1983 Moldenke, Notes on Aeglphila 309 
ABQIPHILA VITELLINIFLORA var. BGLERI Mold. 
Additional bibliography: Mold., Phyi-ologia 27: 376. 1973; Mold., 
Phytol. Mem. 2: 138 & 522. I98O. 
ABGIPHILA WIGANDIOIDES Lundell 
Additional bibliography: Mold., Phytol. Mem. 2s 59 & 522. I98O} 
Mold., Phytologia li75 137. 1980. 
ADDITIONAL NOTES ON THE GENUS BOUCHEA . VII 
Harold N. Moldenke 
The last previous installment of these notes was published in 
Phytologia UO: 1^3 — ii23 (1978) . For a detailed explanation of the 
herbarium acronjrms used in this and all others in my series of 
papers, see Phytologia Memoirs 2: U63 — U69 (1980) with supplement 
in Phytologia 50: 268 (1982) . 
BOUCHEA Cham. 
Additional & emended bibliography: Sandmark in L., Amoen. Acad. 
5: 375. 1759} J. F. Gmel. in L., Syst. Nat., ed. 13, imp. 2, 2: ia. 
I79IJ Meian., PI. Vase. Gen. 2: I98. iSUOj Reichenb., Deutsch. Bot. 
[Repert. Herb. Nom.] IO8. iSUj D. Dietr., Syn. PI. 3: 370, 596, & 
605. 18U3; Lindl., Veget. Kingd. 661i. l8U6j A. L. Juss, in Orbigny, 
Diet. Univ. Hist. Nat. 13: 185. 181|9; Wight, Icon. PI. Orient, h 
(3): 10—11, pi. IU6I & IU62. 18U9J C. Muell. in Walp., Ann. Bot. 
Syst. 5 J 706. 1860| Eocq. in Baill., Adansonia, ser. 1 [Rec . Obs, 
Bot.] 2: 89, 110, 115, 12U, 125, 127, 128, 132, I3U, 1U3, « IU6— 
lliS (1862) and 3: 180—182, I8U, 185, 235— ?37, & 2Ul, pi. 16. 
1863} Lindl., Treas. Bot., imp. 1, 1: I6O. 1870} Pfeiffer, Nom. 
Bot, 2 (1): 133 & 759. I87U} Lindl., Treas. Bot., imp. 2, 1: 160 
(1876) and imp. 3, 1: I60. I88U} A. Gray, Synop. Fl. N. Am., ed. 2, 
2: 333—335. 1886} Durand, Ind. Gen. Phan. 320. 1888} Baill., Hist. 
PI. 11: 9U & 102 (1891) and 11: U88. 1892} Briq. in Engl. & Prantl, 
Nat. Pflanzenfam., ed. 1, h (3a): 133, 135, 137, IhO, 1U2— Ihl, lU9, 
& 153— l51i, fig. 59 A & B. 1895} Gtfrke in Engl., Pflanzerw. Ost- 
Afr. C: 338. 1895} Briq. in Engl. & Prantl, Nat. Pflanzenfam., ed. 
1, h (3a): (38I] . 1897} Lindl., Treas. Bot., imp. h, 1: I60. 1899} 
Millsp., Field Columb. Mus . Publ. Bot. 1: 523. 1902} Post & Kuntze, 
Lexicon 76 & 167. 190li} Reiche, Estud. Grit. Fl. Chile 5: 30U. 
1907} J. C. Willis, Diet. Flow. PI., ed. 3, 272. 1908} Urb., Symb. 
Antil. Us 532 — 533. 1911} Fedde & Schust., Justs Bot. Jahresber. 
39 (2): 319. 1913} Thonner, Flow. PI. Afr. Ii68. 1915} Fedde, Justs 
Bot. Jahresber. 39 (2): 132U. 1916} Sanzin, Anal. Soc. Cient. Ar- 
gent. 88: 106 & 133. 1919} J. C Willis, Diet. Flow. PI., ed. 5, 
90. 1925} J. Hutchins., Fam. Flow. PI., ed. 1, 1: 309 & 315. 1926} 
310 P H Y T L I A Vol. 52, No. 5 
Kobuski, Ann. Mo. Bot. Gard. 13: 16 & 23. 1926: Knuth, Feddes 
Repert. Spec. Nov. Beih. U3s [Init. Fl. Venez.J 603. 1927 J Chiov., 
ri. Somala 56, 60, & 27U. 1929} Ekman, Arkiv Bot. Stockh. 22A: $1, 
1929J A, W. Hill, Ind. Kefw. Suppl. 7: 260. 1929} Levyns, Guide Fl. 
Cane Penins. 216. 1929} Good & Exell, Journ. Bot. 68: Suppl. lliO. 
1930} Pio Correa & Pena, Dice. PI. Uteis Bras. 3: 39li & 395. 
1931} Marloth, Fl. S. Afr. 3: 1U6. 1932} A. Ghev., Rev. Bot. Appl. 
Agric. Trop. 15 : 913. 1935} Mold., Suppl. List Comm. Vem. Names 
1 & 15. 19iiO} Worsdell, Ind. Lond. Suppl. 1j 132. 19la} Lemfie, 
Diet. Descrip. Syn, Gen. PI. Phan. 8b: 653. 191^3} Srdtman, Svensk 
Bot. Tidsk. 39: 281— 28U. 19U5} H. N. & A. L. Mold., PI. Life 2: 
20, 22— 2li, 28, 30, 33, 5l, 56—58, 62, 67, 7U, 80, 81, 83, 89, & 
9h. 19U8} Kidd, Wild Fls. Cape Penins. viii, pi. [UO], fif> 6. 
1950} Metcalfe & Chalk, Anat. Dicot. 2: 1031, 1032, & lOUO. 1950} 
Lawrence, Taxon. Vase. PI., imp. 1, 687 & 782. 1951} J. C. V/illis, 
Diet. Flow. PI., ed. 6, 90. 1951} Alain in Le6n & Alain, Fl. Cuba, 
imp. 1, U: 279 & 29U— 295, fig. 127. 1957} Dalla Torre & Harms, 
Gen. Siphonog., imp. 2, 1*30. 1958} Rickett & Stafleu, Taxon 9: 8U. 
