NEW COMBINATIONS AND TAXA IN CUCURBITACEAE R. P. Wunderlin University of South Florida, Tampa, FL 33620 The following new combinations in Psiguria are proposed and new species described preliminary to a treatment of the family for the Flora of Panama. 1. Psiguria bignoniacea (Poepp. & Endl.) Wunderlin, comb. nov. BASIONYM: Anguria bignoniacea Poepp. & Endl., Nov. Gen. Sp. 2: 53. 1838. 2. Psiguria longipedunculata (Cogn.) Wunderlin, comb. nov. BASIONYM: Anguria longipedunculata Cogn., Diagn. Cucurb. fasc. 1, 21. 1876. 3. Psiguria warscewiczii (Hook, f.) Wunderlin, comb. nov. BASIONYM: Anguria warscewiczii Hook, f., Bot. Mag. t. 5304. 1862. 4. Cyclanthera dressleri Wunderlin, sp. nov. Herba scandens; foliis hastatis, 5-9 cm longis, basi 3-6 cm latis, apice acuminatis, marginibus denticulatis ; floribus staminatis in racemosis apice petiolorum productis; corollis albis, lobis 2-3 mm longis; floribus pistillatis solitariis; fructu viridibus, oblique ovoideis, setis ca. 1 mm longis. Herbaceous vines; stems slender, glabrous. Leaves simple, blade hastate, 5-9 cm long, 3-6 cm wide at base, margins dentic- ulate, apex acuminate, basal sinus acute, chartaceous, upper surface minutely pustulate, lower surface glabrous; petiole 1.5- 2.0 cm long; tendrils simple. Staminate flowers produced from the petiole at base of leaf blade, short racemose, 6- to 10-f lower- ed ; peduncle 1.5-2.0 cm long, slender; pedicels 5-8 mm long, fili- form; calyx shallowly cupulate, 3-5 mm wide, greenish-white, lobes obsolete; corolla ca. 6 mm in diameter, white or cream, lobes 2-3 mm long, broadly lanceolate or triangular, outer surface glabrous, inner surface slightly papillate, 3-nerved; androecial column ca. 1 mm long, head ca. 1 mm in diameter. Pistillate flowers solitary in same axils as staminate; peduncle 6-10 mm long; calyx and corolla as in staminate flowers, but slightly smaller; ovary obliquely ovoid, rostrate, ca. 4 mm long, short setose; stigma subglobose. Fruits greenish, obliquely ovoid, ca. 1.5 cm long, explosively dehiscent, seta ca. 1 mm long; seeds not seen. TYPE: Panama, Chiriqui: Cerro Colorado, 50 km N of San Felix on the continental divide. Cloud forest. Altitude 1200- 1500 meters. 17 August 1975. Mori & Dressier 7795 (MO, holotype; USF, isotype). 219 220 PHITOLOGIA Vol. 38, no. 3 This rare species is known only from the type locality. It resembles C. phyllantha Harms in having its staminate inflores- cence produced from the petiole at the base of the blade, but differs in having leaves hastate rather than 3-to 5-angulate. The species is named in honor of Dr. Robert Dressier, one of the collectors of the type. 5. Cayaponia sessilif lora Wunderlin, sp. nov. Suffrutex volubilis ; foliis ellipto-ovatis , 9-21 cm longis, 6-11 cm latis, apice acuminatis, basi truncatis, marginibus remote denticulatis ; floribus staminatis fasciculatis sessilibus vel subsessilibus ; calycibus dense pilosis; corollis viridibus, minutis; floribus pistillatis ignotis; fructu luteo-aurantiacis, ca. 3 mm in diametris, sparse pilosis. Suffruticose vines; stem deeply 5-sulcate, puberulent, often also sparsely pilose, glabrescent in age. Leaves elliptic-ovate, 9-21 cm long, 6-11 cm wide, apex acuminate, base truncate, margin remotely denticulate, subcoriaceous , upper surface sparsely pubes- cent on primary nerves, otherwise glabrous, finely