The carabid beetles of New Guinea. Part IV. General considerations; analysis and history of fauna; taxonomic supplement

THE CARABID BEETLES OF NEW GUINEA. PART IV. GENERAL CONSIDERATIONS; ANALYSIS AND HISTORY OF FAUNA; TAXONOMIC SUPPLEMENr p. J. DARLINGTON, JR. CONTENTS Introduction to Part IV. 133 1. Purpose; previous parts; acknowledg-ments 133 2. Sources, disposition, and adequacy of material 133 3. Preparation for work on New Guinean Carabidae; my collecting 135 4. Basic literature 136 5. Localities 138 Policies and Methods 143 6. Modern taxonomy 143 7. Types 146 8. Taxon concepts: subfamilies and tribes 147 9. Genera 147 10. Species 148 11. Subspecies 149 12. Nature of taxonomic characters 150 13. Secondary sexual and genitalic char-acters 151 14. Preservation of material 152 15. Taxonomic methods and procedures __ 153 16. Data sheets 154 Analysis and Discussion 156 17. New Guinea 156 ^The author suggests that, when the four parts of TJie Caiabid Beetles of New Guinea are bound together, the present part be divided, and the whole bound in the following order: general con-siderations, analysis, and history of the fauna from Part IV; Part I; Part II; Part III; and the taxonomic supplement from Part IV. ■ Work and publication supported by National Science Foundation Grant GB-12346. Bull. Mus. Comp, 18. The New Guinean carabid fauna: tax-onomic composition 157 19. Numbers of species 158 20. Size of individuals 161 21. Wings and wing atrophy 165 22. Explanations of wing atrophy 170 23. Summary of wing state and wing atrophy 172 24. Ecology: habitats 172 25. Ecologic composition 173 26. Altitude 175 27. Ants : 176 28. Ecologic interactions 176 29. Ecologic ranges 177 Zoogeography: Existing Geographic Patterns .. 178 30. Geographic patterns, relationships, and origins 178 31. Existing geographic relationships: problems and procedures _. 178 32. Distribution of carabid tribes from Asia to Australia 180 (Cicindelinae) _ 182 33. Ozaenini 183 34. Paussini 183 35. Scaritini 183 36. Bembidiini 184 37. Trechini 185 38. Panagaeini 186 39. Pterostichini 186 40. Agonini 188 41. Perigonini 189 42. Licinini 189 43. Chlaeniini 190 44. Oodini 190 45. Harpalini 190 46. Anaulacini 192 47. Gyclosomini 192 48. Lebiini _____ __. 193 49. Pentagonicini 196 50. Hexagoniini 197 Zool., 142(2): 129-337, October, 1971 129