1989]PRICE, PINACEAE247 THE GENERA OF PINACEAE IN THE SOUTHEASTERN UNITED STATES' ROBERT A. PRICE2 PINACEAE Lindley, Nat. Syst. Bot. ed. 2. 313. 1836, nom. cons. (PINE FAMILY) Evergreen [or deciduous], monoecious trees (occasionally shrubs), usuallypyramidal in form. Leaves linear to acicular [rarely lanceolate], spirally ar-ranged (often apparently 2-ranked). sessile or short-petiolate; foliage leaveseither borne on long-shoots or clustered tightly on short-shoots. Pollen cones(microsporangiate strobili) bearing spirally arranged, bilaterally symmetricalmicrosporophylls; microsporangia 2 on the abaxial surface of each sporophyll,longitudinally or transversely [or obliquely] dehiscent; pollen grains bisaccate.or less commonly with the saccae reduced to a frill (in Tsuga sect. TSUGA) [ornonsaccate in Larix and tPseudotsuia] prothallial cells 2. Ovulate cones with 'Prepared for the Generic Flora of the Southeastern I nited States, a long-term project made possiblethrough the support of National Science Foundation (rant BSR-8415769 ((. E. Wood. Jr., principalinvestigator), under which this account was prepared, and BSR-8415637 (N. G. Miller, principalinvestigator). The 127th in the series, this paper follows the format established in the first one (Jour.Arnold Arb. 39: 296-346. 1958) and continued to the present. The area covered by the Generic Floraincludes North and South Carolina. Georgia. Florida. Tennessee. Alabama. Mississippi. Arkansas,and Louisiana. The descriptions are based primarily on the plants of this area, with informationabout extraregional members of a family or genus in brackets. The references that I have not veritiedare marked with asterisks. I thank Carroll Wood and Norton Miller for the opportunities alforded by participation in theGeneric Flora project and for their guidance and editorial help in the study. I am grateful to IhsanAl-Shehbaz for his generous help during my visits to Harvard University: to William (ritchlield, M.Thompson (onkle, and Eugene Za;arin for sharing insights and prepublication manuscripts on thePinaceae: and to Rudolf Schmid for bibliographic assistance. Library and herbarium collectionsassociated with Harvard University and the University of California, Berkeley, were consulted inthis study, and I wvish to thank the librarians and curatorial staffs of these institutions. ElizabethSchmidt and Stephen Spongberg added their editorial expertise to the manuscript. The illustrations were drawn by Karen Stoutsenburger in 1977 under the supervision of CarrollWood. who also made the dissections. Living, liquid-preserved, and some of the dry materials usedwere variously collected by Walter Judd. Norton Miller, and Carroll Wood. Herbarium materialsinclude an unattributed sheet [] wiith mature seeds of /'inus Strohus utilized by (. E. Faxon for theillustration in Sargent's S.ilva of \iori lr mrica, as well as specimens collected by Adams & Wherry(22'2 [(H]). R. M. Harper (2023, 2027 [GH]). and J. D. Smith (s.n., Aug. 1881 [(n]). The diagrammaticcross sections of leaves of Pinus are after J. W. Lanyon's A Card Key to Piius Based on Needlelnaion1m . The pollen grain of I' Stroius is after a photograph made by Norton Miller. 'University Herbarium. Department of Botany. University ofCalifornia. Berkeley, California 94720.Please address reprint requests' I)r. Norton Miller. Room 3132 CEC, New York State EducationDepartment. Albany. New York 12230.c President and Fellows of Harvard College, 1989.Journal of the Arnold .Arboretum 70: 247-305. April. 1989.
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