I960} Dalla Torre & Harms, Gen. Siphonog., imp. 3, U30. 1963} 
Erdtman, Journ. Indian Bot. Soc . Ii2A: 35 — 38. 1963} Jafri, Fl. 
Karachi 286, 287, & 352. 1966} Martin & Drew, Jo^rm. Ariz. Acad. 
Sei. 6: Iii0~l6l. 1970} Rouleau, Guide Ind. Kew. 28 & 352. 1970} 
Lawrence, Taxon. Vase. PI., imp. 2, 687 & 782. 1971} Mukhopadhyay, 
Pollen Morph. Verb, [thesis]. 1971} Mukherjee, Trans. Bose Res. 
Inst. 35: 37—142. 1972} J. Hutehins., Fam. Flow. PI., ed. 3, U87 
& 917. 1973} D. Powell, Bull. Inst. Jam. Sei. l5 (2); h2h* 1973} 
Thanikaimoni, Inst. Fran?. Pond. Trav. Sect. Scient. Tech. 12 
(2): 20. 1973} Garcla-Earriga, Fl. Med. Colomb. 2: U97. 1975} 
Thanikaimoni, Inst. FranQ. Pond. Trav. Sect. Scient. Tech. 13s 39, 
328, & 359. 1976} Ozenda, Fl. Sahara, ed. 2, kOJ . 1977} Anon., 
Roy. Bot. Gard. Kew Lib. Curr. Awaren. 11: 21. 1978} Bhandari, 
Fl. Indian Des . 308—309. 1978} Foumet, Fl. Illustr. Phan. Guad. 
Mart. 1391 & IUI9. 1978} Mold., Phytologia UO: ia3— U23, U80, 
505, & 507. 1978} Miikherjee & Chanda, Trans. Bose Res. Inst, lil. 
Hi, U7, U8, 5U, & 57. 1978} Steyerm. & Huber, Fl. Avila 863, fig. 
lUa. 1978} Hocking, Excerpt. Bot. A.33: 90. 1979} Holm, Pancho, 
Herberger, & Plucknett, Geogr. Atlas World Weeds 51. 1979} Klein, 
Sellowia 31: I63. 1979} L6pez-Palaeio8, Revist. Fac . Farm. Univ. 
Andes 20: 17. 1979} Milz & Rimpler, Zeitschr. Naturforseh. Wiesb. 
3UC: 32U & 325. 1979} Mold., Phytologia hhi 506. 1979} Rogerson, 
Becker, & Prince, Bull. Torrey Bot. Club 106: 62. 1979} Troncoso 
in Burkart, Fl. Ilustr. Entre Rios 5» 230. 1979} J. T. & R. Kar- 
tesz, Syn. Checklist Vase, Fl. 2: U66. I98O} Mold., Phytol. Mem, 
2: U, 25, hS, 5U, 55, 59, 60, 70, 73, 75, 77, 78, 80, 85, 87, 93, 
9U, 97—101, lOU, 107, 127, 132, 139, 173, I80, 185, 3li5, 375, 
376, 396, khS, Ui8, U53, U63, & 52U. 1980} Mold., Phytologia h5» 
liO, 352, & 50U (1980), U7: li56, U58, U59, & 503 (1981). U8: U5l, 
U52, & 506 (1981), 50: 2ljO, 2U3, 2hh, 257, & 50U (1982), and 52: 
116 & 117. 1982. 
The genus Bouchea is classified in the Boraginaeeae by Rickett 
& Stafleu (i960), apparently as the result of a typographic or 
1983 Moldenke, Notes on Bouchea 311 
editorial error. 
The Graham & Johnston W^S, distributed as Bouchea sp., actu- 
ally is Ghinia curassavica f , parvifolia Mold. 
BOUCHEA AGRESTIS Schau. 
Additional bibliography: Mold., Ph(jrtologia UO: la5. 1978} 
Hocking, Excerpt. Bot. A. 33* 90. 1979; Mold., Phytol. Mam. 2i 139 
& 52li. 1980. 
BOUCHEA BOLIVUNA (Kuntze) Mold. 
Additional bibliography: Mold., Phytologia UO: Ul?. 1978} 
Mold., Phytol. Mem. 2: 173, 185, 375, 376, & S2h* 1980. 
Beck refers to this plant as a common herb, 1.2 m. tall, on 
riverbanks, at UOO ra. altitude, and states that the corollas 
are "blue". 
Additional citations: BOLIVIA: Santa Cruz: S. G. Beck 6617 (Ld) . 
BOUCHEA BOTACANA Mold. 
Synocymyt Buchea boyacana Mold, ex L6pez-Palacios , Revist. Fac . 
Farm. Univ. Andes 20: 17, sphalm. 1979. 
Additional bibliography: Mold., Phytologia liO: la5 & hl9. 1978} 
L6pez-Palacios, Revist. Fac. Farm. Univ. Andes 20: 17. 1979} Mold., 
Phytol. Mem. 2: 107, 132, 376, & 52U. 1980. 
BOUCHEA BOYACANA var. GLABRATA Mold. 
Additional bibliography: Mold., Phytologia kO: la5 & lQ.9» 1978} 
Mold., Phytol. Mem. 2: 132 & 52l|. 1980. 
BOUCHEA CHASCANOIDES Mold. 
Additional bibliography: Mold., Phytologia 29: Uli. 197U} Mold., 
Phytol. Mem. 2: 139 & 52U. 1980. 
Hatschbach & Kasper describe this plant as a shrub, 1,$ m. 
tall, with "blue" corollas, and found it growing in "encosta roch- 
osa de morro", flowering in October. 
Additional citations: BRAZIL: Minas Gerais: Hatschbach & Kasper 
la633 (Ld) . 
