145 
REVISION OP THE GENUS BRACHYCOME CASS. 
PART III. DESCRIPTION OF THREE NEW AUSTRALIAN SPECIES AND SOME NEW 
LOCALITY RECORDS. 
By Gwenda L. Davis, B.Sc, Lecturer in Biology, New England 
University College, Armidale. 
(Seven Text-figures.) 
[Read 25th May, 1949.1 
Introduction. 
Recent intensive collection of this genus by Victorian botanists has brought to hand 
several specimens which are quite distinct from any described species. Similar collecting 
in other States will probably increase the number of known species considerably and 
at the same time lead to a more accurate knowledge of the distribution of those 
already described. 
Taxonomy, 
compositae, tribe asteroidea. 
Brachycome Cass., Diet. Sci. Nat., xxxvii (1825), 471. 
Subgenus Eubrachycome G. L. Davis. 
Superspecies tenuiscapa. 
brachycome stolonifera, sp. nov. (Text-figures 1-3.) 
Holotype: Summits of Kosciusko plateau, N.S.W., ca. 7,000 ft., 3, 1947, A. Costin (MEL). 
Paratypes: Eleven, loc. cit. (MEL). 
Herba perennis, erecta, glabra, stolonifera, ad 6-7 cm. alta; folia ad 3-2 cm. longa, 
1-2 mm. lata, radicalia, lato linearia, integra, obtusa, basibus involutis; omnis herba 
1-4 stolonibus quorum folia, brevia sed typica, ultima parti radicata, moriuntur. 
Capitula sola, 6-7 mm. transverse lata; pedunculi uno phyllo robusti; involucri phylla 
18-24, circiter 4-5 mm. longa, 1-4 mm. lata, lanceolata, acuminata, marginibus fimbriato- 
ciliatis; flores radii circiter 38, ligulis 6 mm. longis, 2-1 mm. latis, albis; receptaculum 
2 mm. latum, 2 mm. altum, praecipiter conicum, punctum; achaenia 1-8 mm. longa, 
0-9 mm. lata, fusca, cuneata, in septato-piloso disco turgida, margine angusto et tenui 
et in labro septato-piloso; pappus setis, sericis, albis, conspicuis. 
Erect glabrous stoloniferous perennials up to 6-7 cm. high. Leaves up to 3-2 cm. 
long, 1-2 mm. broad, radical, broad-linear, entire, obtuse with sheathing bases. Each 
plant bears 1-4 stolons on which the small but typical leaves die off when the tip 
becomes rooted. Inflorescences solitary, 6-7 mm. diameter. Scapes robust and provided 
with a single bract. Involucral bracts 18-24, about 4-5 mm. long, 1-4 mm. broad, 
lanceolate, acute, with torn-ciliate margins. Ray florets about 38, rays 6 mm. long, 
2-1 mm. broad, white. Receptacle 2 mm. broad, 2 mm. high, steeply conical, pitted. 
Fruit 1-8 mm. long, 0-9 mm. broad, dark brown, cuneate, thick in central region, with 
a narrow thin margin. Septate hairs are scattered over the central area and along 
the edge of each margin. Pappus of silky white conspicuous bristles. 
Habitat : Not recorded. 
Range : Only known from the type locality. 
Specimens examined : Type series only. 
This species is vegetatively very similar to certain specimens of B. nivalis var. 
alpina (P. Muell. ex. Benth.) G. L. Davis, and B. raclicans Steetz, which is possibly the 
reason it has been overlooked for so long. The fruit structure is closest to B. Stuartii 
Benth., but the margin is thinner and wider and the longitudinal folds are absent. 
146 
REVISION OF THE GENUS RRACHYCOME CASS. PART III, 
Text-figures 1-7. 
1-2. B. stolonifera. Holotype and paratype. x J. 3. B. stolonifera. Fruit, x 20. 
4. B. obovata. Holotype. x J. 5. B. obovata. Fruit, x 20. 6. B. petrophila. Holotype. x i. 
7. B. petrophila. Fruit, x 20. 
brachycome obovata, sp. iiov. (Text-figures 4-5.) 
Holotype : Echo Flat, Lake Mountain, about 4,700 ft., margin of alpine sphagnum bogs, 
25.1.1948, J. H. Willis (MEL). 
Paratypes: Two, loc. cit. (MEL). 
