Rahowardiana globifera (Solanaceae), a New Species from Colombia Sandra Knapp The Natural History Museum, Cromwell Road, London, SW7 5BD, United Kingdom William G. D'Arcy Missouri Botanical Garden, P.O. Box 299, St. Louis, Missouri 63166-0299, U.S.A.ABSTRACT. Rahowardiana globifera, newly de-scribed from Antioquia, Colombia, is a second spe-cies for this striking, high-climbing solanaceous ge-nus. Rahowardiana is a small group of relatively rareepiphytic shrubs first described from Panama(D'Arcy, 1973: 670). It is a member of tribe Juan-ulloeae, whose members are all neotropical woodyepiphytes. Rahowardiana is most closely related toJuanulloa, sharing with that genus narrow tubularflowers and highly branched inflorescences. Workin progress by V. Persson, S. Knapp & S. Blackmoresuggests the genera share several pollen characters.Rahowardiana is distinctive, however, in its con-gested globose inflorescence with highly contrastingcorollas and calyces.Rahowardiana globifera Knapp & D'Arcy, sp. nov. TYPE: Colombia. Antioquia: Parque Na-cional Natural "Las Orquideas," Sector Ven-ados, 890 m, 6033'N, 76019'W, 1 Apr. 1988 (fl, fr), Cogollo, Ramirez & Alvarez 2895 (holotype, JAUM-018167; isotypes, COL not seen, FMB not seen, MO). Figure 1. Frutex scandens, foliis obovatis coriaceis magnis pe-tiolatis. Inflorescentia congesta globosa, pedicellis brevis.Flores calycibus tubulosis, coriaceis purpurea, corollis al-bis, antheris linearis. Bacca anguste turbinata. High-climbing, unarmed, scandent shrub, twigsstout, drying strongly angled; pubescence (seen onlyon leaf undersides) of slender, erect, weak, collaps-ing, simple, uniseriate hairs 3-5 cells long, the distalcells smaller. Leaves obovate, 15-30 x 10-21cm, apically rounded, basally obtuse, veins obscureabove, prominently elevated beneath, the lateral veins4 on each side, looping and anastomosing 5-10 mmfrom the margin, the minor venation mostly per-pendicular, the margins slightly revolute, subcor-iaceous, concolorous, glabrous above, beneath softlyand evenly pubescent; petioles drying dark, 15-25mm long, ca. 5 mm thick, drying longitudinallyridged and crossed with numerous transverse fis-sures. Inflorescence a condensed, crowded, globosepanicle or raceme ca. 15 cm long, forming spheresca. 35 cm diam.; peduncle not evident, pedicelsstout, ca. 5 mm long, glabrous. Flowers with calyxpurple, tubular, strongly angled, 7-8 cm long, ca.15 mm wide apically, coriaceous, irregularly sinuatelobed, the tips of the lobes green; corolla white, thetips of the lobes purple dorsally, ca. 6.5 cm long(immature), the lobes 0.5-1 cm long; stamens 5,the filaments straight, glabrous, inserted at or nearthe base of the corolla tube, anthers linear, ca. 15mm long, 2-3 mm wide, yellow, the connectivedrying dark, the thecae confluent at the apex, de-hiscing longitudinally; ovary pale brown, conical, ca.1 cm long, narrowly beaked, glabrous, surmountinga distinct, slightly lighter colored, slightly undulatingannular nectary, style white, markedly different incolor from the ovary, straight, 5-6 cm long, stigmastrongly bifurcate, the lobes ca. 1 mm long, minutelypapillate. Fruit a pale yellow, narrowly ovoid berry,ca. 9.5 cm long, basally 1 cm wide, expanding toca. 2 cm wide about 4 mm from the base, thennarrowing into a slender beak 4-5 cm long; seedfabiform, ca. 6 x 3.5 mm, tan colored, the surfaceminutely pitted. Rahowardiana globifera resembles Rahowar-diana wardiana D'Arcy, from Panama. It differsin its stouter twigs; larger, more coriaceous leaves,which are puberulent beneath; in its short pedicels;larger calyces; white instead of yellow flowers; andlarger fruit. The inflorescences of this species aretwice or three times as large as those of R. war-diana. Recent collecting in northwestern Colombia hasuncovered many interesting new plants, one of whichis described here. The collecting area, Parque Na-cional "Las Orquideas," is on the Pacific slope ofthe Western Cordillera, an area of high rainfall andhigh relief. This species was found in premontanerainforest, climbing in the overstory within hand'sreach. It is locally abundant, and the large, unusualinflorescences can be seen at a distance (Cogollo,pers. comm.). NovoN 3: 429-430. 1993.