Five New Species of Chusquea (Poaceae: Bambusoideae) and a New Combination Lynn G. Clark Department of Botany, Iowa State University, Ames, Iowa 50011, U.S.A.ABSTRACT. Five new species, Chusquea /iilcata.(. subulata, C. barbata, C. spathacea, and C.pulchella, are described and illustrated. One newcombination, Chusquea attenuata (Doell in Martius)I. G. Clark, is proposed for a Brazilian speciesoriginally described as an Arundinaria, based on avegetative specimen. During the course of floristic and phylogenetic studies in the neotropical woody bamboo genus Chusquea Kunth, I have discovered a number of species new to science, of which five are described here. A previously poorly known species of Arun-dinaria Michaux from Brazil is transferred to Chus-quea, and the new combination is made. Chusquea falcata L. G. Clark, sp. nov. TYPE: Ecuador. Loja: Parque Nacional Podocarpus, above Nudo de Cajanuma, wet montane forest around "Centro de Informaci6n," 2,800 3,000 m, 14-15 May 1988 (fl), H. 0llgaard, J. E. Madsen & L. Christensen 74116 (holotype, QCA; isotype, AAU). Figure 1A-D. Culmi ca. 1-1.5 cm diametro, 1.5-3(6-8) m alti; erectiad basim, interdum arcuati ad apicem. Folia culmorum18 32 cm longa, fragilia, glabra; vaginae 17-30 cmlongae, (7-)10-20-plo longiores quam lamina; laminae1.2 2.7 cm longae. Gemma centralis rotundata. Rami-ficatio infravaginalis; rami subsidiarii cujusquisque nodi30-60. Folia cujusquisque complementi 3-4; laminae7.8 20 cm longae, 0.4-0.85 cm latae, ratio long.:lat.= (11-)17.5-35, apice setosae, basi rotundato-attenu-atae. Paniculae (5-)7-16 cm longae, angustae. Spiculae(4.6 )5.1-6.5 mm longae, falcatae, nitidae. Glumae 2,squamiformes. Lemmata sterilia 2, mucronata. Lemmafertile 4.3-5.7 mm longum, mucronatum. Culms ca. 1-1.5 cm diam., 1.5-3(6-8) m tall,erect at base, scandent to arching above. Internodesterete, usually glabrous, sometimes scabrous belowthe nodes. Culm leaves 18-32 cm long, just sur-passing the next node, abaxially glabrous, brittle,eventually deciduous, juncture of sheath and bladeabaxially indistinguishable or nearly so; sheaths 17-30 cm long, (7 )10-20 times as long as the blade,the overlapping margin ciliate, not fused at the base;blades 1.2-2.7 cm long, triangular, erect, persistent,adaxially pubescent, apex subulate; girdle 1-3 mmNovoN 3: 228-238. 1993.wide, glabrous; inner ligule ca. 0.5 mm long, cili-olate, pubescent. Nodes at mid-culm with the cir-cular central bud subtended by numerous smallersubsidiary buds in a constellate arrangement; sheathscar dipping below the bud/branch complement;supranodal ridge a visible but not prominent line.Branching infravaginal; central branch sometimesdeveloping; leafy subsidiary branches 30-60 pernode, 11-27 cm long, sometimes rebranching fromthe base. Foliage leaves 3-4 per complement; sheathsglabrous, the overlapping margin ciliate, a tuft ofcilia present on each side of the sheath apex, ciliaca. 1 mm long; blades 7.8-20 cm long, 0.4-0.85cm wide, L:W = (11-)17.5-35, adaxially usuallyglabrous but sometimes hispid toward the base ofthe midrib, abaxially usually glabrous, sometimessparsely pilose, not tessellate or only weakly so, apexsetose, base rounded-attenuate; pseudopetiole 1.5-2 mm long, adaxially hispidulous, abaxially glabrous;outer ligule a short, usually ciliolate rim, 0.3-0.5mm long; inner ligule 0.5-0.6 mm long, truncate,abaxially pubescent. Panicles (5-)7-16 cm long,narrow, the base usually either retained within thesubtending sheath or just fully exserted; rachis -flattened at the base, angular above, glabrous, theedges scabrous, the basal bract 1.5 4 mm long,papery, pubescent, ciliate; branches 2-3 cm longat the base, angular, glabrous to scabrid, the lowerbranches ascending to appressed, the upper branch-es appressed; pedicels 1-4 mm long, angular, gla-brous to scabrid. Spikelets (4.6-)5.1-6.5 mm long,falcate, shiny. Glumes 2, scalelike, obtuse, abaxiallysparsely pubescent toward the apex; glume I 0.4-0.9 mm long, ca. '/ the spikelet length, 1-nerved;glume II 1-1.8 mm long, ca. ', the spikelet length,1 3-nerved. Sterile lemmas 2, mucronate, glabrous;sterile lemma 1 1.6-2.9 mm long, ca. / the spikeletlength, 3-nerved; sterile lemma II 2.7-4 mm long,ca. / the spikelet length, 3-or 5-nerved. Fertilelemma 4.3-5.7 mm long, mucronate, glabrous, 5-or 7-nerved. Palea 4.2-5.3 mm long, subequal tothe fertile lemma, bimucronulate, 4-nerved, sulcateonly toward the apex, the sulcus scabrous-hispid.Stamens 3; anthers 2.7 mm long. Lodicules 3; an-terior pair 2.1 mm long, posterior one 1.8 mm long,all apically ciliate. Fruit unknown.
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