Herba perennis, glabra, ad 19 cm. alta, radicalibus foliis in basi. Folia ad 10-5 cm. 
longa, 1-1-3 mm. lata, linearia — lineari-obovata, integra, quorum latiora in basi petiolum 
habent, Capitula 1-2; 8-9 mm. transverse lata; pedunculi 4-6, foliosis phyllis; involucri 
phylla 22-26, 4-3-4-5 mm. longa, 1-1-1-5 mm. lata, subacuminata-acuminata, serrata, 
glabra aut in parti exteriora mic. glandulosa; flores radii circa 24, 5 mm. longi, 1-4 mm. 
lati, simulate albi-caerulei; achaenia 2-3-2-5 mm. longa, 1-1-3 mm. lata, fusca, obovata, 
levia et planata; pappus setis brevibus fulvis. 
Erect glabrous perennials up to 19 cm. high with a basal cluster of radical leaves. 
Leaves up to 10-5 cm. long, 1-1-3 mm. broad, linear to narrow-obovate, entire, the broader 
leaves tapering proximally into a petiole. Inflorescences 8-9 mm. diameter, 1 or 2 present 
on each plant. Scapes provided with 4-6 leaf-like bracts. Involucral bracts 22-26, 
4-3-4-5 mm. long, 1-1-1-5 mm. broad, subacute to acute with serrulate margins and 
either glabrous or microscopically glandular on the outer surfaces. Ray florets about 
24, rays 5 mm. long, 1-4 mm. broad, apparently white to bluish. Fruit 2-3-2-5 mm. long, 
1-1-3 mm. broad, light brown, obovate, smooth and flattened. Pappus of short straw- 
coloured bristles. 
Habitat : Apparently swampy conditions at high elevations. 
Range : Eastern Victoria. 
Specimens examined : 
Victoria : Echo Flat, Lake Mountain, about 4,700 ft., margin of alpine sphagnum bogs, 
25.1.1948, J. H.' Willis (MEL. holotype and paratypes) ; Snowy Mountains, 6-7,000 ft., 
2.1890, W. Bauerlen (MEL) ; Genoa district, 3.1885, W. Bauerlen (MEL). 
BY OWETTOA L. DAVIS. 147 
The above series, although small, shows a certain amount of variation in the shape 
of the leaves. Those of specimens from Lake Mountain and Genoa agree in that they 
are linear and grasslike and the plants resemble certain specimens of B. radicans Steetz 
ex Lehmann. On the other hand, the specimens from Snowy Mts. bear narrow-obovate 
leaves and are very similar to B. nivalis var. alpina (F. Muell. ex Benth.) G. L. Davis. 
It is anticipated that more critical collecting in the highlands of Victoria will produce 
specimens intermediate between these two forms of B. obovata. 
The fruits are very distinctive but are superficially similar to those of B. graminea 
(Labill.) F. Muell. They can, however, be readily distinguished by the small but 
distinct pappus. Dissection of the fruit shows that the whole of the interior is 
occupied by the embryo and cotyledons, whereas in B. graminea these are completely 
enclosed by the spongy margins. For this reason the affinities of B. obovata are to be 
found within the superspecies tenuiscapa. 
beach ycome PETROPHiLA. sp. nov. (Text-figures 6-7.) 
Holotype : Little River Falls, about five miles N.E. of Wulgulmerang, E. Vic, "on dripping 
cliff faces", 16.1.1948, J. H. Willis (MEL). 
Paratypes : Four, loc. cit (MEL). 
Herba perennis, inferme erecta aut adscendens, ramosa, glanduloso-pubescens, ad 
33-5 cm. alta; folia caulina, ad 4-5 cm. longa, 1-7 cm. lata, grosse orbicularia, superiora 
plerumque sessilia, dentata, 5 utrimque dentibus, inferiora petiolo brevi et interdum 
fere crenata; capitula 1-3, 6-7 mm. transverse lata; pedunculi graciles, axillares, 
6-5-10-7 cm. longi, uno folioso phyllo; involucri phylla circiter 24, ad 4 mm. longa, 
1 mm. lata, angusto-lanceolata, acuminata, in exteriora parti glandulosa, marginibus 
fimbriato-ciliatis; flores radii circiter 28, ligulis 6 mm. longis, 1-3 mm. latis, albis; 
receptaculum 3 mm. latum, 2-1 mm. altum, hemisphaericum, punctum; achaenia 1-9 mm. 
longa, 0-9 mm. lata, fusca, cuneata, compressa, margine angusto glabro 2 longis sinibus 
utrimque separato; discum tubercula pauca, parva, sparsa, in ultima parti mic. 
glanduloso pilo, habet; pappus clarus, fulvus, setis in fasciculis inaequibus dispositis. 
