AL-SHEHBAZ, CRUCIFERAE THE TRIBES OF CRUCIFERAE (BRASSICACEAE) IN THE SOUTHEASTERN UNITED STATES' IHSAN A. AL-SHEHBAZ The family Cruciferae is represented in the southeastern United States by121 species in 43 genera assigned to seven tribes. The present account includesa family description, with general comments on the group as a whole; selectedfamily references; a key to the tribes; an artificial key to 46 genera (includingthree known as escapes from cultivation to the west of this area); and briefdescriptions of the tribes, each with a list of the representative genera in theSoutheast. When treating a family generally recognized as difficult for generic and tribaldelimitation, one faces the problem of how genera should be arranged. Analphabetic sequence would definitely be incompatible with the scope of ourflora. A few students of the family avoid recognizing tribes because they believethat tribal boundaries are usually artificial. However, these authors arrange thegenera according to their nearest sister relatives-a disposition that often co-incides so well with the tribal classification that ignoring or totally abandoningthe tribes is unreasonable. Nearly half of the genera of Cruciferae occurring inour area belong to the tribes Thelypodieae, Brassiceae, and Lepidieae, whichare widely recognized as natural groups. The majority of the remaining generafall within the presumably well-defined centers of four other tribes. For thesereasons I favor the use of tribes to provide a workable framework, even though 'Prepared for the Generic Flora of the Southeastern United States, a long-term project made possibleby grants from the National Science Foundation and currently supported by BSR-8111520 (C. E.Wood, Jr., principal investigator), under which this research was done, and BSR-8303100 (N. G.Miller, principal investigator). This account, the 103rd in the series, follows the format establishedin the first paper (Jour. Arnold Arb. 39: 296-346. 1958) and continued to the present. The areacovered by the Generic Flora includes North and South Carolina, Georgia, Florida. Tennessee.Alabama, Mississippi, Arkansas, and Louisiana. The descriptions are based primarily on the plantsof this area, with information about extraregional members of a family or genus in brackets [ ]. Thetwo references that I have not verified are marked with an asterisk. I am most grateful to Carroll Wood for his support, guidance, and help with many aspects of thispaper, and particularly for his critical review of the manuscript. Reed C. Rollins, Norton G. Miller,George K. Rogers, and Elizabeth A. Shaw were helpful throughout this study, and I am indebted toBarbara Nimblett for her careful typing of the manuscript. I am grateful to Elizabeth B. Schmidt andStephen A. Spongberg for their editorial help. The illustrations were made by Karen Stoutsenberger (primarily under earlier grants), with theexception of FIGURES 1, S (by Rachel A. Wheeler) and 3, a-c (by the late Dorothy H. Marsh). CarrollWood and/or Kenneth R. Robertson prepared the materials and supervised the illustrations. Freshmaterial was contributed by K. R. Robertson, R. C. Rollins, and C. E. Wood, and the fruits andseeds are largely from herbarium specimens of the Arnold Arboretum and the Gray Herbarium.C President and Fellows of Harvard College, 1984.Journal of the Arnold Arboretum 65: 343-373. July, 1984.3431984]
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