JOURNAL OF THE ARNOLD ARBORETUMVOL. 50 JULY 1969 NUMBER 3 A REVISION OF THE MALESIAN AND PACIFIC RAINFOREST CONIFERS, I. PODOCARPACEAE, IN PART * DAVID J. DE LAUBENFELSDacrycarpus (Endlicher) de Laubenfels, stat. nov. Podocarpus sect. Dacrycarpus Endlicher. Syn. Conif. 221. 1847. Type species: Podocarpus imbricatus Blume [Dacrycarpus imbricatus (Blume) de Lau-benfels]. Podocarpus sect. Dacrydioideae Bennett ex Horsfield, PI. jav. rar. 41. 1838. Type species: Podocarpus dacrydioides Rich. [Dacrycarpus dacrydioides (Rich.) de Laubenfels]. Folia parva vel squamata. Strobili feminei terminales; receptaculumverruculosum, tandem carnosum; unus vel duo bracteae terminalae fer-tiles, cum ovulo connatum in forma crista superans; ovulum inversumepimatium contingens. The distinguishing character of Dacrycarpus is the union of the bractwith the seed and seed scale on one side, forming a projecting crestparticularly noticeable on immature fruit. As in most of the family, theseed is inverse and Dacrycarpus resembles Podocarpus, of which it haslong been treated as a section, because of the union of the fertile scalewith the seed and its distinct receptacle. In addition to the fusion of thefertile bract with the corresponding scale and seed, however, is the factthat the cone is produced terminally on leafy branches and not onspecialized shoots or peduncles. The leaves of Dacrycarpus also differmarkedly from Podocarpus and resemble those of Dacrydium, beingsometimes difficult to distinguish in the sterile form. There are, in addi-tion, New Zealand species of Dacrydium in which the seed is covered bythe scale and remains inverted. A fleshy development upon maturityof the structure below the seed is normal in Dacrydium. In short. Dacry-carpus is as closely related to Dacrydium as to Podocarpus while theunion of the bract with the seed sets it sharply apart from either ofthem. The wartyness of the receptacle is unusual, a character shared onlywith Acmopyle. In the juvenile form Dacrycarpus is interesting in that * Continued from volume 50, p. 314
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