An apparently weakly erect or ascending branching ? perennial, shortly glandular 
pubescent all over and up to 33-5 cm. high. Leaves cauline, up to 4-5 cm. long, 1-7 cm. 
broad, orbicular in gross outline, mainly sessile but the lower ones tapering proximally 
into a short petiole; upper leaves dentate with usually five teeth on either side, 
lower leaves sometimes almost crenate. Inflorescences 1-3 on each plant, 6-7 mm. 
diameter, and borne on slender axillary peduncles 6-5-10-7 cm. long, each of which 
is provided with a small leaf-like bract. Involucral bracts about 24, up to 4 mm. long, 
1 mm. broad, narrow-lanceolate, acute, glandular on the outer surface and with torn 
ciliate margins. Ray florets about 28, rays 6 mm. long, 1-3 mm. broad, white. 
Receptacle 3 mm. broad, 2-1 mm. high, hemispherical, pitted. Fruit 1-9 mm. long, 
0-9 mm. broad, dark brown, cuneate, flat, with two longitudinal folds down each 
side, cutting off a narrow smooth margin. Central region of fruit bears a few small 
scattered tubercles, each of which is tipped with a microscopic glandular hair. Pappus 
conspicuous, straw-coloured, the bristles grouped in bundles of irregular length. 
Habitat: "In moist shaded places in soil amongst the rocks, on ledges or in crevices" (N. A. 
Wakefield). 
Range : Eastern Victoria. 
Specimens examined : 
Victoria: Little River Falls, about five miles N.E. of Wulgulmerang, E. Vic, "on dripping- 
cliff faces", 16.1.1948, J. H. Willis (MEL, holotype and paratypes) ; Little River Falls, 
"Wulgulmerang, "on ledges and in crevices amongst moist porphyry rocks, 2,500 ft., 
17.1.1948, N. A. Wakefield (NAW) ; Murrundal River, "in shaded crevice of porphyry 
rock, about 500 ft.", 19.10.1947, N. A. "Wakefield (NAW). 
Vegetatively this species is very similar to B. Nova-Anglica G. L. Davis, and the 
leaves, though larger, are of the same type. Both species appear to occupy the same 
type of habitat. 
In details of the fruit (i.e. shape, presence of longitudinal folds and a narrow 
margin) a definite affinity is indicated with B. Stuartii Benth,, but a pappus of the 
type present in B. petrophila has not been previously described in the superspecies. 
148 REVISION OF THE GENUS BRACHYCOME CASS. PART III, 
Superspecies tenuiscapa. 
Amended Key to the Species. 
(1). Herbs with a glandular indumentum. Fruit cuneate and flat. 
(2). Leaves radical, oblanceolate to ovate-cuneate, distally dentate, with sheathing bases. 
Fruit 1-2 mm. long, 0-9 mm. broad, smooth, with slightly thickened margins. 
Pappus short and white B. tenuiscapa. 
(2).* Leaves cauline, orbicular in gross outline, dentate to almost crenate, usually sessile. 
Fruit 1-9 mm. long, 0-9 mm. broad, with a longitudinal fold down each side 
and smooth margins. A few tubercles are present on the central area. Pappus 
conspicuous and straw-coloured, bristles grouped in bundles of irregular length 
B. petrophila. 
(1).* Glabrous herbs with radical leaves. 
(3). Leaves entire, linear to oblanceolate or narrow-obovate. 
(4). Base of the plant surrounded by the dead remains of former leaves. Fruit 
2-3-3 mm. long, 0-8-1-2 mm. broad, glabrous, the margins slightly thickened 
and smooth. Pappus short B. scapigera. 
(4).* Bases of dead outer leaves not persistent. 
(5). One to four leafy stolons present on each plant. Fruit 1-8 mm. long, 
0-9 mm. broad, cuneate, thick in central region, with a narrow margin. 
Pappus conspicuous B. stolonifera. 
(5).* No development of stolons. Fruit 2-3-2-5 mm. long, 1-1-3 mm. broad, 
obovate, smooth and flattened. Pappus short B. obovata. 
