437 THE CARBONIFEROUS TRTL0RTTE8 OF AUSTRALIA. By John Mitchell, late Principal, Technical College at Newcastle, New South Wales. (Plates xlvi.-liii.) Introduction. For many years, m\' collection has contained a number of Carboniferous Trilobites new to science. When the opportunity of describing tliem presented itself, my first intention was to confine myself to these specimens. On subsequent consideration, it occurred to me that palaeontoloi^ical knowledge would be increased by extending the scope of my paper so as to embrace all previous work on the Carboniferous Trilobita of Australia, to describe all new forms that might be available from other sources, and review, redescribe, and redetermine, where it might seem necessary, those species referred to by previous authors. It will be found, in the course of this paper, that, in a number of instances, my views are widely divergent from those of pre- vious authors; and, while regretting this divergence, it has not arisen through rashness or want of reflection on my part; and I may candidly affirm that my inability to reconcile the deter- minations of previous authors with my own, in some cases, was the only unpleasant part of the work expended on this paper. My aim is to clear the literature on the Australian Carboniferous Trilobita, to date, of errors which have undoubtedly become in- corporated therein. This aim is one not likely to be fully realised, but it will be my endeavour at least to lessen them, and, if I succeed in doing this, I shall be satisfied. That the present paper is free from errors is not one of my expectations, for with reference to my conclusions in respect to F. woodivardi, I am not by any means satisfied; and I shall await with some anxiety, the discovery of better petrifactions of the remains of this species, to make certain what its complete form actually 33 438 CARBONIFEROUS TRILOBITES OF AUSTRALIA, The discovery of the genus Brachymetopus in the Mount Morgan district will be of much assistance in establishing the geological age of that area, and also in correlating its sedimentary rocks with those of other localities. For reasons which will be obvious, the name Phillij^sia dubia has been discarded by me. To Messrs. W. S. Dun, Palaeontologist, Department of Mines, Sydney, B. Dunstan, Chief Government Geologist, Brisbane, and H. A. Longman, Director of the Queensland Museum, Brisbane, I am indebted for specimens placed with me for inspection and description, and for which I tender to them my thanks. To Mr. R. Etheridge, J. P., Director and Curator of the Australian Museum, and to Mr. Dun, my thanks are tendered for affording me access to literature. I am very grateful to General A. W. Vogdes, of San Diego, U.S. A, Bibliographist and Authorit}^ on Palaeozoic Crustacea, for supplying me with notes on the classifi- cation, and a list of the Carboniferous Trilobites described to date. Lastly, I wish to express my thanks to Mr. F. K. Cowper Heed, M.A., F.G.S., Acting Keeper of the Sedgwick Museum, Cambridge, England, for supplying me with a number of liis valuable papers, from which I have received much help. Bibliography of Australian Carboniferous Trilobites. (I). — 1847. Prof. McCoy (^.im. Mag. Nat. Hist., xx., p.231, PI. xii., fig. 1) described Brachymetopus strzehckii and referred to the occurrence of the genus Fhil/ipsia in Australian Carbonifer- ous rocks. (2). — 1872. R. Etheridge, Senr., (Quart. Journ. Geol. Soc, Vol. xxviii., p 338, PL xviii., fig.7) described Griffithides dubins, which, from the description, would appear to be an abnormal species. (3). — 1877. De Koninck (Foss. Pal. Nouv. Galles du Sud, Pt. 3, p. 348, etc., PI. 24, figs. 8, 9, 9a, 10, 10a, 106, and 10c) de- scribed and figured Griffithides (PhiUipsia) elchwaldi Fischer, Phillipsia (Griffithides) seminifera Phillips, and Brachymetopus strzeleckii McCoy. (4)._1892. R. Etheridge, Junr., (Geo), and Pal. Queensland and New Guinea, pp. 214-216, PI. xxi., figs.U-H; PI. viii., figs. BY JOHN MITCHEJ.L. 439 5, 6; and PI. xl., figs. 4, 5, 6) described the following: — PluUipsid dubia, P. woodwardi Eth. fil., PhiUipsia{h sp. ind., for wliich he suggested the specific name Griffithides fieininiferus vai*. (lus- tralasicd, from Queensland. Also in the same year [Mem Geol. Surv. N. S. Wales, Pal., No.5, Pt. ii., pp. 126-1 30, PI. xxi., figs. 1-4, 5 (and Text-fig. f)), 6, 7,8; Pi. xxii., figs. 12-15], the same author described PhiUipsia dubia, P. grandts, J'hillipsia spp. ind. (two, a and b), and Grijfithides, sp. ind. (5). -1903. F. R. C. Reed, M.A., F.G.S. (Geol. Mag., Dec. iv., Vol.x., No. 467, pp. 193-197) redescribed BracJtymetopus strzeleckii McCoy, and added valuable remarks on its generic and specific position. (6). — 1917. R. Etheridge, Junr., (Geol Surv. of Queensland, Publication No.260) made additional reference to the occurrence of PhiUipsia grandis in Queensland. (7). — Besides the above descriptions and determinations, R. Etheridge, Junr., referred a fragment of a pygidium from West Coerdawandy and the Yaltra Mtns., on the Gascoigne River, W^est Australia, to his species PhiUipsia grandis (MS.). Critical Review of the Carboniferous Trilobites of Australia, previously described. Order TRILOBITA. Family PROETID^. Genus Brachymetopus McCoy, 1847. 1. Brachymetopus strzeleckii McCoy, Ann. Mag. Nat. Hist., XX., 1847, p.231, PI. xii., fig.l. Since this species was described by McCoy, no specimens of it, or of any other member of the genus, appear to have been found by collectors, though on the occasion of the first find at Dun- vegan, N. S. Wales, quite a number seem to have been obtained. Genus P h i l l i p s i a Portlock, 1 843. To Prof. McCoy we owe the first reference to the occurrence of this genus in Australia {loc. cit. antea). 440 CARBONIFEROUS TRILOBITES OF AUSTRALIA, 2. Phillipsia eichwaldi De Kon., (non P. eichwaldi Fischer) Foss. Pal. Nouv. Galles du Sud, 1877, Pt.3, p.348, t.24, f.9. Under the name of Griffithides {Phillipsia) eichwaldi, De Koninck described a trilobite from Colo Colo, N.S„W., and de- termined it to be P. eichivaldi Fischer; but, as the writer points out further on, this determination was evidently incorrect. Mr. Etheridge, Junr., expressed a similar view (Etheridge, Junr., Mem. Geol. Surv. N. S. Wales, Pal. No.5, Pt. ii., 1892, p.l24). There is little doubt that the presence of this trilobite in Australia remains yet to be established. 3. Phillipsia dubia Etheridge, Junr., (non Griffithides diihins Etheridge, Senr.) Geol. and Pal. Queensland and New Guinea, 1892, pp.214-215, PL vii., fig.5; PL xliv., fig.4. Phillipsia dubia Etheridge, Junr., (non Griffithides dnhiiis Etheridge, Senr.) Mem. Geol. 8urv. N. S. Wales, Pal. No. 5, Pt. ii., 1892, p.l26, PL xxi., figs.1-4, PL xxii., figs.l2, 13. The above two forms were considered bv Mr. Etheridge, Junr., to be specifically identical with Griffi'thides dnhiiis Etheridge, Senr.; and the latter to be generically misplaced. The writer regrets that, with this determination, he cannot agree with Mr. Etheridge. It appears to him that neither of the above trilobites is specifically identical with Grijfithides duhius Eth. Senr., nor with each other. Mr. Etheridge, Junr., wlien describing the trilobite from Binge Berry, Rouchel Brook, New South Wales iloc. cit. antea) pointed out that it diftered from the Queensland form in having eighteen to twenty axial rings in the pygidium instead of eight to ten, which is the number in that of the Queensland form. This alone is suflficient to separate them specifically; but the head-shields of the two also differ. Both forms are here treated as separate species. The writer's view regarding the Queensland Phillipsia dubia is, that it cannot at present be generically or specifically joined with Griffithides dubius; and that the latter species must still be recognised. Only the discovery of a number of specimens of this form, show- ing the stages of development from the immature to the mature state, can satisfactorily establish the proper position generically and specifically of Griffithides dubius Eth. Senr. fiY JOHN MITCHELL. 44l 4. Phillipsia grandis Etheridge, Jurir., Geol. and Pal. Queens- land and New Guinea, 1892, p.215. Etheridge, Junr., Mem. Geol. 8urv. N. S Wales, Pal. No.o, Pt. ii., 1892, Text-6g.5, and Pi. xxi., fig. 5. Etheridge, Junr., Geol. Surv. Queensland, Pub- lication No.260, pp.11-12, PI. iii., fig 3. It is unfortunate that Mr. Etheridge placed tliese two forms specifically together. They are quite difterent, and are so treated by the writer.. In describing them, their differences will be fully shown. As the Queensland form was the one for which the specific name grandis was first suggested by Mr. Etheridge, it is proposed that it should continue to be known by this name. To tlie New South Wales one, a new specific term will be given. The name grandis is quite appropriate for either of the forms; because their pygidia are larger than those of any other known Carboniferous trilobite. 5. Phillipsia woodwardi Etheridge, Junr., Geol. and Pal. Queensland and New Guinea, 1892, p. 215, PI. 7, figs. 11, 13; PL 44, figs. 5, 6. This is a singular species, to which some reference will be made later on. 6. Besides the foregoing species of Phillipsia, which have been specifically determined, Mr. Etheridge described two pygidia from Binge Berry, Rouchel Brook, and near Paterson, respect- ively, and referred them to this genus, but did not give them specific rank. 7. (a) Phillipsia sp. ind. (a), Etheridge, Junr., Mem. Geol. Surv. N S. Wales, Pal. No. f., Pt. ii., 1892, p. 1 29, PL xxi , figs. 6-8. (b). Phillipsia sp. ind. (6), Etlieridge, Junr., op. cit., p. 129, PL xxiL, fig. 14. Both of these jvill be redescribed and named. Genus G R i f f i t h i d e s Portlock, 1843. 8. (^iRiFFiTHiDES DUBius Etheridge, Senr., Quart. Journ. Geol. Soc, 1872, xxviii., p.338, t.l8, f.7. 442 CARBONIFEROUS TRILOBITES OP AUSTRALIA, The writer is of the opinion that this species should still be recognised. Its description is the first record of the genus in Australian rocks. 