New Species and Nomenclatural Notes in Mesoamerican Ardisia (Myrsinaceae) John J. Pipoly II/ Fairchild Tropical (arden, I 1935 )ld Cutler Road, Coral (ables (Miami), Florida 33156-4299. U.S.A. jpipolyCifairchilgardten.org Jon M. RicketsonMissouri Botanical Gartlde. P.O. Box 299. St. Louis. Missouri 63166-0299. U.S.A. j on.ricketso)
[email protected] the Flora ' levOt(ne'ricatit( treatiimentet resulted inthel dis(coery of1 three new species from Panama.which are descriIbed and illustrated: Ardisia iherrer-InII Pipohl & Iicketson of subigenus Anrdisii. and1. nanoa Pipoly 1 lickltson amil 1. pfse.udcrus/dpiPipoly i Ilicketsoni, bolh of s lubgenus Icmaorec. Inadtlition. two new c(ombinations within |rdisiastl i. I/(corH'e are mt llad: Ardisi. n.xictan( snl)sJ.silth''cala (I.unndell) Pipoly & licketson and 1r-d(lisia fir/firacella subsI). reratgn.wensis (Lundlell) 1i-poly & Iticketsoin, the latter witlh two heterotpiicI)imoinials nri\l\ relegateil to syinonyiv (4. oerscte-diani I.undell anti A. <(r(foti/ali. I.unhdell). Final-Iv. we lectoihpil ird(isia conmpress. Kunlh var. mex-ilain(i Oerstedl. i new ltaonomic svnon11\ 1. andplace A. jldis1'1nsis lIundell into sIynonyIIy of 1.mnexictana Lundell suhsp. mnexicant.{':I MXl :N. El estluio taxondtillc o dte la familiaMyrsinaceac pmra el p)roiecto Flora Mesotunericainiresult5 'n el l(scnubrimiento le tires nnivas espe-cies lpanImieras lasI cnials se 4lesc(rillen e islustran:Ardisia herrerana I'ipoly & Rticketson del sulbge-nerl Anrdisii 1. nintlna Piploly & IKicketson l A. )se.u-(docusliddat Pipoly I Iicketson, almb)as del sul)g'-Ilnro Ictrcorea. Adll'rmis, sc proponen dlosc<'ominlaciois ncltvas dceitro de Arilisin sub g. /c( -corea: Ardlisim mexicrana suhl)s. sil/ternatm (lann-tdll) Pi"poly & iticketson y Irdlisida ij/iitracellaIsulspl. rerLagusensis (Lrulell) Pipoly 1 & Iickelson.y crm1o novedadl se relega a la sinonimia dc< la til-tima subespetcie a dos linoriios hIetrotflpios (L1.oferstedinda Ilntldll y A. candmf(/dii/l Iunlell). Fin-almentc, se ctloltipifica Irdisid c'nl/r.ess.t Kunthl\ar. n?'x'icnlt ()Oersted. IIn nIul1c 1 sinonimo1 taxon-omico. y, se ulica a i.I. jliscen.sis l.uindell coii si-noniino d� e A. mrexicana ILundell. Ke )uord.s: Irdi/.si. lrdisil subg. rdisia. Ar-Nol)ON 15: 190-201. 2005.(lisia sublg. I((iorcaV. conservation assessment. F'lo-r(i :'leso.arnericana. le(ctotvpilications. Mexico. Mvr-sinaceae. Paatlina. 1The I'anltroical 'gels ltrdifsi Swarlz is the larg-est in the fa1unil Mvrsinaceae. containing Iperhapsas many as 5()0 species (Chen & Pip)oly. 190). Itscircumscription has been problematic due to thelack Io a o lmprehensive treatmentl since that of Mez(1902). The genus has traditionally been separatedfront all others in the Mvrsinaeeae bI what he in-lerpreted as free filaments, and byv pluriserialeovules (Mez. 1902). Pipll and Ricketson 199i)8)fil(nd tlha the stamens in all sulbgenera are actuallyconnate hasally b) their filaments to form a hyaline.inconspicuous tube. but tlhat the tube is free fromthe corolla. The hyaline staminal tube. free fromnthe corolla tube. is the only unique characterislicthat distinguishes \rdisia from the other genera ofthe family, and thus defines it. While preplaring the taxonomic rieatment of Ar-disia for inclusion in the Mvrsinaceae for F/onr Me-souneiri(cti1. one new species was discovered with-in Arda sub. isi, a1ml two addlitional onesfrom subgenus Ic((or'ea (Aubilet) Mez. Also, twonew combinations were found to be needed, andthree binomials are relegated to synonymy. Finally.while working tiup tihe synonml for Irdisia nexi-(ca(t.( it was defler tilledI that lone of its newly pro-Iposed taixonomic( synlonyllms, Al . colnmpresst' var. 4 n'x-ic.ar, needed to be lectotypified.Ardisia Swartz sublg. Ardisia Ardisia subg. Ardisia. as will be circumscribedin thie lpcomling /lora ll(es(lrnlritcarn treatmenll.comprises I 1 species in Mesoarnerica. Members ofthis group are mostly locally common, occur in for-est margins anl survive moderate levels of distur-bance (Pipoly. unpulblished). and their ripe black
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