PROC. BIOL. SOC. WASH. 103(4), 1990, pp. 955-961 CROCODYLUS RANINUS S. MULLER AND SCHLEGEL, A VALID SPECIES OF CROCODILE (REPTILIA: CROCODYLIDAE) FROM BORNEO Charles A. Ross Abstract. — Cwcodylus raninus S. Miiller & Schlegel 1844, described on the basis of two syntypes from Borneo, is resurrected and diagnosed. Squamation and cranial morphology of C. raninus are compared with other sympatric and neighboring Indopacific species. The systematics and distribution of In-dopacific Crocodylus are inadequately known. Many museum specimens identi-fied as the widely distributed Crocodylus po-rosus Schneider, 1801, actually represent unrelated and poorly known insular species. This is particularly true in the Indonesian Archipelago, where specimens of unde-scribed Crocodylus have been examined from Banka Island and Sulawesi (Ross 1986). Unfortunately, diagnoses of many of these populations are precluded by insufficient material and complicated by a long history of misidentifications in the literature. One exception is the palustrine crocodile of Bor-neo which, although equally confused in the older literature and known from very few specimens, can be diagnosed on the basis of limited museum material and as an une-quivocal name is available, its resurrection is uncomplicated. Methods Scale counts used in this study were made on 481 individual crocodiles from the In-dopacific region. Specimens were examined from museums, private collections, and from the wild. Some specimens critical to this study in European museums were not examined autoptically but photographs of them were examined and scale counts made from the photographs. Scale counts follow Ross & Roberts (1979), King & Brazaitis (1971), and Bra-zaitis (1973, 1974). Dorsal and neck armor terminology of F. Ross & Mayer (1983) is given in parentheses where pertinent. Museum abreviations follow Leviton et al. (1985). Historical Discussion S. MuUer & Schlegel (1844) described Crocodylus raninus (as a race of C. bipor-catus Cuvier = C. porosus) on the basis of an adult skull and a whole juvenile from Borneo. They differentiated C. raninus from C porosus on the basis of snout shape and neck squamation but the name was not for-mally proposed and specific locality data and disposition of the syntypes were not included. Many crocodile specimens col-lected by M. S. Miiller from Borneo in 1 836 were in the Museum d'Histoire Nalurelle des Pays-Bas and listed by Lidth de Jeude (1898) in the "Catalogue Oslcologiquc." These specimens are individualh identified by letter of the alphabet and are now in the Rijksmuseum van Naluurlijkc Historic, Leiden. The syntypes of C\ raninus were noi specifically noted by Lidth de Jeude ( 1 898) and have not been located. They were, how-ever, clearly figured (S. Miiller & Schlegel 1844, pi. 3, figs. 7, 8), and show an adult skull (fig. 7) and the neck and head of a juvenile (fig. 8). There is no indication that any subsequent worker examined the s-



Crocodylus raninus S. Müller and Schlegel, a valid species of crocodile (Reptilia: Crocodylidae) from Borneo

Charles A Ross
Proc. Biol. Soc. Washington 103(4): 956-961 (1990)

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