9. Gkiffithides skminiferus De Koninck (non Pliillips, species) Foss. Pal. Nouv. Galles du Sud, 1877, Pt.3, p.348, t.24, f.9, 9a. That this determination on the part of ])e Koninck was in- correct, there is really no doubt. It is fully dealt with in the observations on the relationship of Phi/lipsia collitisi to other species, described further on. 10. Gkiffithides seminifeuus var, austkalasica Etheridge, Junr., Geol. and Pal. Queensland and New Guinea, 1892, p. 216, PI. vii., fig. 14. The material used by Mr. Etheridge for his description of the above is before me, and, after close examination of it, I am doubtful of the correctness of Mr Etheridge's conclusions. The pygidia, by which his conclusions were greatly influenced, have each thirteen and eleven axial and pleural divisions respectively; and, in this respect, agree with pygidia }>laced by me with his P. woodwardi. All these pygidia agree in tlieir ornamentation. As regards the cephalon associated on the specimen with these pygidia, it does not appear to differ in any essential from two of those included by Mr. Etheridge in his F. ivoodwardi, except that, in the latter, the granulation has been worn off by weather- ing. On the glabella of the former, it is true, no anterior or ftiesial glabellar furrows are visible, but this glabella is a very imperfect intaglio. If it should ultimately be shown that this fossil is a good species, it will not, even then, be closely related to Gr. semitiiferns, because it possesses supplementary basal lobes, and the latter does not. This difference certainly places them specifically apart. It may be stated that, so disposed was I to the opinion that this was a separate species, that I had written a description of it under a new name, but not being able to discern any difference in the pygidia now under discussion, and those I have placed with F. woodtvardi, I deferred final judgment. BY JOHN MITCHELr.. 443 11. Griffithides sweeti Etheridge, Junr., Proc. Linn. Soc. N. S. Wales, 1894, p. 528, PI. xxxix., f.3. The writer lias not seen the original specimen. Judging from the description and illustration, it is a very unusual type. See description further on. 12. Griffithides sp. ind., Etheridge, Junr., Mem. Geol. Surv. N. 8 Wales, Pal. No.5, Pt. ii., 1892, p.l30, PI. xxii., figs.lo, 16. It is not improbable that this may be identical with Phillipsia collhisi. From, the foregoing review, the following conclusions have been drawn, viz. : — i. That, of the nine species of Australian Carboniferous trilo- bites previously described and named, only the five following are worthy of recognition. 1. lirachymetojnis strzeleckii McCoy. 2. Griffithides dubius Eth. 8enr. 3. Griffithiiies sweeti Eth. Junr. 4. PhiUipsia icoodfvardi Eth. Junr., Geol. et Pal. Queens- land and New Guinea, p. 215, PI. 7, figs. 11, 13. b. PhillijJsia yrandis l^lh. iuin\, oi which the type is the Mt. Morgan form. Mem. Geol. Surv. N.S. Wales, Pal. No.5, Pt, ii., 1892, p. 128, PI. xxi., fig.5. ii. That the two determinations made by the late Prof. i)e Koiiinck are incorrect, and scientifically valueless. Family PROETID^ Steiiib. Genus Phillipsia Portlock, 1843. Dr. H. Woodward's* description of the genus is as follows: — "General form oval; glabella with nearly parallel sides, marked by either two or three pairs of short lateral furrows; the posterior angles, forming the basal lobes, always separated by a circular furrow from the rest of the glabella; eyes large, reniform, surface delicately faceted; cervical furrow deep; free cheeks separated from the glabella by the axal (facial) suture, which forms an acute angle with the circular border of the cheek in front of the * Moil. Brit. Carb. Trilobites, 1883-4, p. 11. 444 CARBONIFEROUS TRILOBITES OP AUSTRALIA, glabella; whilst the facial suture cuts obliquely across the pos- terior margin, just behind the eye, leaving a small pointed por- tion fixed to the glabella b}' the neck lobe; angles of the cheeks more or less produced, mart^in of head incurved, forming a striated and punctured rim. Thoracic segments nine in number, the axis distinctly marked off from the side-lobes or pleurae by the axal furrows; the abdomen or pygidium usually with a rounded border, the axis composed of from 12 to 18 coalesced segments." Phillipsia collinsi, sp.nov. (PI. xlvi., figs. 1-5; PI. xlviii., figs. 8, 9; PL lii., fig. 4). Sp. CVtars. — Complete form elongately ellipsoidal. Cephalon sub-semielliptic, moderately inflated, densely tuber- culated; glabella sub-oblong or sub-bellshaped, narrow, upper surface flat, steep and straight sided, densely crowded with pro- minent tubercles of variable sizes; medially the tubercles are arranged in four sublongitudinal rows, those on the anterior pari are subconcentric; front gently rounded and bearing a concentric row of tubercles; lateral lobes and furrows distinct, the basal pair altogether circumscribed, and bearing a varying number of tubercles, one usually much larger than the others; limb narrow, witli a gentle upward turn, smooth or very finely granulate; neck-furrow distinct, narrow, and deep behind the basal lobes; neck-ring strong, moderately arched and tuberculate, its lateral extensions being narrow; fixed cheeks very small, eye or palpe- bral lobes bounded by tubercles; eyes apparently crescentic; free cheeks unknown. Axial furrows distinct; facial sutures ante- riorly very gently sinuate (almost straight) and close to the axial furrows; posteriorly straight, and almost at right angles to the axial line, and at last cutting the fixed-cheeks with a sharp backward turn. TJiorax oblong, moderately inflated and granulated; axis pro- minent, the rings centrally and at the bases mildly curve forward; bases feebly tuberculated, width equal to that of one side-lobe, median sulci of the rings narrow; pleura? convex, medial furrows of each segment wide and shallow, and reaching the distal end. BY JOHN MITCHELL. 44 5 which is rounded; each posterior segmental ridge bears a row of tubercles, as du also the axial rings; axial furrows distinct. Pygidinvi semi-ellipsoidal, moderately tumid and strongly tubercuJate; proportion of length to width equals 3 : 4 approxi- mately; axis has fourteen rings, prominent, dorsally depressed, sides steep, contracting very gradually posteriorly to about half the anterior width, terminating bluntly and somewhat overhang- ing the border, the rings bearing tubercles which vary in number with their spread from two or more on the posterior ring, to eight on the anterior ones, and not exactly forming longitudinal rows; axial furrows pronounced; pleurae very convex, arching from the axial grooves at first gently and then steeply to the narrow furrow separating them from the border, where all the segments, except the anterior pair, end, segments ten in number, each succeeding one having a gently backward trend, and dimin- ishing in length till the last two pairs are represented by mere tubercles. Each segment bears a row of tubercles along the posterior ridge varying in number with its length, from one to ten, and perhaps in some instances more; medial segmental furrows wide and shallow, reaching to the faint furrow within the border; the border itself is fairly wide, and thickened, and- in normal specimens, has the same convexity as the pleura] ribs; it bears a row of conspicuous tubercles usually placed in line with each rib, and three to five behind the axis, these tubercles being occasionally elongated or double; when the thickened por- tion is removed, the undersurface shows concentric striations, which is a common feature in the Proetidte. Only the first pair of the pleural segments interrupts the continuity of the l)order. Ohs —This species is (juite singular. Its outstanding features are:— 1. The narrow, straight-sided glabella. 2. 'J'he crowded and conspicuous tuberculation uf the glabella. 3. The equality of the lengths of thepygidium and thorax. 4. The width of the thorax, which equals half the total length of the complete indi- vidual. 5. The closeness of the eyes to the axial groove. G. The straightness of the anterior branches of the facial sutures, and their parallelism to the axial furrows. Dimensions: total length of an individual not quite mature. 446 CARBONIFEROUS TRILOBITES OF AUSTRALIA one inch: widtli, half an inch; cephalon, one-quarter inch: thorax and pygidiuH), each three-eighths of an inch. In the proportions of length to width, and the character of the pvgidial granulation, this trilobite agrees with P. gemmu- lifera Phillips, with which it also agrees in the equality of the lengths of the thorax and pygidium. In other specific features, thev differ. In the cephalons and pygidia of F. collinsi and F. truncdtuld, there is much common to the two. They agree in possessing narrow, straight-sided and tul)erculate glabella, eyes close to the axial furrows, and similarly tuberculated pygidia: but the tuberculation on the glabella of the former is much more pronounced than it is on the latter, and the sinuate course of the anterior branches of the facial sutures of the latter is more pro- nounced than that of the former. Thepj^gidium of F truncatula has seventeen or eighteen annulations in the axis, and fourteen divisions in each pleura ; but in F. collinsi^ these parts number fourteen (or thirteen and a terminal piece), and ten respectively. In the former, the pygidium has no border, and the pleural ribs extend to the periphery; on the other hand, the latter species has a very distinct pygidial border, which, except in the case of the anterior pairs, the pleural ribs do not cross. The tubercles of the pygidial axis of the former are arranged in longitudinal rows; this is not quite so with those of the latter. Of the North American Phillipsif^, the nearest relative to ours appears to be P. insignis, which apparently has the same number of divisions in the axis and pleurae of the pygidium, as are in the similar parts of P. collinsi. The tuberculation of the axial rings is much alike in the two species. In other respects, they differ rather widely. The only Australian Carboniferous trilobite fragments bearing any resemblance to P. collinsi are two pygidia described and figured, but not named, by Mr. Etheridge, Junr.