BOUCHEA CIPOENSIS Mold. 
Additional bibliography; Mold., Phytologia 29: hh» 197hi Mold., 
Phytol. Mem. 2: 139 & 52U. 1980. 
BOUCHEA DISSECTA S, Wats. 
Additional bibliography: H. N. & A. L. Mold., PI. Life 2: 7U. 
19U8} Mold., Phytologia UO: la5. 1978} Mold., Phytol. Mem. 2: 59, 
375, & 52U. 1980} Mold., Phytologia 50: 2liO. 1982. 
Recent collectors describe this plant as an erect, tall herb, 
30 — 80 cm. tall, and have found it growing in secondary vegetation, 
in soil from metamorphic rocks ( mirmol ) , and abundant in sandy clay 
soil with matorral vegetation of the low, spiny, deciduous type, 
along with Stenocereus quevedonis , at 100 — 1x00 m. altitude, flower- 
ing and fniiting in September and October, The corollas are said 
312 P H Y T L G I A Vol. 02, No. 5 
to have been "purple" on Soto Nuflez 10 7U and Soto Nufiez & al. 
Ilia & 16U2 and "lilac" on Delgado & al. 77. The Soto Nuflez & 
Ramirez S_. 16U2 collection, cited below, exhibits leases which in 
some cases have only a very few large lobes . 
Additional citations: MEXICO t Jalisco: Delgado , Hernindez , & 
Trejo 77 (Me— 200550) . MichoacAn: Soto Nuflez 107U (Me--290292T; 
Soto Nuflez & Ramos lllg (Me— 281a3U) } Soto Nuflez & Ramirez S. 
16U2 (Me--289868) . 
BOUCHEA FLABELLIFORMIS M. E. Jones 
Additional bibliography: Mold., Phytologia 29: U5. 197U; Mold., 
Phytol. Mem. 2: 60 & 52I4. I98O. 
BOUCHEA FLUMINEieiS (Veil.) Mold. 
Additional synonymy: Buchea fluminensis Veil, ex L6pez-4*alacios, 
Revist. Fac. Farm. Univ. Andes 20: 17, sphalm. 1979. 
Additional & emended bibliography: D. Dietr., Sjm. PI. 3: $96, 
18U3} Walp., Repert. Bot. Syst. U: 11—12 & 3li. I81i5j Bocq. in 
Baill., Adansonia, ser. 1 [Rec . Obs. Bot.] 3: 237. 1863} Bocq., 
Rev. Verbenac. 237. 1863; Baill., Hist. PI. 11: 9U. I89IJ Eriq. 
in Engl. & Prantl, Nat. Pflanzenf am . , ed. 1, U (3a)» lU3. 1895 j 
Herzog, Meded. Rijks Herb. Leid. 29: ii6. 1916 j Grenz., Ann. Mo. 
Bot. Card. 13: 85, pi. 11 & 12. 1926 j Pio Corrfea & Pena, Dice. 
PI. Uteis Bras. 3: 395. 1931; Mold., Phytologia UO: la5— I4I6. 
1978; Mukherjee 4 Chanda, Trans. Bose Res. Inst, ijl: U7. 1978; 
Klein, Sellowia 31: 163. 1979; L6pez-Palacio3, Revist. Fac. Fanm. 
Univ. Andes 20: 17. 1979; Mold., Phytol. Mem. 2: 127, 132, 139, 
173. 180, 185, 3U5, 376, & 52U. 1980. 
Additional illustrations: Grenz., Ann. Mo. Bot. Gard. 13: pi.. 
11 & 12. 1926. 
Pio Gorrea & Pena (1931) say of this plant: "Reputada anti- 
em^tica e estimulante do aparelho digestivo", giving its natural 
distribution as "Amazonas e desde a Bahia at6 S. Paulo e Minas 
Gerais, talvez em todo o Brasil". He avers that in SSo Paulo it 
is called "gervfio de folha grande", while in Kinas Gerais it is 
known as "gervSo de fQlha larga". He seems to recommend the name, 
"gervSo fadso". 
Erdtman (19U5) reports that in B_. fluminensis , as well as in 
B. prismatica (L.) Kuntze and B^ rusbyi Mold., the pollen gradns 
are "usually triporate and of a rather strange appearance" — 
this is in contradistinction to the tricolporate grains in 
Chascanum and the tetra- or pentacolpate grains in B^ linifolla 
A . Gray . He suggests that the genus Bouchea should, perhaps, be 
divided into 2 sections, the one with colpate grains (as in B. 
linifolia or probably also the related B. spathulata Torr.), and 
the other with porate grains . "Pollenmorphologically this 
would form some sort of a parallel to the two sections of Morina 
mentioned in Erdtman 191^5." 
Recent collectors refer to this plant as an herb, less than a 
meter tall, and have found it growing in more or less open campo 
1983 Moldenk©, Notes on Bouchea 313 
with secondary vegetation, in clay soil at the edge of pastures, 
in cacao plantations, and in matorral , at 350—800 m. altitude, 
flowering from February to April as well as in November and Decem- 
ber. Other collectors refer to it as as herb, O.U — 1 m. tall, 
perennial . 
The corollas are said to have been "blue" on Gentry 10063 , 
Pinheiro & Santos 2283 , and Rimachi Y.U129 , "rose with a white 
center" on Santos 20^1 , "pale purplish^nauve, white in the throat" 
on Renvoize 3U8$ , "dark-blue" on Beck 7U9U, and "violet" on 
Steinbaeh 789 . 
The Sggers lU6l8 specimen in the United States National Her- 
barium is not B. fluminensis, as are duplicates of this number 
elsewhere, but is Stachytarpheta cayennensis (L. C. Rich.) Vahl, 
30 the number seems to represent a mixture. 
Additional citations: ECUADOR: Guayas: Gentry IOO63 (Go). 
