Ianthopappus, a New Genus of the Tribe Mutisieae (Compositae) N. Roque Departamento de Ciencias Biol6gicas, Laborat6rio de Taxonomia Vegetal, Universidade Estadual de Feira de Santana, Bahia, Brazil D. J. N. HindThe Herbarium, Royal Botanic Gardens, Kew, Richmond, Surrey, TW9 3AE, United KingdomABSTRACT. The new genus lanthopappus Roque &D. J. N. Hind (Compositae, Mutisieae) is described,and a new combination, lanthopappus corymbosus(Lessing) Roque & D. J. N. Hind, is made. The singlespecies is easily distinguished by its subshrubby hab-it, discolorous leaves with actinodromous venation,corymbifonn inflorescences, and a 3-seriate pappuswith ca. 130 free, purple setae. Comparison is madebetween lanthopappus and other members of the Mu-tisieae that possess apiculate to acuminate apical an-ther appendages and are placed in a more narrowlydefined subtribe Gochnatiinae. lanthopappus is dis-tributed in southern South America from Brazil (RioGrande do Sul), Uruguay, and northern Argentina. Key words: Argentina, Brazil, Compositae, lan-thopappus, Mutisieae, South America, Uruguay. The generic position of Gochnatia corymbosa Les-sing (Compositae, tribe Mutisieae Cassini) has beendebated since its initial description (Lessing, 1830).Bentham (1873) considered that G. corymbosa, withits habit and distinctive capitulum, was better as-signed to the genus Onoseris Willdenow. This positionwas maintained by Baker (1884), but later contestedby Ferreyra (1944) in his revision of Onoseris. Fer-reyra (1944), after having examined Sellow 3479 (laterthe neotype of Gochnatia corymbosa assigned by Ca-brera, 1970), concluded that the species was not anOnoseris but probably a species of Gochnatia, the ge-nus in which it was initially described. Cabrera (1970)re-assessed the tentative generic position of this spe-cies and treated it as a species of Actinoseris (Endlich-er) Cabrera, transferring it from Gochnatia, basedprincipally on the heterogamous radiate capitulum. According to Bremer (1994), Gochnatia is thecrucial taxon in understanding the Mutisieae. InGochnatia, which is the largest genus of the tribe,all species have discoid homogamous capitula andthe apical anther appendages are acuminate toapiculate. Bremer (1994: 80) also stated that "anumber of genera apparently have their sistergroups within Gochnatia." The examples givenwere Actinoseris, Chucoa Cabrera, CnicothamnusGrisebach, Cyclolepis Gillies ex D. Don, Hyalis D.Don ex Hooker & Amott, and Nouelia Franchet.These are distinguished mainly by capitula char-acters and geographic distribution (Table 1). In the present paper, it is proposed that Actinoseriscorymbosa (Lessing) Cabrera is recognized as be-longing to a new genus, based upon our extensivemorphological studies. lanthopappus corymbosus ischaracterized by its striate leafy, silvery stem, thetruncate elliptic to orbicular leaves, which are dis-colorous, white-tomentose to glabrescent above, sil-very-sericeous beneath and have actinodromous ve-nation, the corymbiform inflorescences, and the3-seriate pappus, with ca. 130 purple setae (Table2). The new genus is described below.lanthopappus Roque & D. J. N. Hind, gen. nov. TYPE: lanthopappus corymbosus (Lessing) Roque & D. J. N. Hind. Figure 1. Tribus Mutisieae. subtribus Mutisiinae Lessing. Suffrutex:caules omnino foliosi: folia e base valde 3-nervia inflores-centia laxe corymbosa, pauciceps vel multiceps; capitula ra-diata, heterogama, pedicellata, pedicellis bracteolatis: invo-lucrum 5-7 seriatum; flosculi purpurascentes, eis marginatisbilabiatis, feminina, eis centralibus profunde 5-lobatis etiambisexualibus: rami stylorum bifidi, appendices antherarumbasales caudatae, longe attenuatae, pilosae, apicales itemlonge attenuatae; achenia longe denseque setulifera: pappus3-serialis, purpurascens, setis apice dilatatis.lanthopappus corymbosus (Lessing) Roque & D. J. N. Hind, comb. nov. Basionym: Gochnatia cor-ymbosa Lessing, Linnaea 5: 263. 1830. Onoseris corymbosa (Lessing) Bentham & Hooker f., Gen. pl. 2(1): 487. 1873. Actinoseris corymbosa (Les-sing) Cabrera, Bol. Soc. Argent. Bot. 13: 52. 1970. TYPE: Brazil. "Sellow in Brasilia meri-dionali (v.sp.s.oo)." Syntypes: B destroyed. Sellow 3479 (neotype, designated by Cabrera (1970), K; isoneotype, P (designated here), photo FI, NY, US). [Following Herter & Rambo (1953), Sellow 3479 was collected in January 1826 between Sao Gabriel and Ibicuf in the state of Rio Grande do Sul, Brazil.] Figure 1. NOVON 11: 97-101. 2001.
BioStor
