SYSTEMATIC STUDIES OF DARTERS 3 Paratypes. — A total of 87 specimens deposited as follows: 18 — Illinois Natural History Survey (INHS 75735, 10 specimens, 36-5S mm SL, same collection data as holotype; INHS 75576, 8 speci-mens, 35-64 mm SL, same locality as holotvpe, 11 Dec. 1976); 6— U. S. National Museum (USNM 216895, .34-46 mm SL, same collection data as holotype); 6 — University of Michigan Museum of Zoology (UMMZ 200208, 35-48 mm SL, same collection data as holotype); 6— University of Tennessee (UT 91.1332, 37-45 mm SL, same collection data as holotype); 6— Tulane University (TU 101391, 37-50 mm SL, same collection data as INHS 75576); 6— University of Alabama ( UAIC 5341.01, 39-48 mm SL, same collection data as LNHS 75576); 6— Northeast Louisiana University (NLU 35160, 41-51 mm SL, same collection data as INHS 75576)'; 33— The University-of Kansas (KU 11442, 34-59 mm SL, Dry Fork, 3 km W Gordonsville, Smith Co., TN, 7 April 1966). Etymology. — The name olivaceum refers to the olive color of nonbreeding individuals. The common name, dirty darter, was suggested by Dr. David A. Etnier, University of Tennessee, and refers to the drab color and indistinct mottling on the sides. Diagnosis. — The subgenus Catonotus of Etheostoma was diag-nosed by Kuehne and Small (1971). The presence of an uninter-rupted infraorbital canal in E. olivaceum and E. neopterum Howell and Dingerkus ( 1978 ) requires that the subgeneric description of the infraorbital canal be modified to "infraorbital canal interrupted or uninterrupted." In addition, the cheek, breast, nape and prepectoral area may be scaled or unsealed. Etheostoma olivaceum is distinguished from all other members of the subgenus by the following combination of characteristics: an uninterrupted infraorbital canal with eight infraorbital pores, 10 preoperculomandibular pores, 13 or more pored lateral line scales, scales present on nape and prepectoral area, scales absent on cheek and opercle, distinct black vertical bands on caudal fin, branchiostegal membranes slightly connected, no bar pattern on cheek, no red or blue pigments, no dark suborbital bar, usually nine dorsal spines, 12-13 dorsal rays, and seven or eight anal rays. Comparisons.— Etheostoma olivaceum is separated from all species of Catonotus except E. neopterum by the uninterrupted infraorbital canal and from E. neopterum by the absence of scales on the opercle. E. olivaceum is further separated from the barcheek species of Catonotus (virgatum, obeyense, barbouri, smithi, striatulum) by the lack of the bar pattern on the cheek, lack of red and blue pigments, and modally 7-8 anal rays; from E. flahel-lare, E. kennicotti, and the barcheeks by the presence of scales on the nape and prepectoral area; and from its closest relative, E. squamiceps, by the absence of scales on cheeks and opercles, ab-sence of a dark suborbital bar, modally seven scales above the
BioStor