PERU: Madre de Dlos: Vargas C_. 18826 (W~2702696) . San Martin: 
Rimachi Y.m29 (N) . BRAZIL: Espirito Santo: Pinheiro & Santos 
2283 (W— 28li9207) I T. S^ Santos 20^1 (W— 281^9210) . BOLIVIA: La 
Paz: S. G. Beck 7U9£"(Ld) . Santa Cruz: R. F. Steinbaeh 789 
(Lb~6ll883). ARGENTINA: Jujuyj Renvoize 3U85 (N, W— 289U517) . 
Misiones: Krapovickas , Grist6bal , & Maruflak 1^732 (Ws, Ws). 
BOUCHEA FLUMINENSIS f . ALBIFLORA Mold. 
Additional bibliography: Mold., Phytologia 29? U8~U9. 197U} 
Mold., Phytol. Mem. 2: 139 & 52U. 1980. 
BOUCHEA FLUMINENSIS var. PILOSA Mold. 
Additional bibliography: Mold., Phytologia hO: UI6. 1978j 
Mold., Phytol. Mem. 2: 139, 18^, 376, & 52U. 1980. 
BOUCHEA INOPINATA Mold. 
Additional bibliography: Mold., Phytologia kOt lO-S. 1978} 
Mold., Phytol. Mem. 2: 139, 376, & $2U. 1980. 
BOUCHEA LINIFOLIA A. Gray 
Additional bibliography: Mold., Phytologia UO: ia6— m7. 1978| 
J. T. & R. Kartesz, Syn. Checklist Vase. Fl. 2: U66. I98O} Mold., 
Phytol. Mem. 2: U8, 5U, 60, & 52li. 1980. 
Recent collectors describe this plant as a small perennial 
herb or woody-based subshrub and have found it growing in crev- 
ices in limestone arroyos with Quercus invaginata , Acacia berlan- 
dieri, Gochnatia , Helietta , Brahea , Condalla , Brlckellia , etc. 
in a general area of Tamaulipan scrub, at 1100 — I66O m. altitude, 
in flower in May, and in both flower and fruit in June and Sep- 
tember . 
The corollas are said to have been "lavender, lighter in the 
throat" on the Wendt & Riskind collection, cited below, and 
"bright-reddish" on the Gentry & Engard collection. The two lat- 
ter collectors encountered it on sunny rocky slopes, while Stenr- 
3lli P H Y T L G I A Vol. 52, No. 5 
art refers to it as a conmon shrub on dry rocky hillsides . 
Erdtman il9h^) describes the pollen grains of B_. linifolia as 
tetra- or pentacolpate and suggests that this species, and 
probably the related B. spathulata Torr., should perhaps consti- 
tute a separate subsection or section in the genus. 
Additional citations: TEXAS: Pecos Co.: Correll & Hanson 
2988$ (Mi). Val Verde Co.: Roirell 1$27U (Ne— 128181) . MEXICO: 
Coahuila: Gentiy & Engard 23233 (Me---l6387g); Gould 1060$ (Mi)-, 
R. M. Stewart 6b (Me— 20373U) } Wendt & Riskind 1726 (Au, Me— 
2^3229). 
BODCHEA NELSONII Grenz . 
Additional & emended bibliography t Grenz., Ann. Mo. Bot. Card. 
13: 83— -811, pi. 10 & 12. 1926 J Worsdell, Ind. Lond. Suppl. 1j 132. 
19la: H. N. & A. L. Mold., PI. Life 2: 7h. I9U85 Mold., Phytolo- 
gia UO: m7. 1978; Mold., Phytol. Mem. 2: 60 70, 75, 78, 8O, 3U5, 
& 52U. 1980 J Mold., Phytologia 50: 2U3 (1982) and 52: 117. 1982. 
Additional illustrations: Grenz., Ann, Mo. Bot. Gard. 13: pl. 
10 & 12. 1926. 
Recent collectors have found this plant growing along open 
roadsides, in low grazed deciduous forests, gravelly soil in 
Acacia thorn forests, and on steep slopes with tropical deciduous 
forest in ravines, at 80 — 8 30 m, altitude, in both flower and 
fruit from August to October. They desciT-be it as an herb, 30 
cm. tall, the leaves gray-green beneath, and the U fertile stamens 
with yellow anthers. Webster and his associates report it a con- 
mon roadside weed in Oaxaca, while Stevens refers to it as "uncom- 
mon along roadsides on floodplain" in Nicaragua, Matuda I6U & 
6119 exhibit especially large leaves. 
The corollas are described as having been "lavender" on Breed- 
love U2288, "purple" on Breedlove & Thome 20872 , "pale-purple" 
on D'Arcy 12033 , "lavender-pink" on Breedlove 2827^ , "purple, 
throat white but red above" on Rohweder 3362, and ••lobes pink, 
tube white" on Clausen & Cervantes G. 6125 ^ while on Stevens 
3030 the corolla is said to have been "purple with a white center". 
On Zizumbo & Colunga 188 it is also described as "purple" and on 
Stevens 9980 "pale-purple" . 
The additional vernacular name, "xepepiend", is reported from 
Oaxaca. 
Material of B. nelsonii has been misidentified and distributed 
in some herbaria as B_. prismatica (L.) Kvintze, " B. priamatica 
(Jacq.) Kuntze", B_^ prismatica var. longirostra Grenz., Stachy - 
tarpheta sp., and Labia tae sp. 
Additional citations: MEXICO: Chiapas: Breedlove 28276 (Me — 
228538), li2288 (Me— 2U425)j Breedlove & Thome 20872 (Me— 200017) } 
Clausen & Cervantes G. 6125 (It, Me— 21U886, N)j Matuda 16922 
(Me— 81718). Guerrero: Boege I9I6 (Me— 96O89) . Oaxaca: D'Arcy 
12033 (Ld)j HernAndez M. 50a (Me— 1377i;5) ; Matuda 16U (Me— 8I686), 
1983 Moldenke, Notes on Bouchea 31$ 
6119 (Me— 81687)} Pennell , Dunn , & Dziekanowski 370 (Me--258735)> 
Webster , Miller , & Miller 13000 (Me— 133600); Zizumbo & Colunga 
188 (Ma— 26823U). NICARAGUA: Estell: TJL D. Stevens 9980 (Ld) . 
