AL-SHEHBAZ, ANCHONIEAE THE GENERA OF ANCHONIEAE (HESPERIDEAE) (CRUCIFERAE; BRASSICACEAE) IN THE SOUTHEASTERN UNITED STATES'2 IHSAN A. AL-SHEHBAZ3 Tribe Anchonieae A. P. de Candolle, Syst. Nat. 2: 152. 1821. Annual, biennial, or perennial herbs [rarely shrubs]; trichomes unicellular,eglandular, simple or furcate, stellate, dendritic, or malpighiaceous, sometimesmixed with multicellular glandular ones. Inflorescences ebracteate [rarely brac-teate], corymbose racemes [rarely panicles], usually elongated in fruit. Sepalserect, free or sometimes connivent, usually unequal and slightly to conspicu-ously saccate at base. Stamens 6, often strongly tetradynamous; median fila-ments free [rarely connate], unappendaged [or dentate]. Fruits dehiscent orindehiscent and usually lomentaceous, breaking into 1-seeded parts [rarelynutlike], terete, tetragonal, or flattened parallel [or at right angles] to the septum;styles conspicuous [or obsolete]; stigmas strongly 2-lobed [rarely entire], thelobes connivent or spreading, decurrent or not [rarely forming conspicuoushorns or appendages]. Seeds few to numerous, uniseriately [or biseriately] ar-ranged in each locule, wingless [or winged], nonmucilaginous [sometimes mu-cilaginous] when wet; cotyledons accumbent or incumbent. (Including Buni-adeae DC., Cheirantheae Webb & Berth., Erysimeae Dumort., HesperideaePrantl, Matthioleae O. E. Schulz.) TYPE GENUS: Anchonium DC. 'Prepared for the Generic Flora ofthe Southeastern United States, a long-term project made possibleby grants from the National Science Foundation and currently supported by BSR-8415769 (C. E.Wood. Jr.. principal investigator), under which this research was done, and BSR-8415637 (N. G.Miller, principal investigator). This account, the 121st in the series, follows the format establishedin the first paper (Jour. Arnold Arb. 39: 296-346. 1958) and continued to the present. The areacovered by the Generic Flora includes North and South Carolina, Georgia, Florida. Tennessee.Alabama, Mississippi. Arkansas, and I ouisiana. The descriptions are based primarily on the plantsof this area. with information about extraregional members of a family or genus in brackets. Thereferences that I have not verified are marked with asterisks. I am most grateful to Carroll Wood for his continuous support. advice, and help during thepreparation of this paper, and particularly for his critical review of the manuscript. I should also liketo thank Norton G. Miller for reviewing the paper. Reed Cf. Rollins and Robert A. Price for theirvaluable discussions, and Barbara Nimblett for typing the manuscript. I am grateful to Elizabeth B.Schmidt and Stephen A. Spongberg for their editorial ad ice. 2For an account of the family and its tribes, see AI-Shehba,. The Tribes of('ruciferae (Brassicaceae)in the Southeastern t nited States. Jour. Arnold Arb. 65: 343-373. 1984. 'Arnold Arboretum. Harvard I niversity. 22 l)ivinity Avenue. Cambridge. Massachusetts 02138.� President and Fellows of Harvard College. 1988.Journal of the Arnold Arboretum 69: 193-212. Jul., 1988.193
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