(3).* Leaves pinnatisect, crenate or acutely toothed. 
(6). Leaves oblong-cuneate to elliptical, crenate to acutely toothed distally. Fruit 
3 mm. long, 1-4 mm. broad, with relatively long straight hairs on each flat 
surface and a shallow longitudinal groove down the outer edge of each 
slightly thickened margin. Pappus short B. decipiens. 
(6).* Leaves pinnatisect, the segments entire or again divided. Fruit 0-9-1-7 mm. 
long, 0-5-0-9 mm. broad, with two longitudinal folds on each face. Pappus 
short, bristles of unequal length B. Stuartii. 
Since the publication of Part I (Proc. Linn. Soc. N.S.W., lxxiii, 1948, 142-241) of 
this series further specimens have been examined from the herberia of two private 
collectors. In addition, a large amount of unworked material collected by Mueller 
and his contemporaries has recently been found in the National Herbarium, Melbourne. 
Although the majority of these specimens, are from well-collected localities, and therefore 
of no great interest, a number of them establish new locality records and extend the 
known range of certain species, in some cases, considerably. These fresh records are 
recorded under the appropriate species, the present location of each specimen being 
indicated as follows: 
National Herbarium, Melbourne (MEL). 
F. A. Rodway (FAR). 
N. A. Wakefield (NAW). 
Superspecies tenuiscapa. 
erachycome tenuiscapa Hook, f., var. a tenuiscapa (Hook, f . ) G. L. Davis. 
Proc. Linn. Soc. N.S.W., lxxiii, 1948, 153. 
New South Wales: Moona R., Walopa (MEL). 
Tasmania: Cradle Mt., 12.1915, F. A. Rodway (FAR); Cradle Valley, 12.1915, F. A. Rodway 
(FAR). 
This species has not previously been recorded from New South Wales, and 
unfortunately the locality has not been traced. Since "N.S.W." is recorded on the 
label it is assumed that Walopa is either too small to be indicated on the maps 
consulted or is now known by another name. 
brachycome tenuiscapa Hook, f., var. j3 pubescens (Benth.) G. L. Davis. 
Proc. Linn. Soc. N.S.W., lxxiii, 1948, 153. 
Neiv So%ith Wales: Swamp near Tenterfield, C. Stuart, n. 925 (MEL). 
brachycome scapigera (Sieb. ex Spreng) DC, 
Prod, vii, (1838), 277. 
New South Wales: Braidwood district, 3,000 ft., 10.1886, W. Bauerlen (MEL), 
BY GWENDA L. DAVIS. 149 
BRACHYCOME DECIPIENS Hook, f., 
London Journ. Bot., vi, (1847), 114. 
Neiv South Wales: *Near Dubbo, 1882, J. M. Curran (MEL); Upper Macquarie R., 10.1882, 
J. M. Curran (MEL). 
Victoria: Hume R., 1883. Jephcott (MEL); *Upper Livingstone, Omeo, 1882, Howitt (MEL); 
Slopes around Omeo, metamorphic schist for 2,200 ft. above sea level, 26.9.1882 (MEL) ; 
Gippsland, 1882, Howitt (MEL) ; Daylesford, 1880, Wallace (MEL) ; Ballarat, 9.1884, 
D. M. Spence (MEL) ; near Geelong, 1884, Wilson (MEL). 
Tasmania: *Victoria Valley, past R. Ouse crossing on road to L. St. Clair, 12.1917, F. A. 
Rodway (FAR). 
This species was previously recorded in New South Wales only from the southern 
highlands. 
BEACHYCOME STUARTII Benth., 
Fl. Aust., iii, (1866), 513. 
New South Wales: Mudgee, Woolls (MEL). 
These specimens depart from the typical condition in that they are glandular hairy 
and the pappus is microscopic. Hitherto this species was thought to be confined in 
New South Wales to the New England Tableland. 
Superspecies leptocaepa. 
BEACHYCOME DEBILIS Sond., 
Linnaea, xxv, (1852), 477. 
New South Wales: Lower Edward's R., Mein (MEL). 
Victoria: Little R., Fullager (MEL); Werribee, Fullager (MEL); Wimmera, 1890, J. Eckert 
(MEL). 
Specimens from the above Victorian localities were intermixed with vegetatively 
identical ones of B. leptocarpa F. Muell., an association previously noted (Davis, 1948). 
