 
opening. The genitalia resemble most those of segrega, from which 
they are readily separated by the shape of the lower margin of the 
genital opening, 
Type.—V.S.'^.U. no. 51895, 
Type locality. — Castro, Parana, Brazil. 
Remarks. — Described from one female. The fore wings of this 
specimen have a faint creamy tint, which may or may not be the 
normal color. I suspect that a series would show most of the speci- 
mens pure Vvdiite. Possibly the female of saetigera or pallidida. 
RUPELA EDUSA, new species 
Plate 30, Figure 29 
Wings white. Fore wing with 11 and 12 separate or approximate; 
4 and 5 separate or connate. Hind wing with 4 and 5 separate or 
connate. Anal tuft wliite. 
380 PROCEEDINGS OF THE NATIONAL MUSEUM vol.84 
Alar expanse, 21-26 mm. 
Genitalia with area behind the genital opening markedly sclerotized 
and pigmented, smooth just behind opening and rugose and finely 
granulate beyond (in the direction of ovipositor). Lower margin of 
genital opening concave with lateral ends somewhat straightened ; the 
ductus at genital opening narrowly sclerotized. Ductus seminalis 
from ductus bursae and forming a loop with it just before genital 
opening. 
Type and paratypes.—V.S.^M. no, 5180G. Paratype also in 
British Museum. 
Type locality. — Castro, Parana, Brazil. 
Remarks. — Described from seven females, the paratypes all from 
the type locality. 
The species is chiefly distinguished by the peculiar shape and junc- 
ture of the ductus bursae and ductus seminalis. This may be the 
female of laheosa. 
RUPELA FAUSTINA, new species 
Plate 29, Figuke 22 
Wings white. Fore wing with veins 11 and 12 anastomosing; 4 
and 5 connate or shortly stalked. Hind wing with 4 and 5 connate 
or shortly stalked. Anal tuft yellow. 
Alar expanse, 21-25 mm. 
Genitalia with a sinjooth, sclerotized, roundly oval plate just behind 
genital opening. Lower margin of genital opening slightly angulate. 
Just within lip of genital opening a pair of short, dark, hooklike 
processes. Ductus weakly sclerotized toward genital opening. 
Type and paratype. — U.S.N.M. no. 51897. Paratype also in British 
Museum. 
Type locality. — Cabima, Panama (Busck, May). 
Remark's. — Described from three females from the type locality. 
From the fore-wing venation, size, and distribution I am inclined 
to believe that this is the female of liberta. 
RUPELA GAIA, new species 
Plate 30, Figure 28 
"Wings white. Fore wing Avith veins 11 and 12 separate; 4 and 5 
closely approximate or connate. Hind wing with 4 and 5 closely 
approximate or connate. Anal tuft yellow. 
Alar expanse, 39-48 mm. 
Genitalia Avith area to the sides and behind the genital opening 
slightly rugose and weakly sclerotized. Lower margin of genital 
opening angulate. the angle bluntly pointed. Within the genital 
opening a pair of dark, small, oblong, sclerotized disks. 
MOTHS OF THE GENUS RUPELA HEINRICH 381 
Tyfe and -paratypes. — U.S.N.M. no. 51898. Paratypes also in 
British Museum. 
Type locality. — Castro, Parana, Brazil. 
Remarks. — Described from six females, the paratypes distributed 
as follows: Brazil, Castro, three; Argentina, one; Paraguay, 
Villarrica (November), one. 
RUPELA HERIE, new species 
Plate 30, Figitee 27 
White wings. Fore wing with veins 11 and 12 sejDarate; 4 and 5 
connate. Hind wing with 4 and 5 connate. Anal tuft j^ellow. 
Alar expanse, 30-33 mm. 
Genitalia with pigmented (yellow) and sclerotized genital plate; 
the plate rugose, especially tovrard margins and with caudal part 
acutely angulate. Genital opening semicircular, the lateral edges of 
its lower margin fusing into the plate. 
ry/;^?.— U.S.N.M. no. 51899. Paratype in Cornell University 
collection. 
Type locality. — Georgetown, British Guiana (April). 
Remarks. — Described from two females, the paratype from Zan- 
derij (Para District), Surinam (April). 
RUPELA JANA, new species 
Plate 31, Figuke 38 
Wings white. Fore wing with veins 11 and 12 closely approxi- 
mate (rarely anastomosing) ; 4 and 5 connate or shortly stalked. 