Matagalpa: W^ D^ Stevena 9828 (Ld) , Nueva Segovia: W. D, Stevens 
3030 (Ld). EL SALVADOR: La Libertad: Rohweder 3362 (E— 268lii86) . 
BOUCHEA NOTABILIS Mold. 
Synor^nny: Buchaa notabilis Mold, ex L6pez-Palacios, Revist. 
Fac. Fam. Univ. Andes 20: 17. 1979. 
Additional bibliography: Mold., Phytologia 29 t 51. 197U} L6pez- 
Palacios, Revist. Fac. Farm. Univ. Andes 20 » 17. 1979 J Mold., Phy- 
tol. Mem. 2: 107, 376, & 52li. 1980. 
BOUCHEA PRISMATICA (L.) Kuntze 
Additional & emended synonymy: Bouchea priamatica var. prismatica 
[(L.) Kuntze] apud Alain in Le6n & Alain, Fl. Cuba, imp. 1, U: 29U. 
1957. Bouchea prismatica var. prismatica Alain apud Mold., Fifth 
Summ. 1: UOO, in syn. 1971. Buchia prismatica (L.) Kuntze ex L6- 
pez-Palacios, Revist. Fac. Farm. Univ. Andes 20: 17. 1979. Bouchea 
enbergii Cham, ex Holm, Pane ho, Herberger, & Plucknett, Geogr. At- 
las World Weeds 5l, sphalm. 1979. 
Additional & emended bibliography: Sandmark in L., Amoen. Acad. 
5: 375. 1759,' J. F. Gmel. in L., Syst. Nat., ed. 13, Imp. 2, 2: Ul. 
1791} Lam. Tabl. Encycl. M^th. Bot. 1: 59. 1791} Spach, V6g. Phan. 
9: 227. 18U0} Endl., Enchirid. Bot. 312. I8la; Reichenb., Deutsch, 
Bot. [Repert. Herb. Nom.] 108. l8Ul} D. Dietr., Syn. PI. 3'. 596. 
18U3} Walp., Repert. Bot. Syst. U: 11—12 & 3U. 18U5} Lindl., Veg. 
Kingd. 66U. I8h7} Bocq. in Baill., Adansonia, ser. 1 [Rec . Obs. 
Bot.] 3: 237. 1863; Millsp., Field Columb. Mus. Publ. Bot. 1: 523. 
1902} Urb., Symb. AntUl. U: 532—533. 1911; Grenz., Ann, Mo. Bot. 
Card. 13: 78—80 & 90, pi. 9 & 12. 1926; Knuth, Feddes Repert. Spec, 
Nov. Beih. 1^3: [Init. Fl. Venez.] 603. 1927; Ekman, Arkiv Bot, 
Stockh. 22A: 5l. 1929} Mold., Suppl. List Caam, Vem. Names Ik. 
19U0} Worsdell, Ind. Lond. Suppl.' 1: 132. 19ia} H. N. & A. L. Mold., 
PI. Life 2: 33, 57, & 58. 19U8} Alain in Le6n & Alain, Fl. Cuba, 
imp. 1, U: 29U--295, fig. 127. 1957; D. Powell, Bull, Inst. Jam. 
Sci. 15 (2): U2li. 1973} Garcia Barriga, Fl. Med. Colomb. 2: U97. 
1975} Fournet, Fl. Illust. Phan. Guad. Mart. Ila9. 1978; Mold., Phy- 
tologia UO: m7— Ii21. 1978} Mukherjee &. Chanda, Trans. Bose Res. 
Inst. la: U7. 1978; Steyem. & Huber, Fl, Avila 863 & 86ii, fig. 
lUa. 1978; Holm, Pancho, Herberger, & Plucknett, Geogr, Atlas World 
Weeds 5l. 1979; L6pez-Palacios, Revist. Fac. Farm, Univ. Andes 20: 
17. 1979; Mold., Phytologia Ui: 102. 1979; J. T. & R, Kartesz, Syn. 
Checklist Vase, Fl. 2: U66. 1980; Mold., Phytol. Mem, 2: 25, li8, 
55, 60, 70, 73, 75, 77, 78, 85. 87, 93, 9U, 97— 101,10U, 107, lUi, 
127, 139, 3li5, 376, 396, Uli6, 0^8, U53, U63, & 52li. 1980; Mold., 
Phytologia U7: 1*58 (198l), 50: 2U0, 2hlx, & 257 (1982), and 52: 116. 
1982. 
Qnended illustrations: Alain in Le6n & Alain, Fl. Cuba, imp, 1, 
U: 29U, fig. 127. 1957. 
316 PHYTOLOGIA Vol. 02, No. 
Recent collectors describe this plant as a "weed" and have 
found it growing in waste places, in loose rocky soil on hill- 
sides, in sedimentary rock creekbeds, and on slopes with Croton, 
Heliocarpus , and Erythrina , at 800 — ?-l50 m. altitude, in flower in 
April, June. August, and December, and in fruit in Jvme and August. 
Ekman (1929) describes it as "a weed, not rare, throughout Haiti". 
The corollas are said to have been "mauve" on Alston 0U38 , 
"purple" on Hernandez ^ Mane las 1129 , and "violet" on Molina R. 7 6 . 
Erdtman (19U5) describes the pollen grains as triporate and "of 
a rather strange appearance", as they also are in B_. flumlnensis 
(Veil.) Mold, and B_. rusbyi Mold., but as they are not in B_. lini- 
folia A . Gray . 
The type of B_. ehrenbergii Cham., a synonym of the present tax^ 
on, was photographed in the Berlin herbarium by Macbrlde as his 
photograph number 17582. 
