New Species of Sphaeratldenia (Cyclanthaceae) from Costa Rica and Ecuador Roger EriA.vson D)eparltment of Plant and Environimtental Sciences, Gioteborg University. Box 461,SE 4105 3() Gtiieborg. Sweden.Awlm l:r. TIwo new species of Splaurndeniri liarling((Cyclanthaceac), .. rsle'/otal H. Eriksson frIom CostaIic atnl .5. mtarr.escens. H. Eriksslon fro, Ii lEcuador, l redescribed andl illustraledl . S/li<aeril'deinii rosllll//in isdistinguished fron m the closely relalted ,'. iprt lerm'! nis. Ii .I'riksson and S. occieiliinlis R. Eriksson L the wvidelbhsepairiatld, uin/itile sligiImas. Spi haruIi e.i1 in rc10 ,en(s Idiffers fronm the supposdlly closely related S. pnilida [{.Iriksson by the persislinlg slaminuate flowers.I,]�siMKN. Se descriiben . iluslrin <los espe|ciesinlllvas (ie .ip>lhn. eruId' ni t llarling ((:ycl' ulaeli'' ). S.rioste/lllni I{. I'rikssotn Icl Costa liga \ .. m,, rtitr 'll tts H.Krikssonl de I'lcullador. .Spht<<erdnl'iii rosle//tl sedistingue de las tisp. ci ss ('sirechaliiet'll relacionadas..5. prl h'lcriniss. IR. I'riiksson v . cci' lerntali. Ia .I'rikssol i, ior Ns t's lignias iiin s1epairadc s \y Illlcinalos.Sphl er, n(l('r ' Iii<i trl.rcc.cens" se dlifer'iie ia d<t la cspecicepllalllll ll nte mins relacionada, S. p, llida I\. Eriksson.p ir las ilorrs eslam inadals i)ersistel ts. cv ior(is: C(osa Iinca. Cv('li lhaeac,. c'(':ua or. 'The genuis Sl)hae'r(i(l'ni<a Harling ((Cvclanllhacae) isclharacterized ly). e.g.. <listichlius lea\c s. pislillateIloer\cs wilh onIe apical placenia. and anelicrs oftenhiaving an apical secrletion globule. I'uiirtlhennore, thefriiliiing spadi s, are olti'n blrightll colo rd, in contrasttio iilny ollther 'ycvlanilis. I is disribuiiIt Ifronisoutll 'rn Nicaraguall iit ort1-hlrn Inlivia, exiltelilneastward into Venezulca Iand adjacent Brazil, and iscolnnonly foundl in lhumid habilalts froi sea level upto ( . 3(00 1 I altitude. or in somelill areas even higher.Sph/ler<'nileni was recentlly revised (l'riksson. 1995).witih 5() recognized species of hllichl 20 weredescribed ihen as ntw. 'lheire are obvionsl\ newSphl er(d< /<'iii sl'ucies tIo he dliscoverled, and Iwo of.lliem are diescrilbed lie l'. Measuremnents were taken on dried materialaccording to Kriksson (1995: fig. I). excepl thatimedian and latl'eral Mloral strucltrets \were not sepa-ralely measured. "hloral and ifruiting slrlictures. and tosome extent vegethativ'e parts, lmay shrink conlsiderablyduring drying.lrlgel.(lison kiiOt(9(J)(s.g l~st.SllIhalradc(ilia roitellala H. Eriksson. slI. nov. T >'P: C(osa tica. Liniuinn: Cantin de Tila-manail. (Cordillera de Talamallliic. Cerro l Coo, l>etw. I jiarrs & Satn .los( Cab(lliltl, ) 2 '23'"N. 83 13' I()"\\. 1500 il. 19 lMar. 1993.1. Frnmin-i (/e. 711 (hololtype. INH: isolylUe. CI noit sieen). I igure 1 A-C. I11 11h 'piii ll tica v tIierreslris. I antiliiu 7 -HI '1 ci nlilng , se nlI iilis t10-I I m lalis,. lainni latis, istai 21-28 iii linngi : pri liit s l(t ad plus qualin 8t 5 -tii ol iilns; I llt si loeii . in sialu Iru'tifi rn runbra,. in sliut floriftro circiii r 1.5 Ic t liait tro. in sli u fIruclif'ero jIlt ni -1..5 nll linta. 1.t 8 rin iliametro in shlill fruclifenit, Ii ir)o l0.3 i 'i Elnga. 2.1 ('tit diamcn o. Fhlores ntscili caducei. 3.,-I nfin longi: antlhrn, ).6-1).8 innt lh gaue. ).5 nn lilac. ghobis s iirr lionis us(ltii t l .1 111111nI iliain2 rt . 2 l orcs l it ini t stylus 2.')-7 n1n lIongus: stigiala l.0-2.1 nit lonih , (1.1-0. untarn Ila, inter apices dlistalcs 3.8-6.6 1111. sti nmnali 'desup r 8isa lin L ria. In inalll . 1i. stric l) \i'l l. rsuni i ri'11vg s. ;i la 1 l illler co la il a I li tlrn i el, I slith I tru li'rl lai sI i alal. i u llnihil ilnfl riorni all altior. 1111 Il l;i le a ll. 'unnan ob>loiga v: l anguisl, ellipsoitlh,. 'piphylitle or terreslrial plant. Leal blades 76-tl ai. tifid 68%6-650 ofI Ihir lenAligh, segiites 10-11 i i i broad, lanceola.te. acnl inilla , iut dge diistance12-15 ) inll. abasxial ridges willh low papillae (soni -liinis almosi smlootlh). miitlrib 2l -28 mi: peliolc 1,(0ior tllian 85 cIn; slieatlh ;33-ci1 . t Pe'ldll.~ h l 21-31 cm, straight: spalltis ;3. whilte tinged withli red.longest one 11 cil. S adlix cylindrical to narrowlyellipsoilld, in fruit red, 1t lntlhesis ca. 1.5 eiti broad, inyoiung fr il 1.5 X I.8 c11. in Iatl'ure fr il :.;3 X2. 1 11111 . Slaliin ale f lowers 'adlliU l.'ous. 3;.7-I1a il1edicel 2.( -3.1 t i nii . rii ecet pt<iacl 2.1-2,2 tII l liroad.� 1lai: ieriantlih lobles 11 to 12. includling 5 to 6ad1xial ones, up to I. I X (). Im ; stainens 13 Io 22.antilhers )0.i-0.8 X 0.5 nmn. secretion globules pale,to 0.05 uni <liamn.. fI'ilaen nts 0.2-0.3 liiun. basalbulbs ().3 X 0.3 m1il. �+ globose. Pistillate flowersand fruits 2.8-7 X 2.7-1.1 nil; tepals 2.8-5.1 uii.1.5-3 unii broad, ).-1 nunll Ihick. free. oblong Itonarrowly\ Irianiigular. + acilte. alex �+ straightl, upperp)all 1 'lepauls incurved al antlihesis. benti outwardi infruitl, iiiargins straight to incurved: style 2.5-7 X 1-No) tN 17: -16-59. t I.lisml:l ()It 20 ,ItM: 2007.
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