GRAHAM, GENERA OF LYTHRACEAE THE GENERA OF LYTHRACEAE IN THE SOUTHEASTERN UNITED STATES SHIRLEY A. GRAHAMLYTHRACEAE Jaume St.-Hilaire, Expos. Fam. 2: 175. 1805, "Lythrariae," nom. cons. (LOOSESTRIFE FAMILY) Herbs or shrubs [rarely small trees], with quadrangulate or teretestems. Leaves opposite, seldom alternate or whorled, simple, entire,exstipulate. Flowers regular or irregular, axillary or in terminal racemesor spikes, sometimes dimorphic or trimorphic with regard to style andstamen length, bisexual, [3]4-6-merous, with perianth and stamensperigynous, the flowers rarely cleistogamous; bracteoles 2, opposite onthe pedicel, rarely none. Floral tube campanulate to tubular, persistent,often conspicuously nerved; calyx lobes 4-6, alternating with 3-5 deltoidappendages, or the appendages wanting. Petals 0-6, distinct, crumpled,deciduous [rarely persistent], inserted on the inner surface of the floraltube between the calyx lobes. Stamens as many as or twice as many asthe petals [or more numerous], often alternately unequal, inserted on theinner surface of the floral tube below the petals, included to exserted;anthers versatile [rarely basifixed], introrse, 2-locular, longitudinallydehiscent. Gynoecium syncarpous, often surrounded at the base by ahypogynous disc, or in irregular flowers the disc on the upper (adaxial)side only; stigma capitate, rarely bilobed; style filiform; ovary superior,free in the floral tube, 2-4[-6]-locular, the septa complete or reduced;placentation axile [rarely parietal]; ovules anatropous and epitropous,with 2 integuments. Fruit a membranaceous [or leathery] capsule in-closed by the persistent floral tube, septicidally or loculicidally dehiscentor indehiscent, splitting irregularly. Seeds 3 to many, minute or up to ca.4 mm. long, pyramidal, ovoid, or discoid, sometimes slightly winged, with 'Prepared for a generic flora of the southeastern United States, a joint project ofthe Gray Herbarium and the Arnold Arboretum made possible through the supportof George R. Cooley and the National Science Foundation and under the directionof Reed C. Rollins and Carroll E. Wood, Jr. The scheme follows that outlined at thebeginning of the series (Jour. Arnold Arb. 39: 296-346. 1958). The area covered inthis, as in former treatments, is bounded by and includes North Carolina, Tennessee,Arkansas, and Louisiana. Material included in the descriptions in brackets applies tospecies outside this area, and references marked by an asterisk have not been seen bythe author. The aid of Dr. Wood and the observations and suggestions of Drs. H. E. Ahles,R. K. Godfrey, D. B. Ward, and C. A. Brown are gratefully acknowledged. Dr.R. B. Channell kindly supplied living material of Cuphea for study.1964]
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