JOURNAL OF THE ARNOLD ARBORETUM THE GENERA OF ARALIACEAE IN THE SOUTHEASTERN UNITED STATES 1 SHIRLEY A. GRAHAMARALIACEAE A. L. de Jussieu, Gen. PI. 217. 1789, "Araliae," nom. cons. (GINSENG FAMILY) Perennial herbs, vines, shrubs [or trees, rarely scandent epiphytes].Leaves alternate on the stem or basal from the rhizome, petiolate,[simple,] palmately compound or pinnately or ternately compound ordecompound, the stipules either adnate (in varying degrees) to the baseof the slightly sheathing petiole or wanting. Inflorescence basically umbel-late, the umbels many flowered, solitary or arranged in panicles, racemes[corymbs, or rarely in compound umbels], the peduncles and pedicelssubtended by lanceolate, deciduous or persistent bracts. Flowers regular,bisexual and/or unisexual, the plants polygamomonoecious, polygamodioe-cious, or dioecious. Floral tube adnate to the ovary, cupuliform, obconic, orcylindrical, articulated or continuous with the pedicel; calyx lobes 5,small, deltoid [or the apex of the floral tube merely undulate], persistent.Petals 5 [-10, valvate or imbricate in bud, broad [or narrow] at base,greenish white to pink, arising from the margin of a fleshy, epigynous,flattened or cupuliform nectariferous disc, free and separately deciduousor � connate and falling as a unit. Stamens 5 [to many], generally asmany as the petals and alternate with them, inserted epigynously; fila-ments filiform [or ligulate]; anthers oblong [to subglobose], dorsifixed,versatile, 2-locular, longitudinally dehiscent; pollen usually 30-40 i long,oblate-spheroid [to prolate], 3-colporate, the pores often transverselyelongated, the exine reticulate. Gynoecium 2-5(6) [-15 ]-carpellate, syn-carpous; styles persistent, generally as many as the carpels, distinct or 1 Prepared for a generic flora of the southeastern United States, a joint project of the Arnold Arboretum and the Gray Herbarium of Harvard University made possible through the support of George R. Cooley and the National Science Foundation and under the direction of Carroll E. Wood, Jr., and Reed C. Rollins. The treatment follows the pattern established in the first paper in the series (Jour. Arnold Arb. 39: 296-346. 1958) and continued to the present paper. The area covered includes North and South Carolina, Georgia, Florida, Tennessee, Alabama, Mississippi, Arkansas, and Louisiana. The descriptions apply primarily to the plants of this area, with supple-mentary information in brackets. References not seen by the author are marked with an asterisk. The author is indebted to Dr. Lily M. Perry for references to Chinese medicinal uses of Panax and to Dr. Wood for his generous assistance and suggestions. Harry E. Ahles and Wilbur H. Duncan have kindly contributed information on Hedera in the Carolinas and Georgia, respectively.[VOL. 47
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