New Species and Combinations in Ceradenia (Grammitidaceae) Alan R. SmithUniversity Herbarium, University of California, Berkeley, California 94720, U.S.A.ABSTRACT. Ceradenia asthenophylla, C. aulaei-folia, and C. tristis are described from Mesoam-erica, and their relationships are discussed. Fifteenspecies of Ceradenia are now known from the re-gion; all are characterized by peculiar whitish glan-dular paraphyses and lack of hydathodes. Three newcombinations are also made in the genus for Andeanspecies: Ceradenia hishopii, Ceradenia intricata.and C. semiadnata. So that names will be available for the pterido-phyte volume of Flora ilesoamericana, three newspecies of Ceradenia are herein described. Thesewere recognized as new on specimens annotated byL. E. Bishop, who began a revision of the genus(Bishop, 1988, 1989) but was unable to completeit. In addition to the new species, three new com-binations are made for species occurring in SouthAmerica. These were also contained in an unpub-lished manuscript by Bishop.Ceradenia asthenophylla L. E. Bishop ex A. R. Smith, sp. nov. TYPE: Colombia. Huila: Cor-dillera Central, E slope between Paramo de las Papas and San Antonio, 2,900 m, Hishop 1984 (holotype, UC). Figure 11-L. Ceradeniae knightii affinis, a qua imprimis differt stip-itibus sparsim setosis (non dense setosis), laminis 6-10-plo longioribus quam latioribus (non 4-6-plo longioribusquam latioribus), laminis tenuis (non incrassitis) et tri-chomatibus numerosis ramosis glandulosis praeditis. Plants epiphytic; rhizome slender, with atropur-pureous scales 2-4 x 0.3-0.4 mm, with concol-orous setulae or glandular hairs; petiole brown, 24 cm x 0.2 0.4 mm, with a few scattered casta-neous setae to 3 mm distally and much more nu-merous simple or branched hairs throughout itslength, 0.25 0.35 times the length of the lamina,not flexed distally; rachis abaxially with sclerenchy-ma exposed, with scattered setae 1.5-3 mm; laminalinear, narrowed at base with 2-4 reduced pinnapairs, apex not seen, deeply pinnatifid, up to 15 cmlong; pinnae to 0.8 cm x 4 mm, entire or faintlysubrepand, set 70-800 to rachis, at base dilated onboth sides or straight acroscopically, apex broadlyrounded, with castaneous setae along margins andNOVON 3: 182-185. 1993.costae abaxially, a few scattered setae and numer-ous, branched, glandular hairs on lamina, costalsclerenchyma exposed; texture thin-herbaceous; soriinframedial to medial, up to 4 pairs per pinna, su-perficial, lacking setae but sporangia intermixed withnumerous, stalked, gland-tipped paraphyses. This species is named for its thin, rather lax leaves(from Greek asthenes, weak). It is most closelyrelated to C. knightii (Copeland) L. E. Bishop, whichhas densely setose petioles, lamina only 4 6 timeslonger than broad, and thick-herbaceous to char-taceous lamina lacking numerous, branched, glan-dular hairs abaxially. Ceradenia knightii is knownfrom Costa Rica, Colombia, and Hispaniola. Paratype. COSTA RICA. Cartago: Cerro de la Muerte, 1 km NW of Villa Mills on Interamerican Hwy., behind Hotel La Georgina, 2,900 in, Mickel 3206 (NY). Ceradenia aulaeifolia I. E. Bishop ex A. R. Smith, sp. nov. TYPE: Costa Rica. Limon: Cor-dillera de Talamanca, SW base of Cerro Ka-muk, in shrub-tree paramo, 3,200-3,350 m, Davidse, Herrera & Warner 25939 (holotype, UC; isotype, MO). Figure 1A H. Inter species subgeneris Filicipectinis frondibus pen-dentibus indeterminatibus et laminis perpinnatis ad C.mayoris et species affines accedens sed costarum abaxi-alium sclerenchymnate manifeste exposito, frondibus ma-joribus, pinnis margine valde sinuatis, laminis chartaceisab eis recedens. Plants epiphytic or lithophytic; rhizome stout,with castaneous scales 5-10 x 0.3-0.7 mm, withhyaline to concolorous setulae or white glands at themargin; petiole atropurpureous, 10-40 cm x 0.6-1.2 mm, at the base with a few to many castaneoussetae to 3 mm usually intermixed with much morenumerous short setulae 0.2 0.3 mm, sometimesglabrate distally, about equaling the lamina length,not flexed distally; rachis atropurpureous to blackish,abaxially with scattered hairs and occasionally a fewsetae, adaxially densely setose; lamina pendulous,lanceolate, usually narrowed at base with 1 4 pairsof reduced pinnae, lacking a distinct apical segmentbut with prolonged (indeterminate?) apical growth,perpinnate throughout, up to 50 cm (or more?) long:
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