PROC. BIOL. SOC. WASH. 100(2), 1987, pp. 358-363 ADDITIONAL MATERIAL OF ANHINGA GRANDIS MARTIN AND MENGEL (AVES: ANHINGIDAE) FROM THE LATE MIOCENE OF FLORIDA Jonathan J. Becker Abstract. —An associated partial skeleton and other previously unknown skel-etal elements of Anhinga grandis Martin and Mengel, 1975, are described from three late Miocene (latest Clarendonian and early Hemphillian) localities in Florida. Derived osteological characters of the tarsometatarsus show^. grandis to be more closely related to the New World A. anhinga than to the Old World A. rufa, A. melanogaster, and^. novaehollandiae. The wing of Anhinga grandis was larger than that of the largest living species, Anhinga novaehollandiae, but the pelvic limb was comparable to that species in size. Anhinga grandis was originally described from a single distal end of a humerus from the late Miocene Cambridge (=Ft.-40) local fauna, "Kimball" formation, Frontier County, Nebraska (Martin and Mengel 1975). The associated mammalian fauna, along with that from several other local fau-nas in western Nebraska, were used to typify a discrete post-Hemphillian, pre-Blancan Land Mammal Age, termed the "Kimball-ian" (Schultz et al. 1970). Subsequent work (Breyer 1981, Voorhies 1984) has demon-strated that no stratigraphic nor biochrono-logic evidence exists to support this pro-posal. The Cambridge local fauna is now considered to originate in the Ash Hollow Formation and to be early Hemphillian in age, about 8.5 MYBP (=million years be-fore present) (Tedford et al., in press). This paper describes new material refer-able to Anhinga grandis from three late Miocene localities that extends the range of the species to Florida and provides addi-tional data on its morphology and system-atic relationships to other species in the ge-nus Anhinga. Materials and Methods Fossil specimens included in this study are housed in the Vertebrate Paleontology collections of the Florida State Museum (UF) and the University of Nebraska State Museum (UNSM). Modern comparative material came from the collections of Pierce Brodkorb; Florida State Museum (UF); Na-tional Museum of Natural History, Smith-sonian Institution (USNM); American Mu-seum of Natural History (AMNH); and Royal Ontario Museum (ROM). Modern species and number of specimens examined are as follows: Anhinga anhinga ( 1 7), A. rufa (10), A. melanogaster (1), A. novaehollan-diae (1). Anatomical terminology follows Baumel et al. (1979) and Howard (1929). Measurements given in Tables 1 and 2 are defined below. HUMERUS: LENGTH -Greatest length from the head of the humerus through the midpoint of the lateral condyle. LATIS— Greatest length from the proximal-most ex-tension of the tuberculum for the attach-ment of the posterior head of the latissimus dorsi through the midpoint of the lateral condyle. W-SH AFT— Transverse width of midshaft. D-SHAFT-Depth of midshaft. W-DIST— Transverse width of distal end from the entepicondylar prominence to the ectepicondylar prominence. D-DIST — Depth of distal end from cranial face of ex-ternal condyle through ridge slightly mediad
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