reticulate- nerved, slightly bullate, lower surface pubescent, densely so on nerves, conspicuously reticulate-nerved; petiole 0.8-2.5 cm long, pubescent, often also sparsely pilose; tendrils simple or unequal- ly 2-branched, slender, pubescent, often also sparsely pilose. Staminate flowers axillary in sessile or subsessile, few-flowered clusters; calyx campanulate, ca. 3 mm long, outer surface densely brown-pilose, inner surface pilose near top, otherwise glabrous, lobes subulate, ca. 3 mm long, densely brown-pilose, spreading or reflexed; corolla greenish, triangular, 0.5 mm long, outer surface pilose, inner surface densely pubescent; filaments stout, ca. 0.5 mm long, pubescent at base, anthers complicate, forming an irreg- ular head 2.5-3.0 mm long and nearly as wide; pistillodium obso- lete. Pistillate flowers not seen. Fruits yellowish-orange, globose, ca. 3 cm in diameter, sparsely brown-pilose, peduncles ca. 5 mm long; seeds not seen. TYPE: Ecuador, Los Rios : Rio Palenque Biological Station, km 56 on road from Quevado to Santo Domingo. Elevation 150-220 meters. 16 August 1976. Dodson 6159 (SEL, holotype; MO, USF, isotypes) . This species is known from Ecuador and Panama. It differs from all other species of Cayaponia in having very small, sessile or subsessile flowers with a densely pilose calyx. 6. Melothria dulcis Wunderlin, sp. nov. Herba scandens; foliis cordatis, 8-14 cm longis, 6-10 cm latis, apice acuminatis, marginibus undulatis, remote denticulatis; floribus staminatis racemosis; corollis albis, lobis 3.0-3.5 mm longis, ref lexis untrinque pubescentibus ; floribus pistillatis solitariis; fructu luteis vel aurantiacis, 5-7 cm longis, 4.5-6.0 cm in diametris, glabris, exocarpis subligneis, mesocarpis aurant- iacis . 1978 Wunderlin, New combinations and taxa 221 Herbaceous vines; stems minutely scabrous, glabrate in age. Leaves cordate, 8-14 cm long, 6-10 cm wide, apex acuminate, base deeply cordate, margin undulate, remotely denticulate, charta- ceous, upper surface minutely white pustulate, slightly scabrous to glabrate, sparsely short strigose on nerves, lower surface glabrate; petioles 3-7 cm long, glabrate; tendrils simple, glab- rate. Staminate flowers axillary, racemose; peduncles ca. 2 cm long; calyx-tube narrowly campanulate, ca. 5 ram long, glabrous, 10-nerved, lobes triangular-lanceolate, ca. 1 mm long; corolla white, lobes oblong, 3.0-3.5 mm long, apex emarginate, pubescent on both surfaces, reflexed; anthers oblong, ca. 2.5-3.0 mm long, papillose at apex, ciliate on margins; pistillodium conical, glabrous. Pistillate flowers solitary in same axils as staminate flowers; peduncle 2-3 cm long; other details unknown. Fruits yellow to orange, ellipsoidal to subspherical, 5-7 cm long, 4.5- 6.0 cm in diameter, smooth, glabrous, exocarp subligneous, meso- carp orange, sweet; seeds white-sericeous, narrowly ovate, 9-10 mm long, 5-6 mm wide. TYPE: Panama, Canal Zone: Pipeline Road, 10 mi. from main gate. 14 August 1971. Croat 16693 (MO, holotype; MO, isotype). Melothria dulcis is endemic to Panama. It has been collect- ed in Panama, Darien, and Colon Provinces as well as the Canal Zone. It occurs in wet forests and along streams at 350-400 meters elevation. This species differs from other Neotropical Melothria species in its large fruit with a subligneous exocarp and orange flesh.