The range of this species is now extended to south-western New South Wales, west of 
Port Phillip and western Victoria. 
BEACHYCOME ANGUSTIFOLIA A. Cuilll. ex DC. Vai\ a ANGUSTIFOLIA (A. Clllin. ex DC) 
G. L. Davis, 
Peoc Linn. Soc. N.S.W., lxxiii, 1948, 161. 
New South Wales: Foot of the Pulpit Rock, 5J miles south-west of Nowra, Sandstone, 
22.3.1947, F. A. Rodway (FAR) ; Nowra, roadside, 1.1921, F. A. Rodway (FAR) ; Grassy 
Gully, Shoalhaven R., 14 miles west of Nowra, moist slope, 3.5.1941, F. A. Rodway (FAR). 
BEACHYCOME ANGUSTIFOLIA A. Cunil. eX DC. Vai'. /3 HETEEOPHYLLA (Benth.) 
G. L. Davis, 
Peoc. Linn. Soc N.S.W., lxxiii, 1948, 162. 
New South Wales: Joadga (Southern Tablelands), 10.1919, F. A. Rodway (FAR). 
Victoria: Prince's Highway at Karlo Crk., Mt. Drummer and Wingan River, E. Vic, 25.1.1947, 
J. H. Willis (MEL). 
Previously this species was only recorded from New South Wales, where the 
southern limit was Newcastle, so the range is extended considerably. These specimens 
differ from the type series in that the leaves are orbicular and regularly dentate, and 
Bentham's type specimens are to be regarded as representing one extreme of a 
variable series. 
BEACHYCOME DISSECTIFOLIA G. L. Davis, 
Peoc. Linn. Soc. N.S.W., lxxiii, 1948, 163. 
Victoria: Moyston, 10.1881, D. Sullivan (MEL); Mt. William Creek, 10.1879, D. Sullivan 
(MEL). 
This species has not been previously recorded from Victoria. 
BEACHYCOME PEOCUMBENS G. L. Davis, 
Peoc Linn. Soc. N.S.W., lxxiii, 1948, 164. 
Victoria: Gorge country of the Upper Snowy River, near Deddick, 21.1.1948, J. M. Bechervaise 
(MEL). 
The fruits of these specimens show a small variation from those of the type series 
in that the wing is approximately equal to the breadth of the body. 
150 REVISION OF THE GENUS BRACHYCOME CASS. PART III, 
This is the first record of this species from Victoria, where it appears to occupy 
a similar habitat to that of specimens in New England. 
BRACHYCOME LINEARILOBA (DC) Dl'UCe, 
Rep. Bot. Excli. CI. Brit. Isles, iv (1917), 610. 
New South Wales: Trundle, 8.1916, H. M. R. Rupp (PAR). 
South Australia: Charlotte Waters, 1889, W. Schwartz (MEL). 
Western Australia: Eucla, 1877, Richards (MEL); Eucla, 18S2, J. Oliver (MEL). 
bbachycome gbaminea (Labill.) F. Muell., 
Frag. Phytog., i, (1858), 49. 
New South Wales: Moonie Creek, Jervis Bay, 4.1916, 2.1921, F. A. Rodway (FAR); Tabourie 
Island, 15.5.1938, F. A. Rodway (FAR). 
This species is not confined to the tablelands of New South Wales, as was previously 
stated. 
Superspecies basaltica. 
BRACHYCOME BASALTICA F. Muell. Vai\ j8 GRACILIS Beiltll., 
Fl. Aust., iii (1866), 515. 
New South Wales: Balranald, 1S7S, Lucas (MEL); Murrumbidgee R., 9.1878, F. Mueller 
(MEL). 
Victoria: Swan Hill district, 10.1888, C. French (MEL); Albury, 1890, J. Wilson (MEL); 
Western border of Victoria, C. Walter (MEL). 
These records are interesting in that they link up those already recorded and 
support the suggestion advanced previously (Davis, 1948) that this variety migrated 
to Queensland along the basin of the Darling River. 
BRACHYCOME MULTIFIDA DC, Vai\ a MULTIFIDA (DC) G. L. Davis, 
Proc. Linn. Soc. N.S.W., lxxiii, 1948, 181. 
Victoria: *Bolwarra, near Ballarat, 1S82, W. H. Wooster (MEL) ; Bendigo, 12.10.1892 (MEL). 
These specimens extend the range of this species into central Victoria. 
Superspecies aculeata. 
