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BULLETIN OF THE SOUTHERN CALIFORNIA ACADEMY OF SCIENCES Volume 74 August 26, 1975 Number 2 TAXONOMIC CHARACTERS OF THE GENUS OCHODAEUS SERVILLE WITH DESCRIPTIONS OF TWO NEW SPECIES IN THE O. PECTORALIS LECONTE SPECIES COMPLEX (COLEOPTERA: SCARABAEIDAE) David C. Carlson^ Abstract: The mandibles, stridulatory peg. and male genitalia of the adults of eight species of Ochodaeiis found in the United States have useful characters for the delineation of species and are described and figured. The most useful taxonomic feature of the male genitalia is the aedeagal sac. The taxonomic characters of Ochodaeiis are reviewed and a key to the species of the O. pecloralis species complex is presented. The Ocliodaeiis pecloralis species complex is erected to contain the nominate species O. pecloralis LeConte and two new species which are described. The other five species considered are O. biariuuliis LeConte. O. iiuirmatiis Schaeffer, O. kansaniis Fall. O. praesidii Bates, and O. simplex LeConte. A lectotype is designated for O. iiuirmatus Schaeffer. The genus Ochodaeiis Serviile is one of six genera included in the subfamily Ochodaeinae. Approxi- mately half of the sixty species of Ochodaeiis occur in the New World (Arrow, 1912) and of these, twenty are from the United States, ten from Mexico and Central America, and five from South America. Species of Ochodaeiis are distributed throughout North America with the greatest num- ber occurring in the arid southwestern United States and northern portions of Mexico. Only three species are recorded from Canada (Howden. 1968). Previously, only a few species were known to be common to the United States and Mexico. Examination of several thousand speci- mens from the southwestern United States and northern Mexico discloses that many species are common to both areas. The adults are nocturnal. In most areas where Ochodaeiis have been collected, more than one species usually is taken, and it is not uncommon to take three or four species at the same locality on the same night. Very little is known about their biology or immature stages. The last revisionary work on the United States species of Ochodaeiis was by Fall (1909) and the genus is cinrently in need of revision. The present work is an attempt to discover new taxo- nomic characters and to evaluate their potential for use in revising the genus. Serviile (1828) described Ochodaeiis and West- wood (1852) published the initial synoptic work. The New World species have been revised by LeConte (1868), Horn (1876), Fall (1909), and Arrow ( 1911). Arrow ( 1904) described the position and mor- phology of the stridulatory peg and pectinate mesotibial spur (Fig. 15). Using characters as- sociated with these structures he erected the sub- family Ochodaeinae with Ochodaeiis as its only included genus. Previously, Ochodaeiis was placed in the Orphniidae. The relationships of Ochodaeiis to other Scarabaeidae have been con- sidered by Ritchcr (196')a. b) and Hollosvay (1972). METHODS Prcpanilion of the male genitalia.

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Taxonomic characters of the genus Ochodaeus Serville with descriptions of two new species in the O. pectoralis Leconte species complex (Coleoptera: Scarabaeidae)

Bulletin of The Southern California Academy of Sciences 74: 49-65 (1975)

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