Hind wing with 4 and 5 connate or shortly stalked. Anal tuft 
yellow. 
xVlar expanse, 29-47 mm. 
Genitalia with large sclerotized plate completely surrounding gen- 
ital opening ; its lateral areas partially detached from central portion 
of the plate; near its caudal margin a strongly sclerotized, brown, 
external pocket. Ductus unsclerotized at genital opening and the 
lower margin of genital opening unpigmented. Genital opening 
nearly circular. 
Type and paratypes. — U.S.N.M. no. 51900. Paratypes also in Brit- 
ish Museum and Cornell University collections. 
Type locality. — Chaco, Argentina. 
Remarks. — Described from 28 females, the paratypes distributed as 
follows: Panama, Porto Bello (May), four; Sueinam (no other local- 
ity, Zeller collection of British Museum), one; British Guiana, Ber- 
bice, one; Brazil. Obydos (Para), one, Rio Madeira, three, Rio 
Cuminae, one. Parti, two. Breves (January), one: Peru, Iquitos, one; 
382 PROCEEDINGS OF THE NATIONAL MUSEUM vol.84 
Argentina. Villa Ana (January. February, March), eight, Goya, one, 
and two specimens without definite locality; Paraguay, Gran Chaco 
(March), one. 
In addition to the type series there is one specimen in the British 
Museum from Sao Paulo, Brazil, that belongs here but is not in- 
cluded among the paratypes. It is abnormal, in that the sclerotized 
pocket is much narrower than in other specimens and the lateral 
areas of the genital plate arc completely fused with the central area. 
The entire plate seems to have been pinched. I believe that these 
diiicrences indicate nothing more than an individual and freak 
development. 
RUPELA CANDACE, new species 
Plate 29, Figures 25, 25f/ 
Wings Avhite. Fore wing with veins 11 and 12 separate; 4 and 
5 closely approximate or connate. Hind wnng with 4 and 5 connate. 
Anal tuft 3^ellow. 
Alar expanse, 38-4:4 mm. 
Genitalia with large sclerotized plate surrounding genital open- 
ing; at its caudal end a strongly sclerotized, internal pocket (see 
projection sketch of pi. 29, fig. 25). Ductus sclerotized at genital 
opening. Margin of genital opening strongly sclerotized and dark 
brown. Genital opening oval. 
Type. — In British Museum. 
Parafype.—\J.^.'^M. no. 61901. 
Type locality. — Castro, Parana, Brazil. 
Described from two females from the type locality. 
RUPELA LARA, new species 
Plate 32, Figuue 40 
Wings white. Fore wing with veins 11 and 12 approximate; 4 and 
5 approximate or connate. Hind wing with 4 and 5 approximate, 
connate or stalked. Anal tuft yellow. 
Alar expanse, 20-36 mm. 
Genitalia with genital plate developed as a stout, blunt, thorn- 
like process just in front of genital opening. On each side of genital 
opening a large, finely scobinate, sclerotized area. Membrane back 
of genital opening unpigmented and unsclerotized except that in 
the area near eighth segment collar there is a small, brown, sclerotized, 
external pocket. Genital opening behind apex of genital plate. Duc- 
tus not appreciably sclerotized. 
Type and pamtype,<^.—V.S.'N.M. no. 51902. 
Type locality. — Cabima, Panama (May). 
MOTHS OF THE GENUS RUPELA HEINRICH 383 
Remarks. — Described from three females, the paratypes distributed 
as follows: Panama, Rio Trinidad (June), one; Costa Rica, Gua- 
piles (March), one. 
RUPELA MAENAS, new species 
Plate 31, Figures 36, 37 
Wings white. Fore wing with veins 11 and 12 approximate; 4 
and 5 closely approximate or connate. Hind wing with 4 and 5 
connate or shortly stalked. Anal tuft yellow. 
Alar expanse, 22-31 mm. 
Genitalia with ductus bursae strongly sclerotized, brown and ex- 
panded into a laterally flattened bulb near genital opening. Genital 
plate developed as a narrow hoodlike piece in front and a broad, 
hooked, truncate flange behind genital opening (see pi. 31, fig. 36), 
the flange protruding from the area between the ventrolateral mar- 
gins of the eighth segment collar. Genital opening nearly circular. 
