JOURNAL OF THEARNOLD ARBORETUMVOLUME 67 JULY 1986 NUMBER 3 THE GENERA OF LEPIDIEAE (CRUCIFERAE; BRASSICACEAE) IN THE SOUTHEASTERN UNITED STATES'2 IHSAN A. AL-SHEHBAZ3 Tribe Lepidieae A. P. de Candolle, Syst. Nat. 2: 151. 1821. Annual, biennial, or perennial herbs [sometimes subshrubs or shrubs], gla-brous or with simple or rarely stellate [or bifurcate or dendritic] trichomes.Inflorescence an ebracteate [or bracteate], corymbose raceme, usually elongatedin fruit; flowers few to many [rarely solitary on long scapes], actinomorphic orslightly [to strongly] zygomorphic, perfect [or very rarely imperfect and plantsmonoecious or dioecious]. Sepals erect to ascending or spreading, not saccate[or strongly saccate] at base. Petals present or absent, equal or unequal. Nectarglands 2, 4, or 6, distinct or connate, subtending [or surrounding] the bases of 'Prepared for the Generic Flora of the Southeastern United States, a long-term project made possibleby grants from the National Science Foundation and currently supported by BSR-8415769 (C. E.Wood, Jr., principal investigator), under which this research was completed, and BSR-8415637 (N. G.Miller, principal investigator). This account, the 11 th in the series, was begun under BSR-8111520(C. E. Wood, Jr., principal investigator). It follows the format established in the first paper (Jour.Arnold Arb. 39: 296-346. 1958) and continued to the present. The area covered by the Generic Floraincludes North and South Carolina, Georgia, Florida, Tennessee, Alabama, Mississippi, Arkansas,and Louisiana. The descriptions are based primarily on the plants of this area, with informationabout extraregional members of a family or genus in brackets [ ]. The references that I have notverified are marked with asterisks. I am most grateful to Carroll Wood for his help with many aspects of this paper, especially hiscritical review of the manuscript. I am variously indebted to Reed C. Rollins, Norton G. Miller,George K. Rogers, R. Dale Thomas, John W. Thieret, and Jimmy R. Massey, as well as to BarbaraNimblett, who typed the manuscript. I also wish to thank Elizabeth B. Schmidt and Stephen A.Spongberg for their editorial advice. The illustrations were made by Karen Stoutsenberger under earlier grants. Carroll Wood and/orKenneth R. Robertson prepared the material and supervised the illustrations. Fresh material wascontributed by Wood, and the fruits and seeds are from herbarium specimens in the Gray Herbariumor the Arnold Arboretum. 2For an account of the family and its tribes, see Al-Shehbaz, The tribes of Cruciferae (Brassicaceae)in the southeastern United States. Jour. Arnold Arb. 65: 343-373. 1984. 'Arnold Arboretum, Harvard University, 22 Divinity Avenue, Cambridge, Massachusetts 02138.� President and Fellows of Harvard College, 1986.Journal of the Arnold Arboretum 67: 265-311. July, 1986.