ARDISIA LUNDELLIANA,A NEW SPECIES OFMYRSINACEAE FROMPANAMA The efforts of the Missouri Botanical Garden'splant exploration program in Panama continue toyield new and phytogeographically interesting taxa,even in areas as well known as western Panama.During routine identification of plants collected byGordon McPherson, I discovered a hitherto un-described species of Ardisia, which is describedbelow:Ardisia lundelliana Pipoly sp. nov. TYPE: Pan-ama. Chiriqui: vic. Fortuna Dam, forested slopes along ridge at S boundary of watershed, 8�45'N, 8�15'W, 1,250 m. Tree 4 m; calyx brown; corolla pale pink; fr. green, 28 Apr. 1986, G. McPherson 9107 (holotype, MO; photocopy, US). Figure 1. Ob ramulos foliorum costas abaxiales necnon inflores-centiarum rhachides rufo-stellato-tomentosos et furfura-ceo-stipitato-lepidotos, inflorescentias cernuas, floressubcorymbosos, pedicellos recurvatos, antheras lineo-lan-ceolatas et longitudinaliter dehiscentes A. brenesii arcteaffinis, sed ab ea foliis praeclare petiolatis (non sessilibus),monomorphis (non dimorphis), ellipticis (non oblanceolatisvel obovatis nec oblanceolatis vel oblongis), basibus cu-neatis (non auriculatis) atque apicibus acutis (non acu-minatis), sepalis 5.2-6.0 (non 1.4-1.6) mm longis mar-ginalibusque integerrimis (non erosis), petalisque 12.0-13.9 (non 3.9-4.5) mm longis, intus prope basin verru-cosis (non laevis) statim cognoscitur. Tree to 4 m, the branchlets, petioles, abaxialleaf costae, inflorescence rachis, pedicels and calyxrufous-stellate tomentose and furfuraceous stipi-tate-lepidote. Leaves petiolate, the petioles mar-ginate, (1.1-)1.5-2.0 cm long, the blades coria-ceous, elliptic, (7.0-)11.0-16.7(-18.0) cm long,(4.0-)-5.2-7.7 cm wide, apically acute, basallycuneate, glabrous above, densely furfuraceous stip-itate-lepidote below, the margins entire, revolute.Inflorescence terminal, pendent, paniculate, 8.0-15.0 cm long, the peduncle 2.3-2.7 cm long, therachis linear in flower, becoming flexuous in fruit.Flowers corymbose; pedicels terete, 0.6-1.4 cmlong; sepals erect, coriaceous, dextrorsely con-torted, widely ovate, 5.2-6.0 mm long, 3.6-4.0mm wide, apically acute, asymmetric, basally au-riculate on one side, the auricle and margin scar-ious, glandular-ciliate, densely rufous stellate andfurfuraceous lepidote without, sparsely stipitate-lepidote basally within, black lineate-punctate; pet-als reflexed at anthesis, chartaceous, lanceolate,12.0-13.9 mm long, 3.9-4.5 mm wide, apicallyattenuate, slightly asymmetric, glabrous, denselyblack punctate and lineate-punctate, pink, verru-culose basally within, the margin white, subentire,irregular; stamens 10-12.5 mm long, the filamentshyaline, 6.0-6.3(-6.9) mm long, glabrous, the an-thers linear-lanceolate, concolorous, 4.0-5.9 mmlong, apically apiculate, basally sagittate, dorsi-fixed, the connective inconspicuously black-punc-tate dorsally, longitudinally dehiscent; ovary glo-bose, 5.3-5.7 mm tall, 3.0-3.1 mm diam., denselyorange-punctate; style 9.1-9.4 mm, exserted atanthesis, eccentric or not, densely and prominentlyorange-punctate; stigma punctiform; placenta glo-bose, obtuse apically, 1.2-1.3 mm long and indiameter, the ovules 19-24, pluriseriate. Fruit glo-bose, 5 7 mm diam. (immature), prominently ver-rucose, inconspicuously punctate. Ardisia lundelliana belongs to a group of spe-cies often referred to the genus AuriculardisiaLundell. Because the salient features defining thisgroup (Lundell, 1981) have been encountered ina number of paleotropical members of the morebroadly defined Ardisia Swartz, this species is as-signed to Ardisia until a more thorough, pantrop-ical survey is completed (Pipoly & Stone, in prep.).With rufo stellate-tomentose abaxial leaf midribs,inflorescence rachises and branchlets, pendent,subcorymbose flowers on panicles, and linear-lan-ceolate and longitudinally dehiscent anthers, A.lundelliana is clearly most closely related to A.brenesii (including A. limonensis Lundell). How-ever, A. lundelliana is readily separable by itspetiolate (not sessile), monomorphic (not dimor-phic), and elliptic (not oblanceolate or obovate)leaves with cuneate (not auriculate) bases and acute(not acuminate) apices. Floral characters allowingeasy recognition include the longer, entire sepalsANN. MISSOURI BOT. GARD. 78: 524-526. 1991.
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