Type and paratypes. — ^U.S.N.M, no. 51903. Paratypes also in 
British Museum and Cornell University collections. 
Type locality. — Ponte Nova (Rio Xingu), Brazil. 
Remarks. — Described from eight females, the paratypes distrib- 
uted as follows: French Guiana, St. Laurent Maroni (March), one; 
British Guiana, Kartabo (October), one, Mackenzie (June), two; 
Surinam, Zanderij (April), two; Brazil, Castro (Parana), one. 
Easily identified by the bulbous, sclerotized ductus. I suspect it 
may be the female of horridida. 
RUPELA NEREIS, new species 
Plate 31, Figures 33, 34 
Wings white. Fore wing with veins 11 and 12 separate; 4 and 
5 connate. Hind wing with 4 and 5 connate. Anal tuft yellow. 
Alar expanse, 40 mm. 
Genitalia with ductus bursae brown, sclerotized and tubular toward 
genital opening. Genital plate consisting of a pair of narrow lateral 
flaps extending backward from margin of genital opening and fusing 
with a flanged and hooked process, which protrudes between the 
ventrolateral margins of the eighth segment collar; strongly sclero- 
tized, brown ; apex of protruding flange abruptly tapering to a blunt 
point. Ventral area between collar and ovipositor strongly sclero- 
tized, developed as a pair of elongate, shallow, somewhat rugose 
depressions. Genital opening semicircular. 
Type. — In British Museum. 
Type locality. — Castro, Parana, Brazil. 
Remarks. — Described from one female. 
384 PROCEEDINGS OF THE NATIONAL MUSEUM vor.. 84 
Apparently nearest to maenas but not to be confused with anything 
in the genus. 
RUPELA ORBONA, new species 
Plate 31, Figure 35 
Wings white. Fore wing with veins 11 and 12 separate; 4 and 
5 connate. Hind wing with 4 and 5 shortly stalked. Anal tuft 
3'ellow. 
Alar expanse, 30 mm. 
Genitalia with genital plate triangular, brown, sclerotized, ap- 
pressed to and no wider than ductus bursae; in ductus (at end of 
genital plate) a pale yellow, round, rugose thickening of the tube. 
Area behind genital opening sclerotized and pigmented, a smooth 
ungulate plate, at its apex a dark brown, heavily sclerotized, elon- 
gately triangular, internal pocket. Genital opening slitlike. 
TyiJe. — In Cornell University collection. 
Type locality. — Mackenzie, British Guiana (June). 
Described from one female. 
RUPELA PROCULA, new species 
Plate 32, Figure 41 
Wings white. Fore wing with veins 11 and 12 separate or approxi- 
mate; 4 and 5 approximate. Hind wing with 4 and 5 approximate 
or connate. Anal tuft yellow. 
Alar expanse, 50-51 mm. 
Genitalia with genital plate surrounding the genital opening, ir- 
regular, strongly sclerotized and pigmented; from the area behind 
opening a deep, strongly sclerotized internal pocket; beyond this, 
the area between the ventrolateral margins of the eighth segment 
collar is rugose, partially pigmented, and sclerotized. Ductus 
strong!}^ sclerotized, dark, and laterally expanded towards genital 
opening. Genital opening large, irregular. 
Type and paratype. — U.S.N.M. no. 51904. 
Type locality. — Sta. Catherina, Brazil. 
Described from two females, the paratype from Jepelacio, Peru. 
EXPLANATION OF PLATES 
The drawings for the phites accompanying this paper were made 
under the author's supervision by Mrs. Eleanor A. Carlin, of the 
Bureau of Entomology and Plant Quarantine. The female genitalia 
were drawn to smaller scale than those of the males. 
EXPLANATION OF SYMBOLS APPLIED TO GENITALIA 
^Male 
CZ=Clasper of harpe. 
CZ/i = Basal projection from costa of harpe. 
cji = Cornutus (cornuti). 
Cu = Cucullus of harpe. 
Gn=Gnathos. 
ZoGn= Lateral arm of gnathos. 
Ma=Manica uniting anellus and aedeagus. 
(Sc = Sacculus of harpe. 
T^sp = Spur from ventrolateral margin of tegumen. 
Tr = Transtilla. 
[7= Uncus. 
Female 
jBc = Bursa copulatrix. 
CZr= Collar of eighth abdominal segment. 
Z)5= Ductus bursae. 
Ds = Ductus seminalis. 
