New Taxa and Nomenclatural Changes in Allophyllum, Gilia, and Navarretia (Polemoniaceae) Alva G. DayDepartment of Botany, California Academy of Sciences, Golden Gate Park, San Francisco, California 94118-4599, U.S.A.ABSTRACT. Studies of certain species problems inAllophyllum, Gilia, and Navarretia resulted in thefollowing nomenclatural changes: Gilia subg. Kel-loggia is reduced to sectional status, two new speciesand one new subspecies are described (Gilia lottiae,Navarretia myersii, N. hamata subsp. parviloba),two subspecies are elevated to species (Navarretiajaredii, N. rosulata), one variety is elevated tosubspecies (N. nigelliformis subsp. radians), andeight new combinations are proposed (Allophyllumgilioides subsp. violaceum, Gilia brecciarum subsp.jacens, G. sinistra subsp. pinnatisecta, avrarretialeucocephala subsp. hakeri, N. leucocephala subsp.minima, N. leucocephala subsp. pauciflora, N. leu-cocephala subsp. plicantha, and N. intertexta subsp.propinqua). These changes are included in treatments of thesame genera contributed by the author to the JepsonManual: Higher P'lants of (California (Hickman,1993). This report is presented in order to validate no-menclatural changes found necessary during prep-aration of treatments of Alloph yllum, Gilia, andNavarretia for the Jepson Manual: Higher Plantsof California (Hickman, 1993). A more detailedaccount of studies that led to some of the decisionswill be forthcoming elsewhere.ALLOPIlILLUM (NUTTAIL) V. GRANT Allophyllum is a small genus occurring in themountains of the western United States. All speciesare annual, the sterns leafy, corollas short-or long-tubed, and blue, purple, pink, or white. Althoughsegregated from Gilia by Grant & Grant (1955), itwas maintained as a section of Gilia by Cronquist(1984). Characters distinguishing Allophyllum from Gi-lia are its nonscapose habit, leafier sterns, and dis-tinctive trichome type (fine-stalked, gland-tipped),seeds generally boat-shaped, and leaf-lobes thick-ened at the tip, not fine-pointed as in Gilia. Illophyllum violaceum (Heller) A. & V. Grantanld .. gilioides (Bentham) A. & V. Grant areclosely related species, with the former occurringgenerally at higher elevations. Typical A. violaceumoccurs from 2,000 to 2,900 m in the high SierraNevada of California and in the mountains of westernNevada. The plants are small (generally 6 12 cmtall), have few-lobed leaves, and 2-3 flowers percluster. Allophyllum gilioides occurs at lower el-evations, mostly below 2,000 m; the plants are larg-er (14-60 cm tall), with larger, many-lobed leaves,and a glomerate inflorescence, 4-8 flowers per clus-ter. The two taxa intergrade in some areas at middleelevations (1,200-1,800 m) where their rangesoverlap, as in Kern Canyon in the southern SierraNevada. Since a clear distinction between them isnot possible throughout, I propose the following newcombination.Allophyllum gilioides (Bentham) A. & V. Grant subsp. violaceum (Heller) Day, comb. et stat. nov. Basionym: (ilia violacea Heller, Muhl. 1: 56. 1904. Allophyllum violaceum (Heller) A. & V. Grant, Aliso 3: 106. 1955. Gilia gilioides Bentham var. violacea (Heller) Cronq., Intermountain Flora 4: 132. 1984. TYPE: U.S.A. California: Nevada Co., lower end of Donner Lake, Heller 6873 (holotype, DS; iso-types, NY, UC).GILIA Ruiz & PAV)N The total number of species of Gilia sensu stricto(excluding Ipomopsis) is � 70, of which 9 are SouthAmerican. In North America the genus is confinedto the western states, with the greatest concentrationof species in California, which has 47 of the ap-proximately 60 species, as treated for the JepsonManual. In this work the species are grouped in sixsections: Arachnion, Gilia, Giliandra, Giliastrum,Kelloggia, and Saltugilia. This is modified fromthe sectional arrangement of Grant (1959), es-pecially in the addition of the new section Kelloggia. Two species groups were originally included inGilia sect. Saltugilia V. & A. Grant: the G. splen-dens group and the G. leptalea G. capillaris group(Grant & Grant, 1954). New studies show that G.NovON 3: 331-340. 1993.
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