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PROC. BIOL. SOC. WASH. 102(2), 1989, pp. 483^90 DESCRIPTION OF TWO NEW SPECIES OF THE ELEUTHERODACTYLUS MILESI GROUP (AMPHIBIA: ANURA: LEPTODACTYLIDAE) FROM NORTHERN HONDURAS James. R. McCranie, Jay M. Savage, and Larry David Wilson Abstract.— Two new species of the Eleutherodactylus milesi group (E. chry-sozetetes and E. cruzi) are described from the Cordillera de Nombre de Dios of northern Honduras. Eleutherodactylus milesi is shown to have a wide range in Honduras on scattered montane ridges and is reported from Guatemala for the first time. The systematics of Eleutherodactylus ma-tudai and E. milesi were reviewed by Savage (1975), who included them in the E. rugu-losus group. These species subsequently were removed from the rugulosus group (Miya-moto 1983, Savage 1987) and placed as a separate infrageneric lineage, the E. milesi group (Savage 1987). Members of this stock differ in karyology (2N = 22; NF = 44) and jaw musculature (DFSQAT + e) from those of the rugulosus group (2N = 20, 22; NF = 36; jaw muscle formula: dfsq; dfsqat + e). Externally, frogs of the milesi group are dis-tinct (features for the rugulosus group in pa-rentheses) in: lacking tarsal folds (a well-developed inner tarsal fold), having a strongly tuberculate dorsum (dorsum smooth to rugose), and having a pair of para-anal light bars (never with such markings). In addition E. matudai has an indistinct tympanum in females and E. milesi has the tympanum indistinct or hidden (distinct, nearly transparent tympanum). Recent fieldwork in Guatemala and Hon-duras substantially increases the known range for Eleutherodactylus milesi and this information and data on variation are pre-sented here. Among other newly collected material from the Cordillera de Nombre de Dios of northern Honduras are two forms, clearly allied to E. matudai and E. milesi, but trenchantly different from them and from one another. These distinctive frogs are regarded as representing previously un-known species and are described below fol-lowing the style and terminology of Savage (1 975); a few features not mentioned in that paper follow the definitions of Savage (1987). Eleutherodactylus chrysozetetes, new species Fig. 1 Holotype.-KU 209035, adult male, from Quebrada de Oro (15°38'N, 86°47'W), ele-vation 880 m, tributary of Rio Viejo, south slope of Cerro Bufalo, Cordillera de Nombre de Dios, Departamento de Atlantida, Hon-duras, 17 Aug 1984, James R. McCranie, Kenneth L. Williams, and Larry David Wil-son. Original number LDW 6529. Diagnosis. —Eleutherodactylus chrysoze-tetes is a moderately large frog and is distin-guished from the two previously described members of the milesi group, E. matudai and E. milesi (characteristics for these species in parentheses), in having extensive toe webbing with about three phalanges free of the web on toe IV (basal to moderate webbing, with four to four and one-half pha-langes free of web); strongly expanded finger and toe disks, about twice width of digit (finger disks definite, less than twice width of digit; toe disks weak, barely expanded);

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Description Of 2 New Species Of The Eleutherodactylus milesi Group

J R Mccranie, J M Savage and L D Wilson
Proceedings of The Biological Society of Washington 102: 483-490 (1989)

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