The haplochromine species (Teleostei, Cichlidae) of the Cunene and certain other Angolan rivers *o\ f "^w /? Peter Humphry Greenwood Department of Zoology, British Museum (Natural History), Cromwell Road, London SW7 5BD . BRIT Contents Introduction 187 Methods and materials 188 The haplochromine species of the Cunene river 189 Thoracochromis Greenwood, 1979 189 Thoracochromis buysi (Penrith), 1970 190 Thoracochromis albolabris (Trewavas & Thys van den Audenaerde), 1969 . 197 Orthochromis Greenwood, 1954 206 Orthochromis machadoi (Poll), 1967 206 Pseudocrenilabrus Fowler, 1934 213 Pseudocrenilabrus philander (Weber), 1897 214 Serranochromis Regan, 1920 216 Serranochromis (Sargochromis) Regan, 1920 216 Serranochromis (Sargochromis) gracilis sp. nov 225 Serranochromis (Serranochromis) Regan, 1920 228 Serranochromis (Serranochromis) thumbergi (Castel.) 228 Serranochromis (Serranochromis} macrocephalus (Blgr) 229 Serranochromis (Serranochromis) angusticeps (Blgr) and S. (S.) robustus jallae(E\gr) 229 Zoogeographical considerations 229 Appendix I 233 The generic status of various Angolan haplochromine species previously referred to Haplochromis by Regan (1922), Trewavas (1973) and Bell-Cross (1975). Appendix II The generic status of Pelmatochromis welwitschi Blgr, 1898 234 Acknowledgements 238 References 238 Introduction L 'Angola est un plateau d' ou descendant de nombreux Jleuves et rivieres qui cachet encore bien des secrets Poll (1967:16) Despite Poll's (1967) extensive monograph on the fishes of Angola, and the work of Trewavas (1964, 1973) and Bell-Cross (1975) much has still to be learnt about the biology, taxonomy and zoogeography of the haplochromine cichlid species in this region of Africa (see Greenwood, 1979). A basic inventory of the species has been worked out, but many taxa are known only from the type specimen, by a limited number of type specimens or by some specimens whose locality is recorded no more precisely than 'Angola'. Above all, almost nothing is known about the phyletic relationships of the species, and hence zoogeographical conclusions based on them are correspondingly uncertain. Judging from the present-day hydrography of Angola, especially the isolated rivers which discharge directly into the Atlantic, and the numerous tributaries emptying into the Zaire system, one might expect a high degree of localized endemicity in the various rivers. In other Bull. Br. Mus. nat. Hist. (Zool.) 47(4): 1 87-239 Issued 27 September 1984 187
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