J. HYM. RES. Vol. 9(1), 2000, pp. 18-28 Revision of the Australian Tiphiid Genus Leiothynnus (Hymenoptera: Tiphiidae: Thynninae) Lynn S. Kimsey Bohart Museum of Entomology, Department of Entomology, University of California, Davis, CA 95616, USA Abstract. — The Australian thyronine genus Leiothynnus Turner is revised, describing five new species from Queensland, including cardalae, ferricolus, linnis, multimaculatus and ochrotarsus, and two previously described species, mackayensis (Turner) and spinigerus Turner, also from Queensland. Distribution maps of species and a key to males are given. Most Australian genera of Thynninae with Agriomyia Guerin de Meneville, in-are widespread, occurring in many Aus-eluding the flat male face (Fig. 5), with ex-tralian states, with some even extending treme reduction of the antennal lobes, and up into New Guinea, New Caledonia and simple male epipygium, lacking the well-adjacent islands. However, there are some developed transverse carina or ridge typ-small genera with relatively restricted dis-ical of Epactiothynnus and related genera, tributions, such as Oncorhinothynnus, However, remnants of this ridge can be which is only found in west central West-seen, particularly in cardalae and ochrotar-ern Australia, or Gymnothynnus Turner, sus. Leiothynnus can be distinguished from from Northern Territory and western these and other genera by the long brush Queensland. The genus Leiothynnus Turn-of setae on the base of the male premen-er is one of these relatively localized turn, the darkly stained marginal cell in groups. All Leiothynnus species have been the forewing, and slender, petiolate male collected from eastern Queensland. In ad-abdomen. dition to the two described species, mack-Leiothynnus species differ from one an-ayensis (Turner) and spinigerus Turner, other in modifications of the male legs, ab-there are five new ones, which are de-dominal apex and to some extent color, scribed below. All of the species treated below have the The specific relationships between male thorax with dense, nearly contiguous Leiothynnus and other thynnine genera are punctation, and the propodeal punctation not fully resolved. Leiothynnus shares a obscured by fine shagreening. These fea-number of features with Iswaroides Ash-tures may or may not be significant at the mead and Epactiothynnus Turner, includ-species level. Collecting seems to be too ing in the male the well-developed and patchy to say whether or not there are ad-somewhat arcuate penis valves, volsella ditional undescribed species. Too few fe-U-shaped in cross-section, stipes arcuate males have been collected to generalize with long marginal fringe of setae, vertex about diagnostic features among the spe-with small reddish spot behind the dorsal cies in females, although there appear to eye margin, and a large oral plate. In the be differences in the overall shape of the female, the pygidium has two submedial head and development of carinae or lobes longitudinal carinae, subtended by a long on abdominal segment V. tuft of setae. A few characters are shared Specimens were obtained from the fol-yHWHUmi^HHHIIIIIflBDHHIBIMUaMHHWBnHiHaHl
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