BRACHYCOME PAPILLOSA G. L. Davis, 
Proc. Linn. Soc. N.S.W., lxxiii, 1948, 191. 
New South Wales: Murrumbidgee R., 9.1878, F. Mueller (MEL) ; Murrumbidgee, 1885, Bruckner 
(MEL). 
BRACHYCOME MUELLERI Solid., 
Linnaea, xxv (1852), 475. 
South Australia: Yorke's Penin, 1888, Beythieu (MEL). 
In this specimen the fruits are smaller than previously recorded, being 1-3 mm. 
long, 0-9 mm. broad. 
BRACHYCOME MUELLEROIDES G. L. Davis, 
Proc. Linn. Soc. N.S.W., lxxiii, 1948, 194. 
Victoria : Granite summits of the mountains on the lower Snowy R., 1.1874, F. Mueller (MEL). 
This species was known previously only from two localities, Wagga and Nathalia, 
consequently further records are of considerable interest in determining the limits of 
variation. The specimens from the Snowy River bear fruit 1 mm. long and 1 mm. 
broad, so that they are slightly larger than those of the type series. A further 
difference is apparent in that the body of the fruit is dark brown and the wings golden 
brown. Vegetative variation is shown in the leaves, which bear 3-6 acute linear lobes. 
brachycome cardiocarpa F. Muell. ex. Benth., 
Fl. Aust., iii (1866), 517. 
New South Wales: Upper Darling R., near Queensland border, 1SS4, L. Henry (MEL). 
Victoria: Lower Glenelg R., 1891, Eckert (MEL). 
Tasmania: *Cape Portland, 1S84, Bandinet (MEL); Lake Crescent, 9.12.1938, E. P. Rodway 
(FAR). 
Hitherto this species has been recorded in New South Wales only from the Southern 
Highlands. 
BY GWENDA L. DAVIS. 151 
brachycome diversifolia (Grah. in Hooker) Fisch and Mey, var. 7 dissecta 
G. L. Davis, 
Peoc. Linn. Soc. N.S.W., lxxiii, 1948, 202. 
Victoria: Treasure's Homestead, Dargo High Plains, E. Vic, 4,500 ft., 28.1.1946, J. H. Willis 
(MEL). 
This variety has not previously been recorded from Victoria. 
.Superspecies tesquobum. 
brachycome blackii G. L. Davis, 
Proc. Linn. Soc. N.S.W., lxxiii, 1948, 206. 
Central Australia: James Range, 3.1883, Kempe (MEL). 
Subgenus Metabrachycome G. L. Davis. 
Superspecies iberidifolia. 
BRACHYCOME IBERIDIFOLIA Benth., 
Enum. PL Hueg. (1837), 59. 
South Australia: Mt. Eba, 1880, E. Giles (MEL) ; between the Alberga R. and Mt. Olga, 1873, 
Giles (MEL). 
BRACHYCOME TATEI J. M. Black, 
Proc. Roy. Soc. S.A., lii (1928), 227. 
Western Australia: Eucla, 1SS9, J. D. Batt (MEL). 
Based on these specimens the following notes can be added to the redescription of 
this species (Davis, 1948). 
"Receptacle 2-5 mm. broad, 1 mm. high, conical, shallowly pitted." 
brachycome parvula Hook. f. var. a parvula (Hook, f . ) G. L. Davis, 
Proc. Linn. Soc. N.S.W., lxxiii, 1948, 212. 
Victoria: Lakes Entrance, 12.1878, D. Sullivan (MEL) ; Gippsland, 1882, Howitt (MEL) ; Nhill, 
St. Eloy D'Alton (MEL) ; Lower Glenelg R., 1891, Eckert (MEL). 
Tasmania: Cape Portland, 2.1885, Baudinet (MEL). 
This variety is now known to extend along the whole coast of Victoria and to occur 
also in the western district. In Tasmania the only definite record hitherto was from 
the Gordon River on the west coast. 
brachycome parvula Hook. f. var. /3 lissocarpa (J. M. Black) G. L. Davis, 
Proc. Linn. Soc. N.S.W., lxxiii, 3-4, 1948, 213. 
Victoria: Port Phillip, 1891, J. G. Luchmann (MEL); Ballarat, 1887, G. Day (MEL). 