* The one pygidium he placed in the genus FhiUipsia {loc. cit., PI. xxii., fig. 14). It agrees with F. collinsi in possessing fourteen rings in the pygidial axis, and in having a tuberculate test; but differs in having a continuous, smooth, flattened border, and '" R. Etheridge, Junr., Mem. Geol. 8urv. N. 8. Wales, Pal. No.5, Pt. ii., 1892, pp. 129-130, PI. xxii., figs. 14, 15. BY JOHN MITCHKLL. 447 twelve pleural segments, and less disparity between length and width of the pygidium. The pygidium {loc. cit., PI. xxii., fig. 15) placed by Mr Ether- idge in the genus Grijfithides, bears a resemblance to the present species in the character of its border and granulation, and in having the same number of pleural segments; but its axis has less rings, and ends in a sharp point. Lastly, referring to De Koninck's P. {Grijjithides) seinhiifera {noil Phillips sp.)* from Colo Colo. The dimensions, number of rings in the axis of the pygidium, character of the granulation generally on the pygidium and thorax agree closely with similar features of F. collinsi: but the two forms widely differ in the outlines of their glabellse and pygidia, if De Koninck's figures are to be relied upon, which is doubtful, as they do not agree with the text. In the former, the glabella is represented as being conical in outline; and, in the latter, it is said to have an anterior width slightly less than that of its base. Whatever Phillipsia [Grijfithides) seminifera De Koninck, may have been, it is practically certain it was not Grijjithides seminiferus Phillips; but it may have been identical with the present species. Named after Mr. Collins, C.C.M., Lecturer in Coalmining and Mine Surveying at the Newcastle Technical College. Loc. and H or. — Glen AVilliam Road, one mile from Clarence- town, Parish Parr, County Durham. Lower(?) Carboniferous. Philijpsia coulteri, sp.nov. (Plate xlvi., figs 6-10;. Sjo. Chars. — Complete form subelliptic. Cej^halon sub-semielliptic, fairly inflated, finely granulated : glabella subrectangular, rounded in front, lateral furrows and lobes not visible, owing to the loss of the part bearing them, and the exposure of the hypostome; neck-furrow shallow, its lateral extensions across the cheek.s shallow and wide; neck-ring narrow; axial furrows faint. Fixed cheeks small. Free cheeks relatively large, strongly inflated, steep between the eye and border-furrow, which is wide and shallow, outer edge of border only moderately * Pal. Foss. Nouv. Galles du Sud, 1877, p.'267, PI. xxix., figs. 9, 9m. 448 CARBONIFEROUS TRILOBlTfiS Of AUSTRALIA, thickened. Eyes prominent, large, crescentic and faceted. Genal angles rounded. Hypostome alate, shield like, and striated in tiie fashion common to Phillipsise. Thoi'dx 11 mm. wide and 9 mm. long, appearing to have been finely granulated, with a few scattered tubercles; straight-sided. Axis very strongly and acutely arched, rings, centrally and basally, strongh^ inclined anteriorly, each one bearing along the medial line a rath--r prominent tubercle, giving the axis through- out a serrated character: spread equal to that of one side-lobe, mildly tuberculate, height and width diminishing gradualh* pos- teriorly, axial grooves faint. Pleurae gently rising from the axial grooves to the fulcra, thence steeply deflected, mesial fur- rows of the somites shallow and wide, and reaching the extremi- ties, which are rounded: along the fulcral line each rib bears a tubercle on the posterior ridge, and there is evidence of the presence of general fine tuberculation or granulation. Pygidium semielliptic, length 8 mm., width 10 mm. Axis very prominent, the first ring having a medial forward trend, rings eight or nine, counting the end-piece, which is buttress- like, and ends at the border in a rather fine point, and bears a subconspicuous tubercle. Pleuriie with seven divisions, all rather inconspicuous in the specimen (decorticated) serving for this de- scription, moderately convex; mesial furrows very shallow and reaching to the faint furrow within the border, each segment bearing a tubercle at the fulcral angle, and showing traces of other tuberculation, as do also the axial rings; the border is relatively wide, steep, gently thickened, and bounded inwardly by a faint furrow, and is crossed only by the first pair of ribs. (9^^^ —Though the glabellar furrows and lobes are not visible in the specimen described, the shape of the glabella and character of the hypostome leave no doubt as to its being a Phillipsia, and the specimen is one of the finest of this genus yet discovered in Australia. It difiers so widely from other Australian species that there is no need to detail its relationship. It bears no close relationship to any of the American Phillipsiai referi-ed to in Brigadier General A. W. Vogdes' Monograph.* * " The Genera and Species of North American Carb. Trilobites,"' Amialb N. Y. Acad. Sci., 1888, vi. BY JOHN MITCHELL. 449 The species has been named after Mr, E. Coulter, of Katoomba, an enthusiastic student of geology, who discovered and presented the specimen to the author. Loc. and /To?*. — Stroud-Dungog Road, near Dungog, Parish Howick, County Durham. Lower Carboniferous. Phillipsia brkvickps, sp.nov. (Plate xlvi., figs.ll, 12; and Plate li., fig.2). S]). CAars. — Complete form unknown. Cephalon apparently subsemicireular, finely granulated. Gla- bella straight-sided, rounded in front, very gently convex, highest just in front of the neck-furrow, smooth, though a lens shows evidence of microscopic granulation, two pairs of lateral furrows visible, basal and mesial. 'J'he former very shallow and circumscribing the basal lobes, which are large and pyriform; the other narrow; neck-furrow narrow and shallow; neck-ring relatively wide, decidedly arched back ward ly and lobed at its l)ases; frontal limb narrow and gently recurved. Fixed cheeks very small, eye-lobes small, and abutting the axial furrows. Facial sutures anteriorly rather straight and close to the axial furrows; eyes of moderate size only, judging by the size of the palebral lobes. Thorax unknown. Pygidium sub-semielliptical, moderately convex, very fineh' granulated; width 10 mm., and length 7 mm. respectively. Axis prominent, consisting of thirteen rings, and a very narrow end- piece, spread equal to that of each side-lobe, contracting very gradually posteriorly, and terminating short of the border, rounded and bluntly, with half its anterior width; some of the rings bear very fine tubercles, barely distinguishable without the aid of a lens; side-lobes moderately convex, with ten, or doubtfully eleven, pairs of segments, of which the medial furrows are distinct, and, in the case of the four anterior pairs, at least reach the edge and interrupt the narrow border; a few of the posterior segments bear very small tubercles at their junction with the border, and three similar tubercles occur behind the axis, on the axis itself centrally the posterior ridges also bear fine tubercles. 450 CARBONIFEROUS TRILOBITES OF AUSTRALIA, 6)6s. — Aswill be apparent from a comparison of the above description with that of P. derbieitsis Martin, the two fossils agi-ee in several important features. For example, in both the glabella is parallel-sided and practically smooth, anterior facial sutures rather straight and close to the axial furrows, eyes reni- form, pygidial axis with thirteen rings, and ten segments in the pleur?e; and the pygidial borders obsolescing before reaching the articulating face; in the equality of maximum widths of the axes and pleurae of the pygidia (this is only true in the case of the above species when the pleurae are measured obliquely, and not the actual horizontal widths: if measured in the latter way, the axis is wider than one pleura) and the faint tendency to orna- mentation on these parts. Against these resemblances, there are the following differences to be considered. The length of the glabella of the local species, measured from the neck-furro\v to its front, is equal to its width between the axial furrows across the basal lobes; in that of J\ derbiensis, these dimensions are respectively about 4 : 2. The eye-lobes are shorter, and the frontal border wider, neck-ring more intensely arched posteriorly in the local form than in the British one. The distinct lobation of the bases of the neck-ring, relatively short cephalon, and the almost equality of the lengths of the cephalon and tail are very marked features in the local species, separating it from the other. Then tliere is the distinct, though fine, granulation, more or less tuberculation of the pygidium of the form now described, as against a practically smooth one in the other. Further, judging from the anterior pair of pleural segments of the pygidium, the thoracic pleural ribs were not imbricated, in the case of the local species. Although undoubtedly closely related to each other, I have concluded that the differences between them are sufficient to justify their separation specifically. The glabellie of the above species and Proetus inissourieiisis are much alike. I am indebted to J^rigadier General Vogdes' paper [op. cit., antea) for the references made to American Car- boniferous trilobites. Lvc. and Uor. — Neighbourhood of Port Stephens, probably from near Bulladelah. Carboniferous. BY JOHN MITCHELL. 