PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES Vol. 42, No. 19, pp. 455^96, 74 figs., 22 tables ^ May 14, 1982 SMALL TERRESTRIAL GROUND-BEETLES OF " ' • ^ j CENTRAL AMERICA (CARABIDAE: JUH J 1 jgg2 BEMBIDIINA AND ANILLINA) j By — :.lzJJ2i:^jj^--iass. ' Terry L. Erwin Department of Entomology, Smithsonian Institution, Washington, D.C. 20560 Abstract: This paper provides a synopsis of the tribe Bembidiini with an overview of the three included subtribes, Bembidiina, Tachyina, and Anillina for Central America. Anillina is thought to be a polyphyletic grade derived from genus Paratachys of the Tachyina, but no formal classificatory action on this hypothesis is taken. Based on structural characters, Horologion, previously classified as Psydrini or Trechini, probably also belongs to this grade. Thirteen new species of Bembidion are described from the following type-localities: purulha, 22.0 km south of Puruiha, Baja Verapaz, Guatemala; lavernae, Cerro Buenavista, San Jose Province, Costa Rica; quetzal, 20.0 km west of San Pedro Sacatepequez at Rio Nahuala, Quezaltenango, Guatemala; diabola, Volcan Irazii, Cartago Province, Costa Rica; edwardsi, Cerro Buenavista, San Jose Province, Costa Rica; aeger, Cerro de la Muerte, Cartago Province, Costa Rica; chiriqui, 8.0 km west of Boquete, Chiriqui Province, Panama; nahuala, 27.0 km northwest of San Marcos, San Marcos, Guatemala; /ramat, 4.8 km east of San Mateo Ixtatan, Huehuetenango, Guatemala; cortes. La Lima, Cortes, Honduras; armuelles, Puerto Armuelles, Chiriqui Province, Panama; barrensis, Barro Colorado Island; ixtatan, 4.8 km east of San Mateo Ixtatan, Huehuetenango, Guatemala. Four new species of Geocharidius are described from the following type-localities: romeoi, 13.0 km south of Purulha, Baja Verapaz, Guatemala; gimlii, 1.1 km south of San Juan Ixcoy, Huehuetenango, Guatemala; phineus, 1.6 km south of Pantin, Baja Verapaz, Guatemala; tagliantii, Totonicapan, Totonicapan, Guatemala. The 22 Bembidion species presently known from Central America are arrayed in nine species groups. Corresponding groups for the five known Geocharidius species are not used because of the small size of the fauna and their questionable relationships with poorly known northern taxa. Distribution and relationships are discussed in general, but detailed analysis is deferred until species-group taxonomy is undertaken on a broader geographic scale. Notes are provided on natural history and distribution for each species; dot maps illustrate the known range of each taxon. Introduction Bates's papers. The anilline genus Geocharidius Nearly 100 years have passed since the last was discussed more recently by Taglianti (1973), comprehensive paper on Central American but his coverage did not add new species for Bembidion was produced by H. W. Bates. Dar-Central America. Presently, five species are lington (1934) described a single species. Other known from the area, an increase of four since than these two contributions, the genus has re-Bates's contributions to Biologia Centrali-ceived no taxonomic attention for Central Americana (Bates 1882, 1884). It therefore can America. Presently, 22 species are known from be assumed that after nearly 100 years the Bem-the area, an increase of 17 since the time of bidion fauna of Central America (defined here [455]
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