New Sections, Combinations, and Varieties in Rosaceae, Potentilleae Barblara Erter Curator of Western North American Botany. University and Jepson Herharia, University of California, 1001 Valley Life Science Bldg. 2456, Berkeley, California 94720-2465, U.S.A., and Research Associate, Snake River Plains Herbarium, Boise State University, Boise, Idaho 83725-1515, U.S.A.
[email protected] James L. Reveal Adjunct Professor, Department of Plant Biology, 228 Plant Science Building, CornellUniversity, Ithaca, New York 14853-4301, U.S.A., Honorary Curator, The New York Botanical Garden, Bronx, New York 10458, U.S.A., and Professor Emeritus, University of Maryland, College Park, Maryland 20742-5815, U.S.A.
[email protected]S-THc. Eight new sectional names are pr11oosc1 lin Rosaceae, tribe Potentilleae: Horkelia Chamisso &Schlechtendal sect. Capitatae (Rydberg) Ertler &Reveal, section Hispidulae Ertter & Reveal. antisection Parrue Eriter & Reveal; Ivesia Torrey & A\.(ray sect. Comnre/ll (Rydberg) Ertter & Reveal.section Sax.osae (Rydberg) Ertler & Reveal, seclionStellariopsis (Baillon) Ertter & Reveal, and sectionUInguiculeate (Ryldberg) Ertter & Reveal; and Poten-till/ L. sect. Hipplianae (Rydlerg) Ertter & Reveal.Thirteen varietial clmbiinations are proposed: Horke-lia californica Chamisso & Schlecbtendal var. relta(Creene) Ertter & Reveal and variety fi'ondosui((reene) Ertter & Reveal; H. clevelandii ((reene)lRydberg var. brevibracteata (Wiggins) Ertter & Re-veal; H. cuneatta Iindley var. puberula (Rydl erg)Ertter & Reveal and variety sericea (A. Gray) Ertler &Reveal; H. daucifolia (Greene) Rydberg var. cnarulilii((Rydberg ex Howell) Ertter & Reveal and varietyindicta (Jepson) Eiter & Reveal; H. /usca LIindley var.brownii (Rlydberg) Ertter & Reveal (linked tosignificantly changed circumscriptions of varietypseudocapitlit (Rydberg ex Howell) M. Peck anlvariety capitat (Lindley) M. Peck); H. tridentainTorrey var. flavescens (Rydberg) Ertter & Reveal: andIvesia gordonii (Hooker) Torrey & A. Gray var.alpicola (Rydlherg ex Howell) Ertler & Reveal andlursinorum (Jepson) Ertter & Reveal; and I. lycopo-dioides A. Gray var. megalopetala (Rydllerg) Ertter &Reveal and variety scandularis (Rydberg) Ertter &Reveal. Two new varieties are proposed in Iresia: I.argyrocoma (Rydberg) Rydberg var. moranii Ertter &Reveal, a taxon confined to the Sierra San PedroMdrtir of northern Baja California. Mexico, and /.gordonii var. wiasatchensis N. H. Holmgren ex Ertter &Reveal of Idaho, Utah, Montana, aml WIyoming. A keyto the four varieties of 1. gordonii is presented, anda lectotype is designated for Ilorkelit gordonii. var.(lpicola Rydberg ex Howell. Key words: Comarella. Horkelit, Iresi.a Polentil-l/. Polentilleae, Rosaceae, Stellalriopsis. New sections, combinations, and varieties, pre-sented below, are necessary for a treatment of variousgenera of Rosaceae Jussieu tribe Potentilleae Sweet ina forthcoming volume of Florna of North Americta Northof/ Mexico (FNA). In our treatment, both HorkeliaChamisso & Schletchendal andl Iresia Torrey & A.Gray are considered to be dislinct from Potentilla I,.following Keck (1938) with some modificationsrecently adopted by Ertter (1993) and Holmgren(1997), even though this results in a paraphyleticPolentilla in a strict sense. In the previous compre-hensive treatments of Potentilleae in North America,Rydberg (1898) initially maintained Ivesia in Horkeliaonly to later accept both genera as distinct (Rydberg,1908). He also recognized some entities (ComoarellaRydberg and Stellariopsis Rydberg) at the genericlevel that Keck subsequently submerged into Iesi'sa.Some of the numerous unranked names proposed byRydberg above the rank of species have been formallyestablished at the rank of secttion by subsequentauthors (e.g., Johnston, 1985); several more arelikewise established herein. Keys and full descriptions will be available in theFlora and are not repeated here (except for a key tothe varieties of Ihesia gordonii (looker) Torrey & A.Gray). The distribution statements given here in somedetail are based on our review of taxa for the FNAtreatment and, in some cases, as noted, are novel.NovoN 17: 315-325. Pt IStt.tSED ON 10 SEItt'EMBEK 2007.
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