451 Philltpsia proxima, sp.nov. (Plate xlvi., figs.lf), 16). Sp. Chars. — Pygidinm semielliptical, fairly convex, practically smooth, so fine is the granulation. Length 12 mm., width 15 mm. Axis strongly convex, rings twenty, tapering posteriorly very gradually, and ending bluntly at the border, with a spread of about half that of the anterior ring; axial furrow shallow, but distinct. Pleurje gently convex, consisting of twelve pairs of ribs, the last one or two very shoit and faint, all terminating at the border, which is continuous to the articulating face and sub- depressed: medial furrows of the segments, except in the cases of a few of the posterior pairs, well defined, anchylosing ridges also fairly prominent; furrow separating border and pleural ribs distinct. Ohs. — This p3"gidium agrees with that of PhiUipsia eiehivaldi Fischer, in the continuous smooth and somewhat depressed border, and the character of the pleural segments. The most important differences between them are— (1) the pygidium of F. eichivaldi is just as wide as long, is distinctly granulated, and the distal axial end does not seem to be prominent. The pygidium above-described has a length only four-fifths of its width, and is practically smooth. The total length of the pygidia of mature individuals of the former, as shown by the beautiful figures in Dr. H. Woodward's work* is 9-5 mm., that of the latter is 12 mm. The large number of rings in the axis of ours places it far apart from the other. Named because of its close resemblance to P. eichivaldi Fischer, in shape. Loc. and H or. — Glen William Road, near Clarencetown, Parish Parr, County Durham. Lower Carboniferous. Phillipsia('?) robusta, sp.nov. (Plate xlvii., figs.l and 8). Phillipsia graridis Eth. fil., Mem. Geol. Surv. N. S. Wales, No. 5, Pt. ii., 1892, p.l28, text-fig.5 {non PI. xxi., fig.5). Sp. Chars. — Cephaloji and thorax unknown. Pygidium subsemicircular, strongly convex, very finely granu- * Mon. Brit. Carb. Trilobites, 1883-4, PI. iv., figs. 9 and 1.3. 452 CARBONIFEROUS TRILOBITKS OF AUSTRALIA, late, greatest width 44 mm., length 26 mm., (the width here given is what the writer considers to be the normal, and is calculated from the width of the left side-lobe and half the spread c)f the axis, because the right side-lobe is distorted). Axis fairly pro- minent, evenly arched without any decided backward or forward inclination of the annulations, of which there are fourteen: the furrows of these are wide and V-shaped in section, ridges high, edged, and bore some small tubercles, bases not tuberculate. Axial furrows shallow. Pleurae very convex, arching strongly from the axial furrows, and showing no decided fulcra; segmental divisions twelve pairs, which reach to the border, and are only wently directed posteriorly, except in the case of the last two pairs; their mesial furrows are V-shaped, wide, and reach to the border, ridges high and chisel-edged, and show no sutures, hence the pleural segments in this oygidium were very completely fused: some of the ridges exhibit evidence of very fine and sparse tuberculation, anterior pair facetti. Border continuous, wide, steep, and its undersurface concentri -ally striated. Qf)s. — 11iis pygidium was described and figured by R. Etheridge Junr.,* and considered by him to be identical with a pygidial fragment from near Mt. Morgan, Queensland, and for which he had previously suggested the name Fhillipsia gi-andis.j Mr. Etheridge figured this Queensland specimen, when dealing with some New South Wales trilobites.| The writer is unable to accept Mr. Etheridge's determination of the identity of these two pygidia, for the following reasons. 1. The part of the pleural lobe preserved on the Queensland pygidium shows fifteen segments, and, when complete, without doubt had several others, as against twelve in the New South Wales one. 2. Though the Queensland tail, when complete, was apparently longer than that of the New South Wales specimen, the greatest Midth of its pleural lobes was only 10 mm., as against 15 mm. for that of " Mem. Geol. Survey N. S. Wales, Pal. No.o, Pt. ii., 1892, p. 128, text- fig. 5 {uoii PI. xxi., fig. 5). t Jack and Etheridge, Cleol. and Pal. of Queensland and New Guinea, 1892, pp.2irj, 216. X Mem, Geol. Survey N. S. Wales, Pal. No.5, Pt. ii., 1892, PI. xxi., fig.o. BY JOHN MITCHELL 453 the latter. 3. In the Queensland one, the pleural segments along the ridges are separated by very distinct sutures; in the other, this is not so, 4. In the Queensland one, the border is not nearly continuous, for the three anterior pairs of pleural segments reach to the edge of the lobes, and these are the only ones which are perfect on the portion of the left and pleural lobe of the specimen. 5. The mesial furrows and ridges of the pleural segments are not alike in the two. 6. In the Queensland tail, there were not less than sixteen or more axial rings, but fourteen only in that of the other. 7. The axial grooves are deeper in the specimen from Queensland, than in the one from New South Wales. The two agree in the character of their granulations; but that of the Queensland one is the coarser. That the Queensland pygidial fragment represents a new species of either Phillipsia or Griffithides. there is no doubt; and, to it, my proposition is to apply the name Phillipsia grandis, first suggested for it by my friend, Mr. Etheridge. P.i'ohnsia differs from all species of this genus, or of Griffith- ides known to me. Mr. Etheridge, too, noted (loc. cit.) how different it was from foreign Carboniferous trilobites. Loc. and Hor. — Swain's Conditional Purchase, seven miles south-east of Carroll, County Buckland (D. A. Porter). Car-, boniferous. Phillipsia(?) stroui>e;nsis, sp.nov. (Plate xlvi., fig.14; Plate li., fig.ll), Sp. Chars.-^Cephalon and thorax unknown. Pygidium semielliptic, densely and finely granulate, moder- ately convex; greatest width 17 mm., length 15 mm. Axis only moderately convex. Consists of twenty annulations, a few of the anterior of which have a very gentle forward inclination, centrally and basally ; contraction posteriorly very gradual, terminating bluntly, and rounded at the border, with half the anterior width (3 mm.), medial furrows of the rings wide and shallow; anchylosing ridges not very prominent, but distinctly granulated, anterior spread slightly less than that of one side- lobe. Axial furrows faint. Pleurae moderately convex, seg- ments fifteen, the last faint, anterior pair faceted, all finely and 34 454 CARBONIFEROUS TRILOBITES OF AUSTRALIA, distinctly granulate, reaching to the border, and gently inclined backward (falcate). Medial furrows wide and shallow, reaching the border, ridges low. Border continuous, narrow anteriorly, gently increasing in width posteriorly, flat, densely and finely granulate. O65. — This is a fairly large pygidium, as the dimensions given will show; but it is, in this respect, exceeded by both of the forms placed by Mr. R. Etheridge Junr., under the name of P. yrandifi; but it has no specific relations with either of them. In the number of axial rings, it agrees with P. elongata, but not in other features. It is different from all the British Phillipsise the writer has been able to compare it with; and the same is true as far as the North American species are concerned. Log and Hot. — Stroud, County Gloucester, N.S.W., associated with Orthis resupiiiata and Spirifer striattis. Lower(?) Car- boniferous. Phillipsia superba, sp.nov. (Plate xlviii., fig. 15: Plate Hi., fig.3). Complete form unknown. Sp. Chars.— (7. Char. Glabella round, without any lateral inflection of the margin, moderately convex in the middle line, and a little arched posteriorly; neck furrow strong and deep, with more or less com- plete basal furrows; anterior furrows present, but faint; anterior border thick and upwardly turned, leaving a wide depression between it and the front of the glabella." In his observations, Mr. Etheridge remarks: — "The pygidium referred to is much larger than that described as P. dubia, but otherwise resembles it." In the text, there is nothing to show what pygidium is here referred to, for no description of it is given; but no doubt he refers to the pygidium illustrated in Plate xliv., fig. 6; and if so, the determination, in my opinion, is incorrect. This pygidium belongs to the trilobite described by me as Phillipsia rockhamptonensis, and is shown in PI. 1., fig.8. The following is a more amplified description of the type- cephalon of the species. Outline subsemicircular, surface gran- ulated. Glabella wider across the basal lobes than it is long, very convex both transversely and longitudinally, the front lobe (the portion anterior to the anterior pair of the lateral furrows) semicircular, highest between the mesial pair of lateral furrows. Three pairs of lateral furrows present, anterior and mesial rela- tively wide, shallow and straight, basal pair wide and deep; anterior and mesial lobes narrow, basal pair large, subfusiform, much lower than the glabellar centre, overhanging the axial and neck-furrows; limb very large, border strongly thickened and upturned, furrow wide and deep. Neck-furrow deep; neck-ring strong, convex, sharply curving anteriorly, as it approaches the axial furrows; sides gradually thickening towards their origin. Frontal limb very wide, border strongly thickened and upturned, and separated from the glabella by a wide and pronounced furrow, strongly expanded antero-laterally. Facial sutures anteriorly directed outwards at an angle of about 25°. Fixed BY JOHN MITCHELL. 