A revision of the spider genus Cocalodes with a description of a new related genus (Araneae: Salticidae) F. R. Wanless Department of Zoology, British Museum (Natural History) Cromwell Road, London SW7 5BD. Introduction The genus Cocalodes Pocock, 1897, known from Amboina, Seram, Halmahera and New Guinea, is comprised of 12 species, six of which are described here as new. Petrunkevitch (1928) placed Cocalodes in the subfamily Boethinae, but as far as I am aware it shows no close affinities with the nominate genus Boethus or related genera (sensu Wanless, 1981). It is, however, closely allied to Allococalodes gen. n., proposed here for two new taxa from New Guinea. Both genera are unique in that the majority of males possess a well developed median horn arising from a sclerite between the lower basal margins of the chelicerae (Figs 8 A, 2 1 B). This structure has not been described in Cocalodes by earlier authors, for in the majority of species it does not protrude beyond the edge of the clypeus and is readily overlooked. The horn is not homologous with the paired horns ofPadilla Peckham & Peckham, which arise from the basal region of the anterior surface of the chelicerae. Neither is it homologous with the horn of Thorellia Keyserling which arises from the clypeus. The male palps of both Cocalodes and Allococalodes are also of interest in possessing a functional conductor and median apophysis which seldom occur in other Salticidae. In Allococalodes the functional conductor and median apophysis are lobe-like (Fig. 16E) and relatively simple when compared with the elaborate fan-like functional conductor and bifid median apophysis characteristic of Cocalodes (Fig. 6G). The less complex structures of Allococalodes are possibly vestigial or germinal in respect of those of Cocalodes, but for the present it is a matter for conjecture as the affinities of both genera are uncertain. The standard abbreviations and measurements are those used by Wanless (1978), but for the leg spination the system is that used by Platnick and Shadab (1975). Genus COCALODES Pocock Cocalodes Pocock, 1897:627. Type species Cocalodes leptopus Pocock, by original designation. Simon, 1901:400, 403^405. Waterhouse, 1902:80. Petrunkevitch, 1928:181. Neave, 1939, 1 : 778. Roewer, 1954 : 936. Bonnet, 1956 : 1 172. DEFINITION. Small or large spiders ranging from about 4*5 to 11 -Omm in length. Most species elongate and narrow with long slender legs, the abdomen usually marked with characteristic dark lateral bands; chelicerae robust, in males usually elongate and porrect with a median horn which sometimes protrudes beyond the clypeus; not hirsute, fringes lacking. Carapace: longer than broad, moderately high, widest at about level of coxae II-III; fovea long, weakly sulciform, positioned more or less midway between posterior margins of posterior lateral eyes. Eyes: with black surrounds except anterior medians; posterior medians and posterior laterals on moderately well developed tubercles; Bull. Br. Mus. not. Hist. (Zool.) 42(4): 263-298 Issued 24 June 1 982
BioStor
