STUDIES ON THE TETRACLITIDAE (CIRRIPEDIA: THORACICA): REVISION OF TETRACLITA ARNOLD ROSS ABSTRACT. — Tesseropora and Tetraclitella. subgenera of Telraclita, are elevated to full generic status ir the absence of intergrading major characters. Newmanella gen. nov. is proposed for Telraclita (Telraclita radiata (Bruguiere, 1789) and Tesseroplax gen. nov. for Telraclita ( Tesseropora ) unisemita Zullo. 1968. On th< basis of shell characters the five genera here recognized fall into two groups. Monometric growth, allometry 01 the carina, and non-tubiferous radii characterize Tesseropora, Tesseroplax, and Telraclita; diametric growth isometry of the wall plates, and tubiferous radii characterize Tetraclitella and Newmanella. Both Tesseropla.\ and Tetraclita are considered later derivatives from the Tesseropora stock; but Tetraclitella and Newmanellc from a tesseroporan precursor. New distributional records for Newmanella radiata include the Bahamas Puerto Rico, Dominica, and Venezuela. Apparently the antenniform ramus of cirrus III in N. radiata play; some role during the reproductive cycle, but its exact function is unknown. RESUMEN. — Tesseropora y Tetraclitella, subgenero de Tetraclita. estan elevados al estado generico en a ausencia de intergradacion caracteres mayores. Newmanella gen. nov. esta propuesto para Tetraclitc (Tetraclita I radiata (Bruguiere, 1789) y Tesseroplax gen. nov. para Tetraclita I Tesseropora ) unisemita Zullo 1968. En el base de caracteres de concha los cincogeneros reconocidos aquT, caen en dos grupos. Crecimientc monometrico, alometria de la carena, y radios no-tubiferoso caracterizan Tesseropora. Tesseroplax. \ Tetraclita; crecimiento diametrico, isometrfa de la laminas de las paredes, y radios tubiferoso caracterizar Tetraclitella y Newmanella. Los dos Tesseroplax y Tetraclita son considerados derivatives mas tarde de lo; progenitores de Tesseropora, pero Tetraclitella y Newmanella de un progenitor de Tesseropora. Registro; distribucionales nuevos por Newmanella radiata se incluyen las Bahamas, Puerto Rico, Dominica, > Venezuela. Aparentemente, la ramal antenniforma de cirrus III en N. radiata hace un papel durante lo* reproducti vos, pero su funcion exacto es desconocido. The Tetraclitidae comprise a group of intertidal balanomorph barnacles occurring in tropical and warm-temperature waters between latitudes 38° north and 52° south. Although! these barnacles are a major component of intertidal faunas they have not received adequate attention either taxonomically or biologically. In this, the first in a series of papers covering the taxonomy and general biology of the tetraclitids, I re-evaluate the status of Tetraclita and its subgenera, and propose a new classification in the light of recent studies (Ross, 1968), and the work of Darwin ( 1 854), Nilsson-Cantell ( 1 92 1 ) and Hiro ( 1 939). There are more than two dozen named taxa distributed presently between Tetraclita Schumacher, 1817 (type species: T. (Tetraclita) squamosa (Bruguiere), 1789), and its subgenera, Tesseropora Pi lsbry, 1916 (type species: T (Tesseropora) rosea (Krauss), 1848), and Tetraclitella Hiro, 1939 (type species: T (Tetraclitella) purpurascens (Wood). 1815). These subgenera are distinct, and their elevation to generic status clarifies relationships that are obscured when all of the species are considered monogeneric. Both Darwin (1854: 344) and Hiro (1939: 270) cited numerous morphological characters that support the generic distinctness of the Caribbean western Atlantic species. / radiata. These characters, coupled with the absence of intergrading major morphological structures, necessitate the proposal of Newmanella gen. nov. for this species. Attempts to assign the recently described Pliocene species from the Gulf of California, Mexico. T (Tesseropora) unisemita (Zullo, 1968: 273), within the proposed classification posed innumerable problems, and consequently I have placed it in a new genus, Tesseroplax. SAN DIEGO SOC. NAT. HIST. TRANS 15(15): 237-251, 22 SEPTI MM R 1969
BioStor