Knuth (1927) cites from Venezuela the following collections: 
Federal District: Moritz 293, Otto 797 , Pittier 7887 & 9729, Var" 
gas s.n. , Wagener Ui. Miranda: Pittier 5830 & 5832 . Margarita 
Island: Miller & Johnston 205. He makes the curious statement 
that B_. prismatica is not based on Verbena prismatica L. and so 
"L," does not belong in the authority citation. This claim ia 
erroneous . 
Material of typical B^ prismatica has been misidentified and 
distributed in some herbaria as its var. brevirostra Grenz . and 
even as " Stac hy tarphaeta " sp. and Verbena sp. On the other hand, 
the Breedlove St. Thome 20872 , Boege 1916 , Clausen & Cervantes G, 
6125, Herndndez M. 50a, Pennell , Dunn , & Dziekanowskl 370 , Webster , 
Miller, & Miller 13000 , and Zizumbo & Colunga 188, distributed as 
B. prismatica , are all B, nelsonii Grenz., while Dziekanowskl & 
al. 3392 is B_. prismatica var . brevirostra Orenz . 
Steyermark & Huber (1978), without justification, reduce B« 
prismatica var. brevirostra Grenz. to synonymy under typical B» 
prismatica . An examination of any series of specimens will show 
that the two taxa are distinct, provided the material is mature. 
Additional citations: MEXICO: Chiapas: Ton 1385 (Mi). More- 
los: Hunt 55 (Me--l6865l) J A. N^ Jackson l8~[Me--52962) j Lyona 
62 (Me— 268153)} ^ 8 (Me--52873) . Oaxacas Rowell , Webster . & 
Barkley 17Mli88 (Me — 175751) . State undetermined: Ehrenberg 112 
[Macbride photos 17582] (Mi— photo) . HONDURAS: Morazfint Molina 
R. 76 (Me— 81717), 27186 (Mi) . Yoro: HernAndez R. & Mancias 
1129 (Ld) . NICARAGUA: Department undetermined: L6v y s.n. (P) . 
VENEZUELA: Carabobo: Mocquerys s.n. [Mariara] (P) .' i)istrito 
Federal: Alston 51^38 (W— 2865582) . 
BOUCHEA PRISMATICA var. BREVIROSTRA Grenz. 
Additional synonyny: Buchea prismatica var. brevirostra Grenz. 
ex L6pez-Palacio8, Revist. Fac. Farm. Univ. Andes 20: 17, sphalm. 
1983 Moldenke, Notes on Bouchea 317 
1979. 
Additional bibliography: Mold., Phytologia UO; la8— k?0. 1978 j 
L6pez-Palacio3, Revist, Fac . Farm, Univ. Andes 20s 17. 1979J J. 
T, & R. Kartesz, Syn. Checklist Vase. Fl. 2: U66. 1980 j Mold., 
Phytol. Mem. 2: U8, 55, 60, 70, 77, 99—101, lOU, 107, llU, 376, 
& 52J4. 1980; Mold., Phytologia $0-. 2hh. 1982. 
Recent collectors refer to this plant as erect, 30 cm, tall, 
and have found it growing on open rocky cliffs, in heavily grazed 
thorn-scrub pastures with orchids and bromeliads on bark of Cassia, 
Acacia , and Crescentia, on semi-xeric rocky hillsides, on dry open 
roadsides, in open chaparral, among Quercus emoryi , in cultivated 
soil with Convolvulus , Cuphea , and Oxalia , and in association with 
cacti, Ipomoea arborescens , Celtis, Acacia , Maurandia , and com- 
posites, from sealevel to 2100 m. altitude, in both flower and 
fruit from July to October. Kishler found it abundant in associa- 
tion with Sanvitalia procumbens and Ipomoea longif olia . Stuesst 
refers to it as "common* in Chihuahua. 
The corollas are said to have been "blue" on Kishler 786 and 
Matuda 21869 , "violet to pale-blue" on Argflelles 8I6, "lavender" 
on Breedlove 20012 & 2765U, and "purple" on Hernandez M. 3655, 
Liogier & al. 29362, and Stuessy 1006. 
Material of this variety has been misidentified and distributed 
in some herbaria as var. longirostra Grenz. On the other hand, 
the Rowell , Webster , & Barkley 17Mli88 , distributed as and even 
previously cited by me as var. brevi rostra , seems, instead, to be 
typical B^ prismatica (L.) Kuntze. 
Additional citations: MEXICO] Aguascalientes s Rzedowski 16193 
(Me— 75181). Chiapas: Breedlove 20012 (Me— 227138), 26366 (Me— 
231860), 2765U (Me— 227115) . Chihuahua: LeSueur 136 (Me— 50919)} 
Stuessy 1006 (Me~«1363l5) . Distrito Federal: Lyonnet 317 (Me— 
239U13, Me— 239li33), 667 (Me— 239U31); Sharp lih217 (MeJT" Durango: 
Xolocotzi, Rupert, & Guevara X. 2616 (Me— 6U8IO) . Guanajuato: 
Barkley . Rowell , & Paxson 737 (Me— 89307); Kishler 786 (Me— 27U936), 
792 (Me— 27U937). Hidalgo: Hernindez M. 2655 (Me— 276501). Jal- 
isco: Barkley . Webster . & Rowell 761^2 (Me— 17389li) J Harker & Mel - 
lowea 29 (Me— 117925); Templeton 12027 (Mi) . Mexico: Gold 283 
(Me— 199553) ; Guizar N. 596 (Me— 28U9U3) J Matuda 19070 (Me— 6U8lli) , 
21703 (Me— 55581, Me— 6U812), 21869 (Me— 6U813, Me— 109170), 2665U 
(Me— 78676); Matuda & al. 28967 (Me— 78675), 293U3 (Me— 78611) . 
