PROC. ENTOMOL. SOC. WASH. 107(4), 2005, pp. 869-882 TWO NEW SPECIES AND A NEW RECORD OF EUCOENOGENES MEYRICK (LEPIDOPTERA: TORTRICIDAE) FROM THAILAND WITH A DISCUSSION OF CHARACTERS DEFINING THE GENUS Nantasak Pinkaew, Angsumarn Chandrapatya, and Richard L. Brown (NP) Department of Entomology, Faculty of Agriculture Kamphaeng Saen, Kasetsart University, Kamphaeng Saen Campus, Nakhon Pathom 73140 Thailand (e-mail:
[email protected]); (AC) Department of Entomology, Faculty of Agriculture, Kasetsart University, Bangkok 10900, Thailand (e-mail:
[email protected]); (RLB) Mississippi En-tomological Museum, Box 9775, Mississippi State, MS 39762, U.S.A. (e-mail:
[email protected]) Abstract. — Two new species of Eucoenogenes Meyrick (Lepidoptera: Tortricidae: Eu-cosmini), E. bicucullus Pinkaew and E. vaneeae Pinkaew, are described and illustrated from montane evergreen forest in Thong Pha Phum National Park, Kanchanaburi Prov-ince, Thailand. Epinotia munda Diakonoff, described from a female collected in Sumatra, is transferred to Eucoenogenes (n. comb.) based on male and female specimens collected in Thailand; the imago and genitalia of both sexes are described and illustrated. Characters for defining the genus are discussed and illustrated. Key Words: new combination, distribution records, generic characters, Acroclita, Epi-notia, Eucosmini Eucoenogenes Meyrick, 1939, was pro-posed as a replacement name for Caeno-genes Meyrick, 1937, which was preoccu-pied by Caenogenes Walsingham, 1887. Meyrick (1937) described the type species, Caenogenes melanancalis, based on two specimens reared from larvae mining leaves of Eugenia jambolana Lamark (Myrtaceae) in India. Meyrick defined the genus by the thickened palpus with densely appressed scales concealing the third segment, the ab-sence of the R4 vein in the forewing, and absence of the M^ vein in the hindwing. Clarke (1958) designated a female as lec-totype, noting that "the male is missing," even though Meyrick indicated that both specimens were female in his original de-scription. However, Clarke (1955) previ-ously noted that Meyrick often misidenti-fied sexes because of his reliance on a hand lens and emphasis on non-genitalic char-acters; thus, the sex and present location of the second specimen is uncertain. In con-trast to Meyrick's description. Clarke's fig-ure of venation showed that both Rj in the forewing and M^ in the hindwing were pre-sent and that all veins were separate except the fused anal veins in the forewing and the stalked Mj-CuA, in the hindwing; this ve-nation has been confirmed by a re-exami-nation of the venation of the lectotype. Diakonoff (1967) considered the female genitalia of E. melanancalis to lack char-acters of generic significance and consid-ered the palpi with appressed scales and concealment of the third segment to be the only character defining the genus. Based on similar palpi, Diakonoff described E. del-tostoma based on a single male specimen from the Philippine Islands. This descrip-
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