^/a%5i Ji//I/ ^Hy OCCASIONAL PAPERS ^ '^59 of the ^A//Ve^4^0 MUSEUM OF NATURAL HISToftV The University of Kansas Lawrence, Kansas NUMBER 131. PAGES 1-12 JUNE 7. 1989 NEW SPECIES OF HYLID FROGS FROM THE ANDES OF COLUMBIA AND VENEZUELA By William E. Duellman^ Until recently, hylid frogs of the genus Hyla were thought to be poorly represented in the Andes. Cochran and Coin (1970) recognized three species in the Hyla variabilis group, one in the Hyla labialis group, and one in the Hyla bogotensis group in the Andes of Colombia. In addition, Hyla platydac-tyla and H. lascinia were recognized in Venezuela (Rivero, 1961, 1969). Including the species named herein, 25 species of Hyla are known in the Andes north of the Huancabamba Depression in extreme northern Peru. These may be arranged in what seem to be four natural groups. The Hyla bogotensis group consists of 13 species, all but one of which are restricted to the Andes. Members of this group inhabit streams in montane cloud forests from Costa Rica to the Merida Andes in Venezuela and the Amazonian slopes of the Andes in Peru (Duellman, 1972; Duellman and Altig, 1978; Ruiz and Lynch, 1982; Myers and Duellman, 1982;LaMarca, l9S5).ThQ Hyla labialis group consists of three species in the Cordillera Oriental of Colombia and Venezuela and in the Merida Andes. The Hyla columbiana group (=Hyla variabilis group of Cochran and Coin, 1970) contains three species in the Cordillera Occidental of Ecuador and Colombia and the upper valleys of the Rio Cauca and Ri'o Magdalena in Colombia (Duellman and Trueb, 1 982). The large stream-dwelUng species of the Hyla larinopygion group were first discovered in 197 1 . The seven known species have restricted distributions in upper montane cloud forests in Ecuador and Colombia (Duellman, 1973; Duellman and Altig, 1978; Duellman and Berger, 1982; Ruiz and Lynch, 'Curator, Divison of Herpelology, Museum of Natural History, and Professor, Department of Systematics and Ecology, The University of Kansas, Lawrence, Kansas 66045-2454 U.S.A.
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