A specimen was examined from the Wimmera district (1893, W. E. Matthews, 
MEL), which in size and general appearance very closely resembled this variety 
except that the leaves were entire. 
brachycome pusilla Steetz, 
PI. Preiss, i, (1845), 426. 
Western Australia: Between Dundas Hills and Lake Lefroy, 1893, J. D. Batt (MEL). 
This constitutes the most easterly record of this species. 
BRACHYCOME EXILIS Sond., 
Linnaea, xxv (1852), 449. 
New South Wales: Between Bogan and Darling Rivers, 1877, L. Morton (MEL). 
Western Australia: Greenough R., 11.1877, P. Mueller (MEL). 
It would now appear that this species occurs in western New South Wales, and 
since there are now records from two widely separated localities in Western Australia 
(Israelite Bay and Greenough River) it is probably to be found also in intermediate 
areas. 
Superspecies trachycarpa. 
BRACHYCOME TRACHYCARPA F. Mliell., 
Linnaea, xxv, (1852), 339. 
Victoria: Werribee, Pullagher (MEL). 
South Australia: Great Bight, 1875, E. Giles (MEL); Eucla, 1889, J. D. Batt (MEL). 
152 REVISION OF THE GENUS BRACHYCOME CASS. PART III. 
Hitherto this species was only recorded from western Victoria, and in South 
Australia no farther west than Denial Bay. 
brachycome ciliaris (Labill.) Less. var. a ciliaris (Labill.) G. L. Davis, 
Proc. Linn. Soc. N.S.W., lxxiii, 1948, 221. 
Victoria: Snowy R., 2.1890, W. Bauer len (MEL). 
Tasmania: Near Swansea, 1882, A. Simson (MEL); Port Arthur, 1892, J. Button (MEL). 
From previous records this variety appeared to be confined in Victoria to the 
western districts and no specimens had been examined from Tasmania.. 
Superspecies ciliocarpa. 
brachycome ciliocarpa W. V. Fitzgerald, 
Jour. W.A. Nat. Hist. Soc, ii (1905), 23. 
New South Wales: Yandarlo via Wilcannia, 1886, B. Kenney (MEL); Cobar, 1S83, H. Andrae 
(MEL) ; Parkes, 22.9.1947, E. F. Constable (NSW). 
The known range of this species is now extended to western and central New 
South Wales. 
Superspecies silphiosperma. 
brachycome glandulosa (Steetz in Lehmann) Benth., 
Fl. Aust., iii (1866,) 521. 
Western Australia: Stirling Range, P. Mueller (MEL). 
No previous records exist of this species occurring in the coastal area, and although 
vegetatively similar to the inland specimens, the growth of some specimens is more 
luxuriant. The largest specimen reaches a height of 26-5 cm. and bears 44 inflorescences. 
The largest fruit examined was 4 mm. long and 3 mm. broad, but although exceeding 
the previously defined upper limits of size shows no structural variation. 
brachycome PERPusiLLA (Steetz) Benth. var. tenella (Turcz) G. L. Davis, 
Proc. Linn. Soc. N.S.W., lxxiii, 3-4, 1948, 231. 
Victoria: Werribee, Fullager (MEL). 
Western Australia: East of York, 8.1889, A. Eaton (MEL) ; Stirling Range, F. Mueller (MEL). 
These specimens extend the range of this species to the Port Phillip district of 
Victoria and to the south-western area of Western Australia, where it was only 
previously recorded from far inland. 
An atypical specimen was examined from Yorke Peninsula (1879, Tepper, MEL) in 
which a height of 5-5 cm. was attained and whose leaves were up to 2 cm. long and 
entire. In all other specimens examined the leaves were pinnatisect, but in view of 
the robust branching habit its affinities seem closest to this variety. 
Acknowledgements. 
My thanks are due to Mr. A. W. Jessep, Director of the National Herbarium, 
Melbourne, for forwarding a number of specimens for examination, to Dr. F. A. Rodway, 
Nowra, and Mr. N. A. Wakefield, Cann River, Victoria, both of whom placed their 
private herbaria at my disposal. 
I am also indebted to Miss Greta Baddams, of New England University College, 
Armidale, for writing the Latin descriptions of new species. 
All examinations and drawings of specimens were carried out with equipment 
provided by a Commonwealth Research grant. 
Reference. 
Davis, G. L. 1948. — Revision of the Genus Brachycome Cass. Part I. Australian Species. 
Proc. Linn. Soc. N.S.W., lxxiii, 142-241.