ROGERS, CASUARINACEAE THE CASUARINACEAE IN THE SOUTHEASTERN UNITED STATES' GEORGE K. ROGERS CASUARINACEAE R. Brown in Flinders, Voy. Terra Austral. 2: 571.July-Aug. 1814. "Casuarineae." [Mirbel, Ann. Mus. Natl. Hist. Nat. 16: 451. 1810. "Casuarinees"], nom. cons. (BEEFWOOD FAMILY, SHE-OAK FAMILY) A unigeneric family of evergreen [shrubs and] trees with minute. whorled,basally connate, scalelike leaves and thin, ridged and grooved [or quadran-gular], articulated branchlets. Plants monoecious or dioecious, with reducedanemophilous flowers, the staminate flowers whorled in terminal, simple[or compound], bracteate spikelets; the carpellate flowers whorled in brac-teate, globose to ellipsoid heads, usually on short, lateral branchlets. Ovulessemi-anatropous, bitegmic, crassinucellate. Infructescence woody and cone-like, bearing winged, indehiscent samaras, each enclosing a single seed with-out endosperm. TYPE GENUS: Casuarina Adanson.1. Casuarina Adanson, Fam. P1. 2: 481, 534. 1763. Plants to ca. 25(-45) m tall, to ca. 70 cm [-I m] in diameter. Trichomesunicellular or uniseriate, often branched. Calcium oxalate crystals often sin-gle or clustered in parenchymatous tissues, sometimes in phloem fibers.Leaves in whorls of [4 or] 5-16, the free tip of each protruding as the ex-tension of a cauline ridge, linear to narrowly elliptic-lanceolate or narrowlydeltoid [or ovate], erect or recurved, usually pubescent along the margins but 'Prepared for the Generic Flora of the Southeastern United States, a project of the ArnoldArboretum currently made possible through the support of the National Science Foundationunder Grant DEB-81-11520 (Carroll E. Wood. Jr., and Norton G. Miller, principal in-vestigators). This treatment, the 89th in the series, follows the format established in thefirst paper (Jour. Arnold Arb. 39: 296-346. 1958) and continued to the present. The areacovered by the Generic Flora includes North and South Carolina. Georgia. Florida, Ten-nessee. Alabama, Mississippi, Arkansas. and Louisiana. The descriptions are based pri-marily on the plants of this area, with information about extraregional members of a familyor genus in brackets [ ]. I thank Carroll Wood and Norton Miller for their advice, suggestions, and careful reviewof the manuscript. The valuable hours that my wife. Donna Rogers, spent translating pa-pers in French are acknowledged with extreme gratitude. George Staples generously sharedliterature, notes, and observations stemming from his own interest in Casuarina. Usefulcomments on the nomenclatural problems were supplied by Edward Voss. Arnold D. Clap-man drew the illustration using specimens collected by George R. Cooley, David Seligson,and Carroll Wood.� President and Fellows of Harvard College. 1982.Journal of the Arnold Arboretum 63: 357-373. October, 1982.
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