467 cheeks appear to have been small, and strongly tumid. Axial furrows deep. Other parts not known. As Mr. Etheridge pointed out, this head-shield bears a strong resemblance to that of some Proeti. The enormous limb is its most striking feature. The discovery of a complete specimen will be of much scientific interest. It may be noticed here that neither of the figures, PL vii., fig. 13, PL xliv., figs. 5, 6 {loc. cit.) are correct representations of the fossils they are intended to depict. For example, in the case of the first of these, the glabella is much too blunt in front; and supplementary lobes, which are present on the fossil, are not shown in the figure. With reference to figs. 5 and 6 of PL xliv., in the case of fig. 5, the glabella is represented as of a battle-axe shape, instead of being rather quadrate, or only gently rounded in front, and the glabellar basal lobes are exaggerated in size. In fig. 6, the axis is shown terminating much too short of the border, and, in the pleural lobes, too many segments are shown. Photographs of the original specimens, from which the drawings of the figures above referred to were made, will be found in the Plates of this paper. At present, I hesitate to recognise the portion of a cephalon determined by Mr, Etheridge to belong to this species, and re- presented in PL vii., fig. 13, for the following reasons. The glabella of this specimen is much more convex and narrower in front than the type-specimen. Further, this glabella is just as long as wide across the basal glabellar lobes. On the other hand, the typical glabella of the species is much wider across these same lobes than it is long. The glabella in question is not correctly represented by the figure above referred to, as will be seen from the photos of it now produced. Since Mr. Etheridge described this species, much additional material has been secured, and this has been placed with me by Mr. Dunstan, Chief Government Geologist, Brisbane, for classi- fication. Included in the collection is a large number of pygidia, all possessing similar specific features; and these were collected from the same localities from which all the known cephalic frag- ments of the species have been obtained. In pait, owing to the 468 CARBONIFEROUS TRILOBITES OF AUSTRALIA, association in this way of these heads and tails, and because the great convexity of the pygidia referred to, which would indicate the possession of a very convex cephalon by the species they belong to, and, further, because no other head-shields reasonably preserved occur in association with these remains, with whicli they appear to have near relationship, I liave determined to place them with Mr. Etheridge's }\ tvoodwardi, and, for this reason, the species is redescribed hereunder. Complete form not yet certainly known, but it may be assumed to have been oblong-ovate. Cephalon sub-semicircular, highly inflated, surface granulated throughout. Glabella subquadrate, wide and mildly rounded in front, strongly convex, highest between the mesial lobes, sloping strongly anteriorly, and overhanging the furrow of the limb in front; three pairs of glabellar furrows present, the anterior and mesial pairs rather wide, shallow and straight; basal pair wide and deep; anterior and mesial lobes narrow; basal pair large, subfusiform, much lower than the glabellar superficial centre, overhanging the axial and neck-furrows latero-posteriorly; limb very large, border strongly thickened and upturned, furrow wide and deep; neck-furrow wide and deep; neck-ring strong, convex, bases strongly curving anteriorly under the glabellar basal lobes; supplementary lobes present, though tjot conspicuous in the most typical specimen. Facial sutures anteriorly directed outwards at an angle of about 25°. Fixed cheeks would appear to have been of moderate size, and posteriorly strongly inflated. Free cheeks fairly large, strongly and thickly granulated, especially on the part between the lateral and ocular furrows; this part, too, is high, and rather acutely angulate postero-laterally, flat superficially, laterally and posteriorly steeply falling into the lateral and posterior furrows, which are wide and deep; border much thickened and upturned, striated, posterior border (exten- sion of the neck-ring) strong, genal angles apparently blunt; eyes of moderate size, apparently crescentic and faceted, some- what sunk into the ocular furrow. I'horax : for certain not known, but the following is the de- scription of one attached to a pygidium identical with a number BY JOHN MITCHKLL. 469 of others assumed to belong to this species : suboblong, gi-aii- ulated, longer than the p\^gidium, all the segmental ridges appear to have borne tubercles; number of segments normal. Axis prominent, ring-bases non-tuberculate, with a strong anterior inclination centrally, ridges and valleys well defined, apparently as wide as one side-lobe. Side-lobes strongly convex and steep between the fulcra and margins, all parts densely granulated, segments anteriorly angulate at the fulcra; mesial furrows wide, ridges relatively narrow and faceted, and mildly thickened posteriorly. Pygidivm very strongly convex, length to width approximately 2 : 3, granulated; axis prominent, evenly arched, anterior spread about equal to the greatest width of one side-lobe, posteriorly diminishing in width very gradually, and ending prominently and bluntly a little short of the bordei-; number of rings thirteen, and each bore a row of small tubercles quite conspicuous on unweathered specimens; axial grooves wide and deep. Side, lobes convex, consif^ting of eleven pairs of segments, the lidges of each pair bearing rows of small tubercles varying from two or three on the shorter, to eight or perhaps nine on the longer ones. Border continuous, mildly thickened, convex and steep. Ohs. — At the outset, I admit that the pygidia here joined with Mr. Etheridge's tj'pe-cephalon of the species may yet prove to be a wrong conclusion; but should it happen so, T am inclined to believe, too, that the heads (PI. li., figs. 1*2, 13) will also prove not identical with the type one; but should that happen, I am fully satisfied that these latter heads, and the tails here dealt with, will prove to ijelong to the one species. The heads now joined with the typical one possessing the very characteristic frontal limb, cannot, for certain, in the absence of this limb, be said to belong to the species. While completing the above description, the specimen No.712, belonging to the Queensland Museum, Brisbane, is before me; and the trilobite fossil remains on it consist of two pygidia, one thorax and pygidium conjoined but incomplete, a fragmentary head in intaglio, and a part of a free cheek. (On the same specimen, occurs the tail of a Brachymetopus, the first recorded 35 470 CARBONIFEROUS TRILOBITES OF AUSTRALIA, from Queensland). These fossils weie those which chiefly served Mr. Etheridge to found his varietal species Griffithides aemini- ferus var. australasica on. After a close study of these remains, I am compelled to dissent from his conclusions concerning them. They cannot belong to the above species, because the glabella possesses supplementary basal lobes, a feature by some palaeon- tologists thought to be of generic significance, and which G. seminiferus does not possess. Again, these pygidia have thirteen rings in their axis, and eleven pairs of segments in the pleursp. In these parts, G. seminiferus has twelve and nine respectively. These differences alone would render the specific identity of the two forms invalid, but these are not the only differences between them. As far, then, as the fossils under present consideration afford evidence, Mr. Etheridge's varietal species fails. The part of the cephalon on this same specimen of rock is very imperfect, but it shows, on the glabella, similar but clearer granulation, because the granules are not worn off by weathering, similar glabellar basal and supplementary lobes lo that and those respectively found on the cephalons here determined to belong to Phillipsia vjoodwardi. The only features not visible on the glabella of the fragmentary cephalon, to make its identity with the latter certain, are the mesial and anterior glabellar furrows; but these are barely visible on some glabellse of P. woodwardi. Lastly, the pygidium represented in Plate viii., fig. 6 (oj). cit.), which was considered by Mr. Etheridge to be a normal pygidium of his P. duhia (P. stanvellensis mihi) is here joined with the present species by me. It is undoubtedly inseparable from the other pygidia now joined with the species above described, and certainly is not at all closely related to the pygidia of P. stan- vellensis mihi [P. duhia Eth. fil., op. cit., PI. viii., fig.o). Mr. Etheridge has already called attention to the Proetus-like aspect of the cephalon of the present species. The singular form of this part of the fossil makes it unnecessary to point out in detail the features which separate it from other species of Phillipsia. Log. and Hor. — Stoney Creek, Stanwell; Crow's Nest and Trilobite Ridge, Mt. Morgan, Queensland, etc. Associated wit |i BY JOHN MITCHELL. 471 Brachymelopus dniififani. Middle or Lower Carboniferous pro- bably. Phillipsia morganensis, sp. nov. (PI. xlix., figs.ll, 12; PI. li., 6g.l). .S';;. CAa/-6\ — 6V/?/irt/o?i sub-semicircular, very finely granulated, greatest length and width 8 mm., and 16 mm., respectively. Glabella subconical or sub-bellshaped, mildly convex, finely and densely granulated, longer than wide ; three pairs of lateral furrows distinctly shown, basal pair very deep and wide, anterior and mesial pairs short and only gently curved posteriorly: anterior and mesial lobes of moderate size, basal pair fairly large, with small complementary lobes present; neck-furrow deep and wide, the extensions laterally also fairly deep and wide; neck-ring strong, moderately convex, strongly curving anteriorly at its bases; facial sutures anteriorly very straight, posteriorly short and oblique, frontal limb narrow and close to the glabellar front; fixed cheeks very small, with small eye-lobes. Free cheeks rather large, only moderately tumid, finely granulated, both borders and lateral furrows large, the former being moderately thickened, finely granulated on the upper and striated on the undersurface, the latter being shallow, ocular furrow wide and shallow. Eyes of moderate size, finely faceted, vertically narrow, and decidedly higher behind than in front. Genal angles obtuse. Hypostome shield- like, narrow, not alate, posteriorly glossiform and corrugated, no striae visible, widest at two- thirds of its length from the front edge. Thorax not known in a complete state, but one, on which a pygidium and a free cheek rest, shows the following features — moderately convex, finely granulated. Axis moderately convex, diminishing in width and height gradually posteriorly, wider than one side-lobe, rings strong, valleys narrow and