OCCASIONAL PAPERS of the MUSEUM OF NATURAL HISTORY The University of Kansas Lawrence, Kansas NUMBER 113, PAGES 1-16 6 SEPTEMBER 1984 TWO NEW SPECIES OF THE LEPTODACTYLID FROG GENUS PHRYNOPUS, WITH COMMENTS ON THE PHYLOGENY OF THE GENUS By David C. Cannatella^ Within the tribe Eleutherodactylini of the Leptodactylidae there are several genera that are presumed to be derived from various groups of the mega-genus Eleutherodactylus. Among these is the Andean genus Phryno-pus, which consists of fourteen species, five of which were named by Lynch (1975) in his review of the genus. Recent fieldwork in the Andes of northern and central Peru by field parties from the University of Kansas has yielded specimens of two new species, which are described below. ACKNOWLEDGMENTS For loan of specimens and/or provision of working space I am indebted to Marinus S. Hoogmoed, Robert F. Inger, and William E. Duellman. The latter and Thomas J. Berger were enjoyable field compan-ions in Peru; fieldwork was supported by NSF grant DEB 76-09986 to William E. Duellman and by a NSF Graduate Fellowship to the author. Linda Ford and Darrel Frost critically read the manuscript. Laboratory facilities were made available by the Center for Biomedical Research of The University of Kansas. Permits for collecting were generously pro-vided by Ing. Carlos Ponce del Prado. Direccion de Conservacion, Lima, Peru. MATERIALS AND METHODS Statistical analyses were done using BMDP statistical software (Dixon, 1981). Cladograms were produced using the WAGNER78 program written by J. S. Farris. Drawings of the skulls were executed by means of a Wild M-8 microscope with camera lucida attachment. Skeletons were ' Museum of Natural History and Department of Systematics and Ecology, The University of Kansas, Lawrence, Kansas 66045 U.S.A.
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