Michoacini Arsene 30l|0 (Me— 185U98) . Morelos: Humphreys 102 (Me — 
108615); A. W. Jackson I8 (Me— I7898I); C. H. Thomas 93 (Me— 
182721). Puebla J Miranda 20U6 (Me— 7321^7 . Quer^taro: Argtfelles 
130 (Me— 89211), 816 (Me— 2IO63I4) . San Luis Potosl: Rzedowski 
3767 (Me— 96603). Sonera: Turner , Dodge , & Mason 2061 (Me— 1^7250); 
S. S. White s.n. [2—3 Sept. 1939] (Me— 89352) . GUATEMALA: Juti- 
apa: Dziekanowski , Dunn , Trott , Thurm , Case , & Hess 3392 (N) . 
PUERTO RICO: Liogier , Liogier . & Martorell 29362 (N) . SOUTHERN 
318 P H Y T L G I A Vol. 52, No. 5 
NETHERLANDS AOTILLES: Curagao: Nagelkerken 127 (Ws) . VENEZUELA! 
Aragua: Tamayo U07$ (N) . 
BOUCHEA PRISMATICA var. LACINIATA Orenz. 
Additional bibliography: Mold., Phjrtologia UO: k20. 1978} 
Mold., Phortol. Mem. 2: 60 & 52U. 1980; Mold., Phytologia 50: 2U0. 
1982 . 
Matuda encoiintered this olant in low spiny matorral at 7^0 m. 
altitude . 
Additional citations: MEXICO: Mexico: Matuda & al. 31306 (Me— 
61i8l8, Me— 109193). 
BOUCHEA PRISMATICA var. LONGIROSTRA Orenz. 
Additional sjrnonymy: Buchea prismatica var, longirostra Grenz. 
ex L6pez-Palaci03 , Revist. Fac. Farm. Univ. Andes 20: 17, sphalm, 
1979. Bouchea prismatica var. longirostra Grezebak. ex Mold,, 
Phytologia 50: 257, in syn. 1982. Bouchea prismatica var, longi - 
rostra Gresebach ex Mold., Phytologia 50» 257, in syn. 1982. 
Bouchea prismatica var. longirostrata Greseb., in herb. 
Additional & emended bibliography: Alain in Le6n & Alain, Fl, 
Cuba, imp. 1, U: 295. 1957; Mold., Phytologia UO: U20— 1^21. 1978} 
Steyerm. & Huber, Fl. Avila 861i, 1978; L6pez-Palacios, Revist, 
Fac. Farm. Univ. Andes 20: 17. 1979} J. T, & R. Kartesz, Syn. 
Checklist Vase, Fl. 2: U66, 1980} Mold,, Phytol. Mem. 2: 25, 60, 
73, 75, 85, 87, 93, 9li, 97, lOU, 107, llli, 376, & 52li, 1980; 
Mold,, Phytologia 50: 2U0 & 257. 1982. 
Vazquez encountered this plant between 1000 and 1100 m, alti- 
tude, in flower and fruit in August. The corollas are said to 
have been "pink" on Xolocotzi & Alexander 111 . 
Steyermark & Huber (1978) without justification reduce this 
taxon to synonyny under typical B. prismatica (L.) Kxrntze, Elxam- 
ination of any large series of specimens with mature fruit will 
show that it is plainly distinct. 
The Kishler 786 & 792 and White s.n. [2—3 Sept. 1939], dis- 
tributed as var. longirostra , actually represent var. brevi rostra 
Grenz . , while Breedlove li2288 is B^ nelsonii Grenz . 
Additional citations: MEXICO: Guerrero: Webster , Rowell , & 
Barkley 17M718 (Me— 175699)} Xolocotzi & Alexander 111 (Me— 
6U811). Morelos: VAzquez 3335 (Me— 157809) . Oaxaca: Miranda lt566 
(Me— 70l|07, Me— 70U08). San Luis Potosl: Alcorn 2073 (Au) . 
BOUCHEA PSEUDOCHASCANUM (Walp.) Grenz. 
Additional bibliography: C. Muell. in Walp., Ann, Bot. Syst. 5» 
706. 1860; Pio Correa & Pena, Dice, PI, Uteis Bras, 3s 39li. 1931} 
Mold., Phytologia UO: U21. 1978; Mold., Phytol. Mem. 2: 139, 3U5, 
376, & 52U. 1980. 
Pio CorrSa (1931) says of this plant "Parece vegetar de prefer- 
Incia em terrenos arenosos" and gives its distribution as from 
Rio de Janeiro to SSo Paulo . He lists the vernacvilar names , 
"gervflc bastardo" and "gervfio false". 
1983 Moldenke, Notes on Bouchea 319 
BOUCHEA RUSBYI Mold. 
Additional bibliography: H. N. & A. L. Mold., PI. Life 2: 80. 
19U8j Mold., Phytologia UO: U21. 1978} Mold., Phytol. Mem. 2: 173 
& 52li. 1980. 
Davidson encountered this plant among "somewhat disturbed 
vegetation on steep hillsides in the Holdridge life-zone Trtiich ap- 
pears to be very moist subtropical rainforest grading into veiy 
moist lower montane rainforest, the latter especially over U500— 
5JOOO feet elevation", in flower in November, and describes the 
"corolla-lobes" as "violet" in color. An in situ photograph ac- 
companies his collection. 
Erdtman (19U5) reports that the pollen grains are "usually 
triporate and of a rather strange appearance", as are those of B. 
prisma tica (L.) Kuntze and B^ fluminensia (Veil.) Mold,, but not 
those of B. linifolia A . Gray . 
Additional citations: BOLIVIA: La Paz: Davidson U869 (N) . 
BOUCHEA SPATHULATA Torr. 
Additional bibliography: Mold., Phytologia i;0: U21— U23. 1978 j 
Hocking, Excerpt. Bot. A. 33: 90. 1979; J. T. & R. Kartesz, Syn. 
Checklist Vase. Fl. 2: h66. 1980 j Mold., Phytol. Mem, 2: U8, 60, 
& 52U. 1980. 