having an anterior inclination, bases non-tuberculate, axial furrows linear. Side-lobes not well shown, but the segmental ridges and furrows were strong and deep respectively. Pygidium widely triangular, mildly convex and granular, length 7 mm., and greatest width 12 mm. Axis mildly and evenly convex, anterior width equal to that of one side-lobe, 472 CARBONIFEROUS TRILOBITES OF AUSTRALIA, diminishing gradually in width and height posteriorly, and end- ing short of the margin inconspicuously with about half its anterior width, rings apparently eleven; ridges not prominent, furrows also faint, some of the anterior ones have centrally a backward trend. Axial furrows linear. Side-lobes convex, con- sisting apparently of nine pairs of segments (eight only are visible on the best available specimen), front pair faceted, ridges and furrows fairly distinct, and having successivelv onlv a moderate backward trend. Obs. — This trilobite belongs to the ty^pe oi Phillipsia which had the glabella narrower in front than posteriorly, which seems a common characteristic among the Australian members of the genus, and, in this respect, shows a closer relationship to the typical genus of the Proetidse, than do the Phillipsiie of Europe, and, perhaps also, America. A singular feature in the present species is the form of the hypostome assumed to belong to it. None similar to it has come under my notice. The one nearest to it is one figured by Dr. H. Woodward* with other illus- trations of Phillipsia eichivaJdi, but evidently not belonging to that species. 'J he straightness of the anterior branches of the facial sutures, and their mild outward divergence as tiiey reach the frontal limb, are striking features of the species. A study of the description and illustrations of the species will render its separation from other Australian Phillipsise simple. In contour, the glabella is not unlike what that of the glabella placed with P. woodtvardi (PI. xlix., fig. 7) would be, were it not so convex; but, in the former, the glabella is longer than it is wide across the basal glabellar lobes; in the latter, these dimensions are equal. No foreign species has come under my notice witli which it seems necessary to compare it. Loc. and Hor. — Trilobite Ridge, Mount Morgan, Queensland. Carboniferous (Lower'?). Phillipsia connollii, sp.nov. (PI. li., figs.4-7). Phillipsia grandis Eth. fil. (MS.). Only an incomplete pygidium known. * Mon. Brit. Carb. Trilobites, 1883-4, PI. iv., fig. 7. 6Y JOHN Mitchell. 4/3 Sp. Chars. — Pyyidiuiu semielliptic, moderately convex, and microscopically granulated. Axis very prominent, consisting of ten, possibly eleven rings, not as wide as one side-lobe, evenly arched, and slightly flattened dorsally, ending at the border very prominently and rather acutely, ridges and valleys of the rings quite distinct. Axial grooves shallow. Side-lobes mildly convex, consisting of eight or nine pairs of segments, -which are only very gently directed backward, and terminating at the border, except in the case of the anterior pair, which encroaches on it. Mesial furrows and the ridges of the segments moderately defined. Border proportionately wide and continuous, mildly convex, horizontal, and apparently wider laterally than behind, separated from the ribs by a shallow furrow. Greatest width 13 mm., length 9 mm., anterior width of axis 3 mm. Obs. — Thispygidium was placed by Mr. Etheridge, Junr., with his Fhillipsia grandis, but to this species I find it has no close relationship. In its wide border, and the number of axial rings and pleural segments, it bears strong resemblances to our Gri(tithides conveoncaudatus, but its axis is much more prominent and relatively narrower, than is that of the latter. The axial and pleural divisions are more clearly defined in the former than they are in the latter; the border in the former is ilat, and in the latter very steep. The shape or outline of the pygidia of these species is very different. In number of axial and pleural divisions, this species is related to both i". cordteri and /-*. staii- reUensls, but not otherwise. In possessing very prominent and relatively nai-row axes, P. stanvellensis and the present species agree. I am not aware of any foreign Carboniferous trilobite having a pygidium closely resembling this one. Named after Mr. Connolly, wlio discovered it. Loc. and llor. — Gascoigne River, AN'est Australia (Connolh'). Carboniferous. Phillipsia kouchklensis, sp.nov. PkUiipsui sp., ind. {a), Eth. til., Mem. Geol. Surv. N. S. Wale.s, Pal. No.5, Pt. ii., 1892, p.l29, PL xxi., tigs. 6, 7. (PI. xlviii., figs. 4, 5, 6, 7). 474 CARBONIFEROUS TRILOBITES OF AUSTRALIA, Cephaloii and thorax unknown. Sp. Chars. — Py^ice Ivoninck states that his specimen has only eight thoiacic segments, that tlie anterior annulation of the thoracic axis has a widthf of 3 mm., and not one of the annulations has a width less than 1 mm. The present specimen has the normal nine somites, and the widest annulation of the thoracic axis does not exceed 1 mm. De Konincks figure of his specimen shows it to have had a \'ery globular frontal glabellar lobe, and, in this, differs from the one above described. Again, De Konincks text does not agree with his illustiation. The former indicates his speci- nien to have eight thoracic somites, and the pleural lobes to have a width equal to that of the thoiacic axis; but his figure shows nine somites, and pleural lobes much narrower than the axis. These important discrepancies make it impossible to accept his " Foss. Pal. Nouv. (dalles du Sud, Pt. i., 1870, pp.278-9, PI. xxiv., tig.8. I The writer assumes the width of tlie axial rings to be their measure- ment alonj.; the lonifjtudinal line of the axis. 478 CARBONIFEROUS TRILOBITES OF AUSTRALIA, description as having any scientiHc value, for there is no evidence to show whether his text or figure is correct, the tj^pe specimen having been destroyed in the Garden Palace fire of 1882. The next consideration is whether the present fossil, or even De Koninck's, is specifically identical with P. eichwaldi Fischer. Accepting the types given by Dr. H. Woodward* as fully reliable, the author does not believe it a difficult task to prove the aegative. 1. Dimensions. — The relative lengths to widths of the parts of the author's fossil and that of P. eichwaldi Fischer, are, respect- ively, for cephalon 14 : 28 and 17 : 27; thorax 17 : 28 and 17 : 31 ; pygidium 9:13 and 1:1. The dimensional differences, as these measurements show, place the fossils apart; but these are not nearly so important specifi- cally as the following. The pygidium of F. eichwaldi is semi- elliptical, and the length equal to the width: that of the author's is sub-semicircular; that is, if the centre of the junction of the second axial ring with the third be taken for centre, the portion of the pygidium posterior to this forms a semicircle; and the length is only two- thirds of the width, approximately. There are up to sixteen rings in the axis, and twelve to fourteen pleural divisions in the pygidium of P. eichwaldi; while, in the local one, these divisions are eleven and eight, respectively. Then, in the former, the pygidial border is depressed; in the latter, tlie curve of convexity of the pleurae continues uninterruptedly across the border to the outer edge. In the former, also, the genal spines are long; in the latter, very short, only extending past the first thoracic segment. These differences are sufficient to prove that the author's specimen is not F. eichwaldi Fischer, and the same conclusion may be drawn in respect to De Konincks fossil. In several respects, this species resembles G'rij/ithides ylobiceps Phillips; but the proportionate lengths of the thorax and pygidium of the latter are different from those of the former; as also is the proportionate length of the cephalon to these same parts. In the former, there is no lobe connecting the eye-lobe with the * Mon. British Trilobites, Pt. i., 1883. BY JOHN MITCHELL. 479 glabella, though, indeed, tlie eye or palpebral lobe is very close to, and only faintly separated from, the basal lobe of the glabella. The eye, too, is relatively larger, the pygidial border is wider, and the glabellar frontal lobe less globular ; the free cheek- border and glabellar limb narrower, and the eyes less prominent and more posteriorly situated in the former than in the latter. Tiie British species, too, was much larger than the New South Wales one. The specific name was chosen for it because of the strong con- vexity of its pygidium. Log. and I/or. —Glen WilUam, about two miles from Clarence- town, Parish Parr, County Durham. Lower Carboniferous. Griffithides sweeti Eth. fil. Grijfithides siveeti Eth. fil., Proc. Linn. Soc. N. S. Wales, (2), Vol. ix., Pt.3, pp.5-28-9, PI. xxxix., fig.3, 1894. (PI. liii, figs.1,2). Through the courtesy of Mr. G. Sweet, of Melbourne, the type-specimen of the species is before me, and I regret that, owing to the incompleteness of its cephalon, it fails to elucidate certain difficulties met with in considering the thoraces and pygidia from the Mt. Morgan area, assumed to belong to PhiUipaia woodtrardi Eth. fil., or to settle whether G. siveeti Eth. fil., and that species are not identical. All the numerous pygidia obtained from the Mt. Morgan area, except those which belong to P. siaitvellensis, P. rockltatnptonensis, and P. inoi-yaiietLsis. have thirteen axial, and eleven pleural divisions; and, aftermost careful and repeated inspection, 1 have to conclude that Mr. Etheridges G. sweeti has the same number of divisions in the axis and pleurie of its pygidium; also the same kind of pygidial border and furrow: and, in fact, its pygidium cannot be sepa- rated from pygidia which are the most plentiful in the Mt. Morgan distiict, and which I have tentatively considered to belong to P. woodwardi, or at least to those cephalons represented on PI. li., figs. 12 and 13, l)ecause these, too, were the most plentiful cephalons occurring in the same area. As regards the