JOURNAL OF THE ARNOLD ARBORETUM THE GENERA OF THE URTICACEAE IN THE SOUTHEASTERN UNITED STATES NORTON G. MILLER URTICACEAE A. L. de Jussieu, Gen. PL. 400. 1789, "Urticae," nom. cons. (NETTLE FAMILY) Taprooted and rhizomatous annual and perennial herbs [or shrubs,rarely large trees], with watery [rarely milky] sap, generally providedwith a vesture of some kind, sometimes with stinging hairs. Stems oftenfluted, mostly nonsucculent, but sometimes fleshy or translucent. Leavesalternate or opposite, sometimes anisophyllous, simple, usually petiolate,blades mostly ovate to lanceolate [rarely lobed]; punctiform, linear [orstellate] cystoliths borne in cells of the adaxial (upper) or abaxial (lower)epidermis or both; margins serrate, dentate, entire, or rarely incised,venation pinnate or palmate; stipules present or absent, if present, eitherpaired at the petiole base or intrapetiolar and partly to entirely connate,fugacious or not. Plants monoecious, dioecious, or polygamous; flowersimperfect, rarely perfect inflorescences basically cymose, axillary, bracte-ate, the flowers arranged in loose to tight clusters, or inflorescence some-times racemose or paniculate [or flowers aggregated on a flat or campanu-late to figlike recenptacle; if monoecious, staminate and carpellate flowersin different inflorescences or intermingled. Staminate flowers small, green-ish or whitish, rarely otherwise with short pedicels or sessile, caducousafter shedding ollen; tepals 4 5 [6, equal, valvate, or partly fused A Prepared for a generic flora of the southeastern United States, a project of the Arnold Arboretum and the Gray Herbarium of Harvard University made possible through the support of the National Science Foundation (Grant GB-6459X, principal in te first contribution o the ser.. The present treatment follows the plan initiated in the first contribution of the series (Jour. Arnold Arbh. 39: 296-346. 1958). The area dealt with includes North and South Carlina, Georgia, Floda, Tennessee, Alabama, Mississippi, Arkansas, and Louisiana. Descriptions are based on species occurring in this region, with additional information from extra-regional taxa in brackets. Ranges of treated species are derived mostly from specimens examined and partly from data already published. References followed by an asterisk are those which I have not been able to check. Sam grateful to Dr. Wood for his continuing advice and assistance; to Dr. Bernice . Schubrt, who kindly examined a number of specimens at the New York Botanical Garden for me; and to Dr. Gordon p. DeWolf, Jr., who helpfully provided living plan of tPrietari Nforidana I have checked ranges against material in the herbarium of the University of North Carolina and thank its staff for giving me access to the letin nde their care. Mrs. Nancy Dunkly has helped greatly with bibliographic matters and in the reparation of the final typescript In the illustration of Piea, or-f ^ r danby Miss Rachel A. Weele script. In the illustration of Pilea, a-f were drawn by Miss Rachel A. of theler; all other drawings are the conscientious work of Miss Virginia Savage. any of the illustrations are based on living or pre-see material suppli by Dr. ments d. Dr. John W. Thieret read the manuscript and I am thankful for his comments.[voL. 52
BioStor
