The phytogeny, classification and evolution of parasitic wasps of the subfamily Ophioninae (Ichneumonidae) Ian D. Gauld Department of Entomology, British Museum (Natural History), Cromwell Road, London SW7 5BD Contents Synopsis 62 Introduction 62 The research potential of Ophioninae 63 The aim of the present work 63 Terminology 63 Material examined 63 Discussion of methodology 63 Choice of parsimony method utilized 64 The advantages and disadvantages of the parsimony method 65 Compatibility methods utilized 66 Value of LEQU . B AS for overcoming some problems of compatibility analysis 67 The operational procedure adopted in this study 70 An apologia for subjectivity in cladistic analysis 70 The holophyly of the subfamily 70 The systematic position of the subfamily 71 Polarity determination and character coding 73 Characters used in the study 74 The characters and their polarity 74 Preliminary remarks on the relative values of the characters 79 The phylogenetic analysis 80 Preliminary analysis of data 80 Analysis using Le Quesne test program, LEQU. B AS 80 Analysis using FOURS program 85 Phylogenetic analyses of groups of ophionine taxa 87 The Euryophion group 87 The Thyreodon group 88 The Dictyonotus/Ophionopsis group 89 The Thyreodon + Euryophion + Dictyonotus complex 90 The Stauropoctonus group 95 The Enicospilus + Stauropoctonus complex 95 The major groups and their relationship with the unassigned taxa 107 The inter-relationship of the two major generic complexes 107 The position of the unassigned taxa 107 Discussion of suggested phylogeny 116 A classification of the Ophioninae 118 Zoogeographic discussion 119 The subfamily Ophioninae 122 The Ophion genus-group 122 The Sicophion genus-group 128 The Eremotylus genus-group 130 The Thyreodon genus-group 133 BRITISH MUSEl {NATURAL HISTORY 30AUGI9& PRESENTED GENERAL LIBRA Bull. Br. Mus. not. Hist. (Ent.) 51 (2): 61-185 Issued 29 August 1985
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