head-shield of G. sweeti., sufficient of it has not been conserved 480 CARBOKiFEKOUS TRlLOBlTES OF AUSTRALIA, to enable one to say positively whether its features place it with the Phillipsia or the Griffithides type; but, except for the un- certainty of the presence on its glabella of the normal lateral furrows of the Phillipsian genus, it does not differ from the head-shields above referred to, which are considered to belong to P. looodwardi Eth. fil. Further, it will be found that G. sweeti, as far as present evidence admits, cannot be separated from Grlffitliides semi)iifenis var. australasica Eth.fi]. This, I think, will be con(;eded after the description and figures of the latter have been studied in conjunction with my remarks on the latter, under P. iroodivardi: and an examination of Plate li., fig. 14^ which is a photo of Mr. Etheridge's type specimen, which shows two tails, the greater portion of a thorax and tail conjoined, an intafylio of a portion of a head, and, between the two tails and on the left top corner, is a view in relief of this intaglio. We are up against a problem here, wliich can be solved only Ijy the discovery of better material: and when it is solved, it seems to me that a new genus or subgenus will be needed for the reception of tliis trilobite with the ten thoracic somites. The following is a fuller description of the species than is given bv Mr. Etheridge It will be seen fronj the photograplis of this species now given, that it was not as correctly figured as it might have been. Complete form suboval. ^}i. Chars. — Cephalon incomplete, apparently granulated throughout, with grHiiules of uniform size. Glabella incomplete, moderately tumid, mesial and anterior furroAvsnot visible (though there appears to be a faint trace of the mesial pair): basnl furrows deep, joining the neck furrow: basal lobes fairly large, suboval: supplementary lobes of moderate size, and suboval : neck-furrow wide and fairly deep: neck-ring stronger than any of the axial rings of the thorax. Fixed and free (;heeks absent. Thorax consisting of ten segments, finely and evenly gran- ulated: axis prominent, diminishing posteriorly very little in prominence, and barely at all in spread: each ring, except the last, has centrally a slightly forward direction, the last ring is stron. C. Reed {op. ci^.) :—" Head-shield semicircnlai', moderately convex, with strong raised rounded border increasing slightly in width towards the front, and separated off by a deep furrow. Genal angles furnished with slender divergent smooth spines, less than half the length of the head shield. At its base is a pair of small nodular basal lobes, in most specimens quite inconspicuous. Two large tubercles are situated in a line down the middle of the glabella, followed by a similar median one on the occipital segment. Occipital segment strong, rounded, separated off by a deep furrow. On cheeks at anterior end of glabella is a pair of large tubercles one on each side. No facial sutures visible. Eyes prominent, reniform, less than half the length of the glabella, distant from the axial furrows about one-third the width of the cheeks, and about their own length from posterior margin. Surface of head-shield, including glabella, border and neck-segment, rather coarsely tuberculated. An indistinct ring of larger tubercles surrounds the eyes, and a large tubercle is situated at each end of eyes on innei- side. Thorax unknown. Pygidium semicircular, slightly convex, with spinose maigin. Axis broad, conical, about one-third the width of the pygidium at front end, tapers rather rapidly to obtuse point, nearh^ touch- ing the border: consists of 9-10 segments, of which eight rings are distinct and completely tuberculated across : the 1st, 3rd, 5th, and 7th have, in addition, a large median tubercle. Lateral lobes consist of six (\ seven in some) pairs of pleurae, of which the last pair is very small: each pleura is gently curved and is divided unequally by a strong longitudinal furrow into a broader, raised, rounded, posterior ridge, and a narrower anterior ridge. The posterior ridge of each pleura crosses a distinct, raised, rounded bordei;, which surrounds the pygidium and bears a large BY JOHN MITCHELL. 485 tubercle at the spot where it crosses, and a single median one behind the axis. The posterior pleural ridges are prolonged into short, recurved, equidistant, and subequal spines, projecting beyond the margin. (In one specimen there seems to be a median spine behind the axis. In another immature example the anterior two or three pairs of spines are half as long as the whole pygidium). Surface of pygidium rather coarsely tuber- culated: the posterior ridge of each pleura bears 4-5 tubercles, and the anterior ridge 5-6 smaller ones. The axial rings bear each 5-7 tubercles." " Dimensions: — Length of head-shield ... ... ... .. 8 'Omni. Width of head-.'^hield ... ... ... ... 4'omm. Length of pyoidiuni ... ... ... ... 2*5 mm. Width of pygidium 4 "0 mm. For affinities, see op. cit. To quote Reed further, he remarks that " McCoy gave as generic characters the circle of tubercles round the eyes and pair of large tubercles at the front end of the glabella, but these may well be considered as of lower classificatory value, and likewise the relatively greater length of the glabella as compared with the European species. It does not, however, seem possible to regard tlie peculiar pygidial characters in the same light, though, as Vogdes (Trans. Acad. So. St. Louis, Vol. v, (1892), p, 617) .says, we have many other genera of trilobites with spinose and non-spinose representatives. The fewer number of segments in the p\'gidium, and the rai.sed spinigerous boi'der separate it from all the European forms. " " The genus or subgenus Phcetonides, as now understood, is partly distinguished for analogous reasons from the typical Proetus; and it seems open to question whether the European species of Brachymetojnis should not be regarded as constituting a distinct group or subgenus for which the name of Brachymeto- pina may be suggested." Personally, I do not think that pygidial characteristics alone are sufficient grounds upon which to found even a 5^ubgenus Loc. and Hor. — Dunvegan, Burragood, and Glen William, 36 486 CARBONIFEROUS TRILOBITES OF AUSTRAF.IA, N.S.W. (Etheridge, Junr., Cat. Austr. Foss., Camb., 1878, p.41). Carboniferous. Bkachymetopus dunstani, sp.iiov. (PI. xlix., figs.15, 16: PI. lii., figs.l, 2). Complete form unknown. Sp. Chars. — Cephalon : length and width 3 mm, and 4 mm., respectively; semielliptic, surface covered with tubercles of vary- ing size, only mildly convex; glabella conical, densely and evenly granulated, very mildly convex, basal furrows well defined and joining the neck-furrow, basal lobes small, greatest width about one-fourth that of the cephalon, length about half that of the cephalon; the surrounding axial furrow relatively deep, cheeks gently convex, bordering the furrow surrounding the glabella; they bear eleven tubercles of uniform size, and follow a course inside of the eyes to the posterior margin: from the front of the eyes there branches from this main bead-like chain of tubercles two other sets (one on each side) of five each, which are arranged along the outer boundary of the eyes; the one or two tubercles in front of, and the one in line with, the back of each eye, appear to be larger than the others of these sets: besides these rows there are a few tubercles bordering the inner edge of the marginal furrow, and the thickened border bears a chain of these small bead-like tubercles of uniform size, and about twenty- five in number, the rest of the surface being finely granulated. The eyes are small, crescentic, close to the glabella and posterior margin; neck-furrow shallow, as are also its lateral extensions: neck-ring fairly robust and granulated, lateral extensions rela- tively strong and narrow; border furrow wide and deep: border strongly tumid and tuberculate; angles, if not spinate, are acute. 06s. — The above Brachymetopus is the first and only specimen of the genus collected from the Carboniferous rocks of Queens- land. It occurs at Trilobite Ridge, Mt. Morgan, Queensland, associated with other genera of trilobites. In several features it resembles lir. strzeleckii Mc(Joy, the prototype, and agrees with this species and Br. Muccoyi in possessing a well-defined cephalic marginal border, bounded inwardly by a well-defined furrow. BY JOHN MITCHELL. 48? The tuberculation, too, on this border is similar in the three species. Dimensionally, this species agrees closely with Br. strzeleckii, judging from the dimensions given by Reed,"^ so also does it in part in the character of the ornamentation, but in part in this respect, it differs very widely, as the description shows. The chief differences between our species and Br. strzeleckii are — 1. The character of the ornamentation. 2. The more conical shape of the glabella in ours, and the absence of the large longi- tudinally placed tubercles on this part. 3. The more acutely rounded cephalon, particularly at the front. 4. The eyes are situated closer to the posterior border, and perhaps to the glabella. 5. The lateral extensions of the neck-ring are narrow and prominent. Without doubt, the two forms are closely related. Since writing the above, I have found, on some of the Queens- land specimens from Rockhampton and Mt. Morgan districts, several pygidia and a portion (cheek) of a cephalic shield, which belong to the genus Brachymetopus. One of these pygidia, which is fairly well preserved, I am assuming to be specifically identical with the cephalon above described. Its description is as follows. Pygidium semicircular, only mildly convex, strongly tuber- culate. Axis moderately prominent, contracting gradually pos- teriorly, ending short of the border bluntly, with less than half its anterior width: it bears five longitudinal rows of tubercles, the individuals of the middle row being much larger than those in the rows on each side of it. There are at least twelve or thirteen rings in the axis, and a central tubercle on ten of these can be clearly seen; this applies also in the case of each row immediately adjacent to the central one. Axial furrows deep. Side-lobes consist of six (? seven) pairs of segments, the ridges of which are very prominent, and bear several relatively large tubercles and some of smaller size: each of the posterior pair merely consists of a tubercle, and each ridge of the anterior pairs bears several tubercles varying much in size, the larger being '■' Geol. .