Go = Genital opening. 
Gp = Genital plate. 
iwi = Lower margin of genital opening. 
p = Sclerotized pocket in area caudad of genital opening. 
Plate 22 
1-1(7. RupeJa leucatca (Zeller) : ], Ventral view of male genitalia with 
aedeagus and one harpe omitted; la, auellus, dorsal view^ lb, 
aedeagus ; Ic, lateral view of tegumen, uncus, and gnathos ; Id, dorsal 
view of uncus and gnathos. 
2-2f/. Riipela aJhinella (Cramer) : 2, Ventral view of male genitalia with 
aedeagus and cue harpe omitted ; 2a, anellus, dorsal view ; 2b, ventral 
and lateral views of aedeagus (the lateral view showdng anellus at- 
tached) ; 2c, lateral view of tegumen, uncus, and gnathos; 2d, dorsal 
view of uncus and gnathos. 
3-3(Z. Rupela labcosa, new species: 3, Ventral view of male genitalia with 
aedeagus and one harpe omitted ; Sa, anellus, dorsal view ; of*, aedea- 
gus ; 3r, lateral view of tegumen, uncus, and gnathos ; 3d, dorsolateral 
view of uncus and gnathos. 
385 
386 PROCEEDINGS OF THE NATIONAL MUSEUM voi.. 84 
Plate 23 
4— 4c. Riipcia libc)-ta, new species : 4, Ventral view of male genitalia with 
aedeagus and one harpe omitted ; 4a, dorsal view of anellns showing 
dorsal part and spines turned outward ; 4?j, dorsal view of anellus 
and spines in normal position ; 4c, aedeagus. 
5-5(?. Ritpcla palUdula, new species : 5, Ventral view of male genitalia with 
aedeagus and one harpe omitted ; 5a, anellus, dorsal view ; 5^, 
aedeagus ; 5c, lateral view of uncus and gnathos ; mI, dorsal view 
of uncus and gnathos. 
6-6(?. Rupela scyreyu, new species : 6, Ventral view of male genitalia with 
aedeagus and one harpe omitted ; 6a, anellus, dorsal view ; G?>, 
aedeagus; 6c, lateral view of uncus and gnathos; 6(/, dorsolateral 
view of uncus and gnathos. 
Plate 24 
1-ld. Rupela yihhera, new species: 7, Ventral view of male genitalia with 
aedeagus and one harpe omitted ; la, anellus, dorsal view ; 7b, 
aedeagus with sclerotized manica ; 7c, lateral view of uncus and 
gnathos ; 7(7, dorsal view of imcus and gnathos. 
8-8c. Rupela saetigera, new species: 8, Ventral view of male genitalia with 
aedeagus and one harpe omitted ; 8a, anellus, dorsal view ; 8&, 
aedeagus with sclerotized manica ; 8c, dorsal view of uncus and 
gnathos. 
9-9c. Rupela tinctella (Walker) : 9, Ventral view of male genitalia with 
aedeagus and one harpe omitted; 9a, anellus, dorsal view; 9&, 
aedeagus ; 9c, lateral view of uncus and gnathos. 
Plate 25 
10-lOc. Rupela n'n-ea Walker: 10, Ventral view of male genitalia with aedeagus 
and one harpe omitted ; 10a, anellus, dorsal view ; lOh, aedeagus ; 
10c, lateral view of uncus and gnathos from abnormal specimen from 
Castro, Brazil. 
11-llc. Rupela vcxativa, new species: 11, Ventral view of male genitalia with 
aedeagus and one harpe omitted; 11a, anellus, dorsal view: lib, 
aedeagus ; lie, dorsal view of uncus and gnathos. 
12-12&. Rupela cornigera, new species: 12, Ventral view of male genitalia with 
aedeagus and one harpe omitted; 12a, anellus. dorsal view; Vlb, 
aedeagus. 
PlATE 26 
13-13c. Rupela imitativa, new species: 13, Ventral view of male genitalia with 
aedeagus and one harpe omitted ; 13a, anellus, dorsal view ; I'-lb, 
aedeagus ; 13c, dorsal view of imcus and gnathos. 
14-I4c. Rupela sejuneta, new species : 14, Ventral view of male genitalia with 
aedeagus and one harpe omitted ; 14a, anellus. dorsal view ; lAb, 
aedeagus with attached manica ; 14c, lateral view of uncus and 
gnathos. 