Eedtman (19U5), while admitting that the pollen morphology of 
this species was at that time still unknown to him, suggests that 
it may belong in a separate section of the genus with B_, linifolia 
A. Gray characterized by sessile leaves of the narrow sublinear 
type and pollen grains that are tetra- or pentacolpate, in dis- 
tinction to a section comprising B. prismatica (L.) Kuntze, B, 
fluminensia (Veil.) Mold., B. rusbyi Mold., etc, 
Stewart describes B^ spathulata as comnon on hillsides in Coa- 
huila, where it grows about 10 — 12 dm. tall, flowering in August 
with "lavender" corollas. 
Additional citations: MEXICO: Coahuila: R. M. Stewart II89 
(Me— 23UU57) . 
BOUCHEA SPATHULATA var. LONGIFLORA Mold. 
Additional bibliography: Mold., Phytologia UO: U22— U23. 1978; 
Hocking, Excerpt. Bot. A.33» 90. 1979j Mold., Phytol. Mem. 2: 60 
& 52U. 1980. 
Recent collectors describe this plant as a common shrub, 1—3 
feet tall, and have found it growing on gravelly- rocky limestone 
south-facing and lower bajada slopes covered by desert scrub of 
Acacia berlandieri , A. neovernicosa . Agave falcata, A_. lecheguilla , 
Bouteloua spp., Bouvardia , Dasylirion , Flourensia solitaria , 
Forestiera angustifolia , Fouquieria , Hechtia, Krameria grayi , 
Leucophyllum minus , Mascagnia cana , Pairbhenivun argentatum , Platy - 
opuntla, Rhus virens , Tiquilia, Viguiera stenoloba, and Yucca car- 
nerosa, at 1^50-162^ m. altitude, flowering in August. The corol- 
las are said to have been "lavender" on both the collections cited 
320 P H Y T L G I A Vol. 52, No. 5 
below. 
Additional citations: WEXICO: Coahuila: Wendt & Lott lU07 (Au)| 
Wendt, Lott, & Miapagel 187$ (Au, \lie—2S9BQor- 
ADDITIONAL NOTES ON THE GENUS CASSELIA . V 
Harold N. Uoldenke 
The previous installment of my notes on this genus vras publish- 
ed in Phjrbologia UO: li77— U79 (1978) . For a detailed explanation or 
the herbarivun acroiiyms employed in this and all others in my long 
series of papers on verbenaceous and eriocaulaceous genera, see 
Phytologia Memoirs 2: hS3 — U69 and Phytologia 50: 268. 1982. 
CASSELIA Nees & Mart. 
Additional bibliography: Cham., Linnaea 7: 36U — 366. 1832; 
Endl., Gen. PI. Is 63U. l838j Heisn., PI, Vase. Gen. 2: 199. I81i0} 
Reichenb., Deutsch. Bot. [Repert. Herb, Nom.] 108. l8Iil; Brongn., 
Enum. Gen. PI., ed. 1, 65. I8lt3j D. Dietr., Syn. PI. 3» 371, 605— 
606, & 618. 18U3; Walp., Repert. Bot. Syst. h'- 39--UO. 18U5; Lindl., 
Veg. Kingd. 66U. l8U6j A. L. Juss. in Orbigny, Diet. Univ. Hist. 
Nat. 13: I81i. I81i9i Brongn., Enum. Gen. PI., ed. 2, 119. 1850; C. 
Muell. in Walp., Ann. Bot. Syst. 5s 705. l860j Miers, Trans. Linn. 
Soc. Lond. Bot. 27: 102. 1871} Pfeiffer, Nom. Bot. 1 (2): 1855 
(187U), 2 (1): 132 (187U), and 2 (2): 1593. 187U} Durand, Ind. Gen. 
Phan. 321. 1888 j Baill., Hist. PI. 11: 80 & 98. I89I} Briq. in 
Engl. 4 Prantl, Nat. Pflanzenfam., ed. 1, U (3a)» 139— lU2, 157, & 
158, fig. Q— J. 1895} S. Moore, Trans. Linn, Soc. Lond, Hot., 
ser. 2, U: Ii39 & 521. 1895} Briq, in Engl, & Prantl, Nat. Pflanzen- 
f^., ed. 1, h (3a)s [38I], 1897} Post & Kuntze, Lexicon 103 & 688. 
190U} D. H. Scott in Solered., Syst. Anat. Dicot. [transl. Boodle 
& Fritsch] 2: 1021. 1908} Nienburg, Justs Bot. Jahresber. 39 (2): 
1051. 1916} J. C. Willis, Diet. Flow. PI., ed. 5, 12li. 1925} 
Lem6e, Diet, Descrip. Syn, Gen. PI, Phan, 8Bj 653. 19U3} H, N. & 
A, L, Mold,, PI. Life 2i 22— 2U, 31, 53, 59, 61, 63, 73, & 83. 
19U8} Metcalfe & Chalk, Anat. Dicot. 2: 1031—1033 & lOUO, 1950 j 
Lawrence, Tax, Vaso . PI., imp. 1, 688. 1951} J. C. Willis, Diet, 
Flow, PI,, ed, 6, 12li. 1951} Dalla Torre & Harms, Gen. Siphonog,, 
imp, 2, Ii31. 1958} Riekett & Stafleu, Taxon 9» 8U. I960} Dalla 
Torre & Harms, Gen, Siphonog., imp. 3, U31, 1963} Rouleau, Guide 
Ind. Ketw. 37, 188, & 352. 1970} Lawrence, Tax. Vase. PI., Inp. 2, 
688. 1971} Napp-Zinn, Anat. Blatt. A (l)i la8. 197U} Mold., ?hyto- 
logia UOi hn — U79 & 506. 1978} Mukherjee & Chanda, Trans. Bose 
Res. Inst. Ul: UO, li5, & U7. 1978} Hocking, ExceiT)t. Bot, A.33i 90. 
1979} Rogerson, Becker, 4 Prince, Bull. Torrey Bot. Club IO61 62.