SYSTEMATICS AND GEOGRAPHIC HISTORY OF THE GENUS PYRGUS IN THE PALAEARCTIC REGION (LEPIDOPTERA, HESPERIIDAE) by R. DE JONG Rijksmuseum van Natuurlijke Historie, Leiden Abstract A systematic and Zoogeographie study on the Palaearctic species of the genus Pyrgus, comprising a taxonomie review, a description of the geographic variation and a reconstruction of the geographic history. Some Zoogeographie problems are discussed and the Pyrgus species are classified as faunal types. Contents 1. Introduction 2 a. Scope 2 b. Taxonomie categories 2 c. Wing markings and genital armature 6 d. Nomenclature 6 e. Material 7 f. Acknowledgements 7 g. Methods, measurements and abbreviations 8 2. The genus Pyrgus Hübner, 1819 8 a. Delimitation 8 b. Classification 10 Subgenus Scelotrix 12 Subgenus Pyrgus 14 The alt eus complex 16 Species of uncertain affinities 23 The American Pyrgus species 24 c. Distribution 24 d. Geographic and evolutionary history 25 Geographic history of the subgenus Scelotrix 27 Geographic history of the alveus species complex 30 3. The species of Pyrgus in the Palaearctic 31 4. Discussion of some Zoogeographie problems 99 a. The geographic origin of species and groups of species 99 b. The geographic history of the Palaearctic Pyrgus species, in relation to the general history of the Palaearctic fauna 100 5. Zoogeographie classification of the Palaearctic Pyrgus species 105 6. Summary 107 7. Literature 115