Ardisia callejasii (Myrsinaceae): A New Species from the Antioquian Choc6 of Colombia John J. Pipoly IllMissouri Botanical Garden, P.O. Box 299, St. Louis, Missouri 63166-0299, U.S.A.ABSTRACT. Continuing studies in Myrsinaceae forFlora de Colombia revealed a heretofore unde-scribed species of Ardisia closely related to Ardisiapittieri Mez of Costa Rica and Panama. While determining specimens in Colombian her-baria for my treatment of the Myrsinaceae for Florade Colombia, I discovered a fine series of specimensrepresenting an unknown taxon, described herewith.Ardisia callejasii Pipoly, sp. nov. TYPE: Colom-bia. Antioquia: Mpio. Mutata, N of Hacienda El Darien, right margin of Rio Chontadural, 28 July 1978 (fl, fr), R. Fonnegra & F. Renteria 953 (holotype, HUA; isotypes, COL, MEIEL, MO). Figure 1. Quoad ramulos furfuraceo-lepidotos, folia magna cor-iacea oblanceolata brevipetiolata, panicularum ramuloscorymbosos, stamina in tubum connata, ad 4. pittierivalde arcte affinis, sed ab ea laminis ad apicem longi-attenuatis (non acutis), sepalis chartaceis (non coriaceis)2-2.8 (nec 3-3.5) mm longis, symmetricis (nec asytm-metricis) ad apices obtusis (nec emarginatis), petalis ovatis(nec ellipticis), 5.3-5.9 (nec 10-11) mm longis, manifeste(nec obscure) punctatis, staminibus 4.9-5.4 (non 8.79.1) mm longis, tubo staminali 1.2-1.3 (nec 1.8-2) mimlongo, antheris ad basem inter se liberis (nec conniven-tibus), denique fructibus luteis (non ruberis) statim separ-abilis. Tree to 7 m; branchlets terete, 0.7-1 cm diamn.,densely and minutely furfuraceous lepidote. Leavescoriaceous, oblanceolate, (25-)31-45.5 cm long,(5-)7-12.2 cm wide, apex long-attenuate, graduallytapering to a cuneate base, costa slightly raisedabove, prominently raised below, secondary veins16-20 pairs, slightly depressed or not visible above,prominently raised below, glabrous and dull above,sordid and sparsely and minutely sessile rufous fur-furaceous-lepidote below, glabrescent, the marginregular, entire, flat; petioles marginate (0.6-)1-1.2cm long, glabrous above, densely and minutely ru-fous furfuraceous-lepidote below. Inflorescence sub-terminal, pinnately or bipinnately paniculate, (7.5)12.5-24.5 cm long, pyramidal, primary and sec-ondary bracts unknown, secondary branches 3-12cm long, the rachis minutely densely furfuraceous-lepidote, glabrescent; floral bracts unknown (pre-sumably early caducous); pedicels cylindrical, 0.71.8 cm long, densely furfuraceous-lepidote, persis-tent. Flowers 5-merous, corymbose, the corymb with7-12 flowers, pink; sepals quincuncial, almost free,chartaceous, ovate, 2-2.8 mm long, 1 1.3 mmnwide. symmetrical, apex obtuse, thickened andprominently black punctate medially, densely andminutely furfuraceous-lepidote, the margin irregu-lar, scarious, hyaline, sparsely glandular-ciliolate,entire; petals quincuncial, basally connivent, cori-aceous, ovate, 5.3-5.9 mm long, apex acute, highlyreflexed in anthesis, medially thickened, prominentlypellucid punctate without, somewhat verruculosermedially within, the margin scarious, hyaline, entire,glabrous; stamens 4.9-5.4 mm long, the filamentsconnate into a coriaceous elobate tube 1.2-1.3 mmlong, epunctate, glabrous, the anthers free, lanceo-late, 3.5-4.2 mm long, 1.3-1.6 mm wide, sessileon the staminal tube, apex attenuate, base cordate,longitudinally dehiscent by narrow, sublatrorse slits,the connective inconspicuously pellucid punctate;pistil obclaviform, 5.3-5.6 mm long, 1 1.2 mmdiam., the ovary 1.5 1.7 mm long, glabrous, thestyle 3.8-4 mm long, erect, inconspicuously pellucidpunctate, glabrous, the placenta ellipsoid, 0.8-1 mmlong, 0.3-0.4 mm diam., apex apiculate, the ovules12 15, pluriseriate, stigma punctiform. Fruit glo-bose, ca. 1 cm diam., green, then yellow at maturity,densely and prominently black punctate, the stylebase persistent. Distribution. ,4rdisia callejasii is endemic to theMunicipality of Mutata, located in the northwesterncorner of the Department of Antioquia, Colombia,at 100 180 m elevation. Ecology and Conservation Status. This new spe-cies occurs in lowland pluvial forests, in a narrowvalley belonging to the Rio Sucio drainage basin,located in the Department of Antioquia between thesouthwestern corner of the Department of Cordobaand the northwestern corner of the Department ofChoco. Label data from the Fonnegra et al. collec-tions indicate it is a common species in forests pre-served on Hacienda El Darien. However, becausethe region is traversed by the Medellin-Turbo high-NovoN 2: 389-391. 1992.
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