IS-loc. Rupela scitula, new species: 15, Ventral view of male genitalia v.ith 
aedeagus and one harpe omitted ; 15a, anellus, dorsal view ; 15b, 
aedeagus ; 15c, lateral view of tegumen, uncus, and gnathos. 
% 
MOTHS OF THE GENUS RUPELA HEINRICH 387 
Plate 27 
1G-16C. Rupela adnnca, new species: 16. Ventral view of male genitalia with 
aedeagus and one barpe omitted ; 16rt, anellus, dorsal view ; 16&, 
aedeagus ; 16c, dorsal view of uncus and gnathos. 
17-170. Rupela lumaria, new species: 17, Ventral view of male genitalia with 
aedeagus and one haipe omitted ; 17a, anellus, dorsal view ; 17&, 
aedeagus ; 17c, dorsal view of uncus and gnathos. 
18-18f. Rupela Jiorridnla, new species: 18, Ventral view of male genitalia with 
aedeagus and one harpe omitted ; 1S«, anellus, dorsal view ; 18?;, 
aedeagus with anellus attached, lateral view; 18c, apex of aedeagus 
lateral view showing cornutus (en) on penis; 18rf, 18c, lateral 
views of uncus and gnathos in two specimens showing extremes of 
variation in the uncus; 18/, dorsal view of gnathos and uncus. 
Plate 28 
19-19r. Rupela spinifera, new species : 19, Ventral vioAv of male genitalia with 
aedeagus and one harpe omitted; 19a, anellus, dorsal view, 19'j. 
aedeagus; 19c, dorsal view of uncus and gnathos. 
20-20C. Rupela monstrata, new species : 20, Ventral view of male genitalia with 
aedeagus and one harpe omitted ; 20a, anellus, dorsal view ; 206, 
aedeagus ; 20c, dorsal view of uncus and gnathos. 
Pl.ATE 29 
21. Rupela iendis, new species: Female genitalia. 
22. Rupela faiistina, new species: Female genitalia. 
23. Rupela canens, new species : Female genitalia. 
24. Rupela drusilla, new species : Female genitalia. 
25,25a. Rupela candace, new species: 25, Female genitalia, the projection to 
the side showing lateral view of genital opening, genital plate, and 
sclerotized pocket; 25ff, dorsal view of coUar of eighth abdominal 
segment. 
26-26?>. Rupela allinella (Cramer) : 26, Female genitalia; 26a, lateral view of 
organs with bursa omitted; 26f/, ventral view of genital plate and 
opening and eighth segment collar, from a specimen showing ex- 
treme of variation from normal specimens. 
Plate 30 
27. Rupela herie, new species: Female genitalia. 
28. Rupela gala, new species: Female genitalia. 
29. Rupela edusa, new species: Female genitalia. 
30. Rupela leucatea (Zeller) : Female genitalia. 
31. Rupela antonia, new species: Female genitalia. 
32. Rupela seyreya, new species: Female genitalia. 
Plate 31 
33, 34. Rupela nereis, new species : 33, Lateral view of femal genitalia ; 34, 
ventral view. 
35. Rupela orljona, new species : Female genitalia, the projection to the 
side showing lateral view of genital opening, genital plate, and 
sclerotized pocket. 
131080—37 3 
388 PROCEEDINGS OF THE NATIONAL MUSEUM vol.81 
36, 37. Rupela maenas, new species : 36, Lateral view of female genitalia ; 37, 
ventral view. 
3S. Rupela jana, new species: Female genitalia. 
Plate 32 
39. Rupela sejuncta, new species : Female genitalia. 
40. Rupela lara, new species : Female genitalia. 
41. Rupela procula, new species : Female genitalia. 
42, 4S. Rupela iinctella (Walker): 42, Lateral view of female genitalin ; 43, 
ventral view. 
Plate 33 
44,45,48. Rupela leucaiea (Zellcr) : 44, Seventh abdominal segment of female, 
showing sclerotization of sternite ; 45, ventral view of eighth abdomi- 
nal segment of male and part of seventli, showing sclerotizod plates 
and scale tuft ; 48, side view of head and expanded thoracic hair tuft. 
46. Rupela tincfclla (Walker) : Seventh abdominal segment of female, 
ventral view, showing sclerotization. 
47. Rupela h07'ridula, new species : Venation of fore and hind wings. 
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