^Ja.^, X.S., Dec. iv., Vol. x.. IIMKS. pp. llKMUfJ. 488 CARBONIFEKOUS TRILOBITES OF AUSTRALIA, situated near the axial furrowy and border; the ribs apparently extend into spines beyond the border, mesial furrows of the ribs deep. Border of moderate width, and apparently only mildly thickened. This pygidium, though it presents in a general way a strong resemblance to that of Br. strzeleckii, exhibits features which help to separate the two forms. The chief differences between their pygidia are — l. The axis of Br. dunstani consists of twelve or thirteen rings, as against ten in the other. 2. Five longitudinal rows of tubercles traverse the axis of the former, while seven rows occur on the latter on the same part. 3. A large tubercle is situated centrally on each axial ring of the former, but only on every alternate axial ring in the latter. 4. There is a smaller number of tubercles on each pleural seg ment of the former than on the similar segments of the latter. The pygidial ornamentation of our species agrees closely with Br. lodiensis Meek. Dedicated to Mr. B. Dunstan, Chief Government Geologist, Brisbane, whose palyeontological discoveries have been numerous and important. Loc. and Hor. — Trilobite Ridge, Mt. Morgan, Queensland, associated with P. ivoodivardi. Lower or Middle Carboniferous. Specimens Nos.F1031, F 1007, Geological Museum, Dept. of Mines, Brisbane; and No.712, Queensland Museum, Brisbane. Brachymetopus, sp. indet. \\\ addition to the head and tail, which served for the descrip- tion of Br. daiisfaui, there occur on specimen No. 712 of the Queensland Museum, Brisbane, no less than three pygidial frag- ments of Brachymetopi, besides the pygidium which served for the foregoing description of Br. dunstani. In the case of two, one is partially superimposed on the other, and the tliird is almost touching these. On specimen F 1007 is a portion of a right cheek, which presents features evidently unlike either Br. strzeleckii or Br. dunstani. Two of the fragmentary pygidia also appear to be new, and maybe specifically identical with the individual to which the cheek-fragment belonged. The material. BY JOHN MITCHELL, 489 liowever, is too fragmentary to describe and figure. The occur- rence of three pygidia in a launch indicates that they may be plentiful, and careful search may bring better specimens to light. KXI^LANATION OF PLATES XLVI.-LIII. Plate xlvi. PliWiima colfinsi Mitchell. I^'ig.s. 1, 2, o, 4. .">. — Fin. 1 i« a photo ( x S) made froni the specimen repre- sented in Fig.'i. The glabellar features are plainly visible. Figs. .'i, 4, T) represent pygidia. and shoM" their characteristic featui-es very well. Figs.4 and "> x | (about). (Coll. Mitchell). PhUlipsia coulteri Mitchell. Figs. (). 7, S. *•. 10. — Different views of one individual. Fig.H x 4, and Fig. 10 nearly natural size. In Figs. 8 and 9, the mild dorsal serration is visil)le. In Fig. 10. the hypostome is plaiidj^ seen, and the latlier large tubercle of the terminal axial ring. (Coll. Mitchell). PJi///ip-^ia hftricejjs Mitchell, Figs.ll. 12.— Cephalon and tail (Colh Mitchell). Grijfit hides conrexicaiidatus Mitchell. Fig. 18.— Dorsal view. (Coll. Mitchell). PJii//ip)s;n (iruiidi.s Ftli. Hi. l''ig.2. — Piioto of the specimen originally descrihetl hy Mr.' H. Flhcridge. Junr. , and for which the specific nam3 r/y-rt/Af//.*.- was first suggested by him. The figure shows how different in character arc the pleural ribs from those of P. rohiisfa; the sutures along the aitit-ulatijig ridges, and the crossing of the border by the anterinr ril»s. (Coll. Geological and Mining Museum, Dept. of Mines, Sydney. Xo.l.lKO). PhiUij,xi, 7. — Four p3-gidia. Thej' exhibit the specific features. Figs. o and 7 are photos of the specimens used bj^ Mr. Etheridge for his figures (PL xxi., figs. 6-7, opj. cit, antea). (Coll. Geological and Mining Museum, Dept. of Mines, Sj'dney. Nos. E 1495, 3534 (2), F 1595). I^hiUipsia colli n-', a pygidium and a portion of a head-shield. (Coll. (Geological and ■Mining Museum. Dept. of Mines, Sydney. Nos. F 14!>6, 1.-)(I0. and l.)0()). J^hi/h'p.-n'a rockhaniptoiitnsis M itchell. Figs.S, 9. — Two pygidia. Fig. 8 is a photo of the specimen sliown in (U'ol. and Pal. (Queensland and New (Guinea, Pl.xliv., fig.G. Fig.9 repre- sents a young individual. (Coll. (Queensland Museum, Brisbane, No.833: and (Jeological Museum. Dept. of Mines, l>risbaiie, No. F7i>2). PhiUipsiai'i) woodwardi Eth. fil. Fig. 10,~(3n this specimen are several pygidia and a remarkable iiypostome, doubtfully assumed to belong to this species; and, in addition, the greater part of a cheek of a Hrachymetopiis; the position of which is indicated by an arrow. (Coll. (Geological Museum, Dept. of Mines, Brisbane. No. 1007). Fig. 11. — Glabella; one of Mr. Etheridge's types. (Coll. (Queensland Museum, Brisbane. No. 707). Plate li. If not otherwise indicated, all the figures are x 'i (about). Phillipsia mori/anensis Mitchell. Fig. 1. — A photo, in two sections, of a specimen on which are parts of at least three individuals, all assumed to belong to this species. The glabellar, pygidial, ocular, and hypostomal features are all dis- cernible, as are also those of the thorax. (Coll. Geological Museum, Dept. of Mines, Brisbane. No.F 1000), BY JOHN MITCHELL 49:; rhilli)>sia brevictp-^ Mitchell. Fi<;K.2, 3. — Photo of a specimen showinj; portions .of two cephalic-shielda and a very perfect pyicidiiini, whose characteristics are represented. Mark the peculiar contraction near the middle of the tail, which suggests a short tail, and a portion of a thorax. Fig.3, a portion of a cephalon. shoN\ inu- limb, mesial furrow on the right, etc. (Coll. Mitchell). riifUipsia connollii Mitchell. Figs. 4-7- — ^A doisal and side-view of the only pygidium known. The pro- portionately very wide border, pi^ominent axis, granulation, and other features are faii-lj- well shown. In Fig. 6, the axial and pleural divisions have been intensified slightl3'. (Coll. (Geological and Mining Museum, Dept. of Mines, Sydney. No. F 1497). Philllpsia sfanvel/ensis Mitchell. Figs.S-lO. — Dorsal and side-view of a very perfect tail, and showing the very prominent, mildly serrated axis clearlj'. In this specimen, all the axial rings (12. doubtfully 13) are visible, as also are eight pleural segments, and the steep striated border. (Coll. (Geological Museum, Dept. of Mines, Brisbane. No.F977). PIn/lipsia xfroiidennx Mitchell. Fig. 11. — A medium-sized tail, having the dorsal part of the axis damaged; but other\\ise exhibiting the normal fetitures. (Coll. Mitchell). Phillipsia icoodicardi Eth, fil. Figs. 12-13.— Portions of cephalons. Fig. 12 shows the glabellar furrows, strong neck-ring, and supplementary lobes, etc., very well. Fig, 13 exhibits these features less clearly. Fig. 12 is from the same speci- men as that figured by Mr. Etheridge, Junr., {op. cit., PI. xliv., fig.5). (Coll. Queensland Museum, Brisbane, No.707; and Geological Museum, Dept. of Mines, Brisbane, No.F 1017). PhiUipsia vmodu:ardi{''.) Eth. fil. Fig. 14. — This photo shows the fragments of trilobite-remains, on which Mr. Etheridge chietly founded his species G. >*emmiferus var. au8- tralasica. On this specimen also occurs the tail of Brarhymetopus dunstani. It remains to be proven that they are not portions of the above species. The tails shown on this specimen have the same number of axial and pleural divisions, and kind of granulation as those included with the cejihalons of P. ivoodwardi. (Coll. Queens- land Museum. Brisbane. No. 712). Plate lii. Phillipsia inoodwardi{'t) Eth. fil., and Brachymetopus dwnstani M'ltaheW. Fig.l. — This represents a portion of specimen No.712 of the Queensland Museum, Brisbane. On it is the pygidium assumed to belong to P. iL'OodAi-a rdi , and a pygidium of Jir. davMani in front of the arrow; ( X 3). 494 CARBONIFEKOUS TRir.OBITES OF AUSTRALIA. Brai-liyine.topus diinstani Mitcliell. etc. Fig.2. — Enlarged poitjon of specimen No. F 1017, (reological Museum, Dept. of Mines, Brisbane. Oh it is the intaglio of Br. dnnstani, showing, fairly well, the chief specific features. The other pygidial and cephalic imprints are assumed to belong to P. iwodirardi Eth. fil. ; ( X 3). Phi/npsia HHperha Mitchell. Fig.3. — Head-shield ( x4) from the same specimen as PI. xlviii., tig. 15. PliiUipskt coUinsi Mitchell. Fig. 4. — A perfect tail of an immature individual; ( x f ), Grijfithides convexicaudatust ^Mitchell. Figs. 5, 6. — Tails about complete. Fig.5 is that of a young individual. Init shows all the axial and pleural divisions. PliUUpsia waterhoiisei Mitchell. Fig. 7. — From a cast; it shows tlie glabellar furrows plainly: and other features. Plate liii. Griffi.thidef>{1) sweet I Eth. fil. Figs. 1,2. — Dorsal aspect, rraces of glabellar furi-ows are visible, especi- ally on the right side generalh'. The important features are well shown. The photos are from the type-specimen; (x2). (Coll. Sweet). Brachymetopus strzeJeckil McCoj'. Figs. 3, 4. — Photos of the enlarged figures of McCoy (lor. cit.). Figs.5, 6.— Photos of Reed's figures of the species (/or rif.). Grijfif hides dnhins Eth. Senr. Fig. 7. — Copy of the figure given by R. Etheridge, Junr.. (in (^eol. Pal, Queensland and New Guinea, 1892, PI. vii., fig. 12). Phif/ipsia iroodwardi Eth. fil. Figs.8, 9.— Copies of Mr. Etheridge's figufes (op. cit., PI. vii.. figs. 13, 15). Griffithidesi'i) seminiferus var. anstralasica. Fig. 10. — Copy of Mr. Etheridge's original figure of a free cheek.