Woods Hoi PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES Vol. 49, No. 11, pp. 405-421, 4 figs., 2 tables. March 21, 1997 SPECIES OF THE HAZELAE GROUP OF PLATYMANTIS (AMPHIBIA: RANIDAE) FROM THE PHILIPPINES, WITH DESCRIPTIONS OF TWO NEW SPECIES By Walter C. Brown Department ofHerpetology, California Academy of Sciences, San Francisco, California 94118. and Rafe M. Brown Department of Zoology, 212 Biological Sciences Building, Miami University, Oxford, Ohio 45056. and Angel C. Alcala Commission on Higher Education, DAP Building, San Miguel Ave., Ortigas Center, Pasig City, Philippines. Three species Groups of Philippine anurans of the genus Platymantis are recognized. The Groups are diagnosed on the basis of combinations of digital characters. Species of the hazelae Group are revised. Eight species are placed in this Group, two of them (reticuLUus and panayensis) previously undescribed. These small to moderate-sized (20-30 mm for males and 25-09 mm for females) frogs are forest species from mountain areas of islands in the Greater Negros and Greater Luzon island groups. Received April 16, 1996. Accepted October 25, 19%. The genus Platymantis, as it is currently diag-nosed, has two centers of diversity, one in the Philippine Islands and one in the Bismarck-Solo-mon Islands. There are three small extensions beyond these archipelagos. New Guinea has three species, Fiji two species, and the Palau Islands one species. The relationships of the three Philippine Groups, recognized in this study, to each other and to the Groups that apparently exist in other parts of the range of the genus are unclear at this time. Phylogenetic studies of the Asiatic island ranids currently underway may provide some answers. The latest revision of Philippine Platymantis is that of Inger (1954). He followed Taylor (1920) in assigning both the large-disked and small-disked species to one genus, whereas Boulenger (19 18) and Noble ( 193 1) had placed these assem-blages in two genera (Cornufer and Platyman-tis). Inger (1954) regarded disk size as simply a measure of the degree of specialization of these structures representing a continuum. Inger recognized seven Philippine species, five with broadly dilated finger disks and two without or with small finger disks. Since 1954, five more Philippine species have been described. Of these [405]
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